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	Author	Ahrendt, L.P.; Christensen, J.W.; Ladewig, J.
	Title	The ability of horses to learn an instrumental task through social observation			Type	Abstract
	Year	2012	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	139	Issue	1	Pages	105-113
	Keywords	Horse; Social learning; Social interaction; Instrumental task; Investigative behaviour; Aggression
	Abstract	The ability of horses to learn through social observation may ease the implementation of new management systems, because the use of automatic feeders etc. by naive horses could be facilitated by observation of experienced horses. However, previous studies found no documentation for observational learning abilities in horses. This study aimed to investigate the ability of horses to learn an instrumental task from a familiar conspecific when social interaction was allowed during the demonstration. Two similar experiments were performed. In the first experiment, Observer horses (n=11) participated in ten successive demonstrations, where a trained Demonstrator opened an operant device by pushing a sliding lid aside with the muzzle in order to obtain a food reward. Immediately after the demonstrations the Observer horses were given the opportunity to operate the device alone. Control horses (n=11) were aware that the device contained food but were presented to the operant device without demonstration of the task. The learning criterion was at least two openings. Accomplishment of and latency to accomplish the learning criterion, and investigative behaviour towards the operant device were recorded. Five Observers and one Control, out of the eleven horses in each treatment group, accomplished the learning criterion. Even though this presents a high odds ratio (OR) in favour of the Observer treatment (OR=7.6), there was no significant difference between the treatment groups (P=0.15). Analysis of investigative behaviour showed, however, that the demonstrations increased the motivation of the Observer horses to investigate the device. Subsequently, a similar experiment was performed in a practical setting with 44 test horses (mixed age, gender and breed). We used the same operant device and the same number and type of demonstrations, although the horses were held on a loose rope to minimise aggression. In this second experiment, six of 23 Observer horses and five of 21 Control horses learned the instrumental task, representing no influence of the demonstration. Thus, this study did not demonstrate an ability of horses to learn an instrumental task through observation.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ S0168-1591(12)00087-1			Serial	5773
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	Author	Ahrendt, L.P.; Labouriau, R.; Malmkvist, J.; Nicol, C.J.; Christensen, J.W.
	Title	Development of a standard test to assess negative reinforcement learning in horses			Type	Journal Article
	Year	2015	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	169	Issue		Pages	38-42
	Keywords	Algometry; Horse behaviour; Learning performance; Operant conditioning; Pressure-release; Horse training
	Abstract	Most horses are trained by negative reinforcement. Currently, however, no standardised test for evaluating horses' negative reinforcement learning ability is available. The aim of this study was to develop an objective test to investigate negative reinforcement learning in horses. Twenty-four Icelandic horses (3 years old) were included in this study. The horses were tested in a pressure-release task on three separate days with 10, 7 and 5 trials on each side, respectively. Each trial consisted of pressure being applied on the hindquarter with an algometer. The force of the pressure was increased until the horse moved laterally away from the point of pressure. There was a significant decrease in required force over trials on the first test day (P<0.001), but not the second and third day. The intercepts on days 2 and 3 differed significantly from day 1 (P<0.001), but not each other. Significantly stronger force was required on the right side compared to the left (P<0.001), but there was no difference between first and second side tested (P=0.56). Individual performance was evaluated by median-force and the change in force over trials on the first test day. These two measures may explain different characteristics of negative reinforcement learning. In conclusion, this study presents a novel, standardised test for evaluating negative reinforcement learning ability in horses.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6650
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	Author	Ajie, B.C.; Pintor, L.M.; Watters, J.; Kerby, J.L.; Hammond, J.I.; Sih, A.
	Title	A framework for determining the fitness consequences of antipredator behavior			Type	Journal Article
	Year	2007	Publication	Behavioral Ecology	Abbreviated Journal	Behav. Ecol.
	Volume	18	Issue	1	Pages	267-270
	Keywords
	Abstract	Behavioral ecologists have long been interested in understanding the adaptive value of antipredator behavior (Sih 1987Go; Lima and Dill 1990Go; Lima 1998Go). A recent review by Lind and Cresswell (2005)Go, however, noted some important difficulties with quantifying the fitness consequences of antipredator behaviors. In essence, Lind and Cresswell suggest that most studies do not provide strong evidence on the adaptive value of antipredator behavior because they do not consider 1) trade-offs between antipredator and reproductive performance, 2) the abilities of organisms to avoid fitness losses associated with constraints on focal traits by employing behavioral alternatives (behavioral compensation), and 3) the effects of behavioral defenses at different stages of the predation sequence. The authors rightfully assert that an understanding of these issues can only be accomplished by measuring multiple traits and fitness components (i.e., survival and reproduction). Nevertheless, the question of how to integrate such data into
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	Notes	10.1093/beheco/arl064			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4087
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	Author	ALEXANDER, F
	Title	Digestion in the horse			Type	Journal Article
	Year	1963	Publication	Zoologische Garten	Abbreviated Journal	Zool. Garten.
	Volume		Issue		Pages	259-268
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	Abstract
	Address
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	Notes	<a href=“http://references.equine-behaviour.de/klingel.htm”>from Prof. Hans Klingels Equine Reference List</a>			Approved	no
	Call Number	refbase @ user @			Serial	636
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	Author	Amé, J.-M.; Halloy, J.; Rivault, C.; Detrain, C.; Deneubourg, J.L.
	Title	Collegial decision making based on social amplification leads to optimal group formation			Type	Journal Article
	Year	2006	Publication	Proceedings of the National Academy of Sciences of the United States of America	Abbreviated Journal	Proc. Natl. Acad. Sci. U.S.A.
	Volume	103	Issue	15	Pages	5835-5840
	Keywords	Animals; Blattellidae/physiology; Choice Behavior; Decision Making; Leadership; Social Behavior
	Abstract	Group-living animals are often faced with choosing between one or more alternative resource sites. A central question in such collective decision making includes determining which individuals induce the decision and when. This experimental and theoretical study of shelter selection by cockroach groups demonstrates that choices can emerge through nonlinear interaction dynamics between equal individuals without perfect knowledge or leadership. We identify a simple mechanism whereby a decision is taken on the move with limited information and signaling and without comparison of available opportunities. This mechanism leads to optimal mean benefit for group individuals. Our model points to a generic self-organized collective decision-making process independent of animal species.
	Address	Service d'Ecologie Sociale CP231, Universite Libre de Bruxelles, Avenue F. D. Roosevelt 50, B-1050 Brussels, Belgium
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0027-8424	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16581903			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2042
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	Author	Amodio, P.; Boeckle, M.; Schnell, A.K.; Ostojic, L.; Fiorito, G.; Clayton, N.S.
	Title	Grow Smart and Die Young: Why Did Cephalopods Evolve Intelligence?			Type	Journal Article
	Year	2018	Publication	Trends in Ecology & Evolution	Abbreviated Journal	Trends. Ecol. Evol.
	Volume		Issue		Pages
	Keywords
	Abstract	Intelligence in large-brained vertebrates might have evolved through independent, yet similar processes based on comparable socioecological pressures and slow life histories. This convergent evolutionary route, however, cannot explain why cephalopods developed large brains and flexible behavioural repertoires: cephalopods have fast life histories and live in simple social environments. Here, we suggest that the loss of the external shell in cephalopods (i) caused a dramatic increase in predatory pressure, which in turn prevented the emergence of slow life histories, and (ii) allowed the exploitation of novel challenging niches, thus favouring the emergence of intelligence. By highlighting convergent and divergent aspects between cephalopods and large-brained vertebrates we illustrate how the evolution of intelligence might not be constrained to a single evolutionary route.
	Address
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	Publisher	Elsevier	Place of Publication		Editor
	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN	0169-5347	ISBN		Medium
	Area		Expedition		Conference
	Notes	doi: 10.1016/j.tree.2018.10.010			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6508
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	Author	Anderson, C.; Franks, N.R.
	Title	Teams in animal societies			Type	Journal Article
	Year	2001	Publication	Behavioral Ecology	Abbreviated Journal	Behav. Ecol.
	Volume	12	Issue	5	Pages	534-540
	Keywords	animal societies, cooperation, division of labor, groups, invertebrates, task types, teams, vertebrates
	Abstract	We review the existence of teams in animal societies. Teams have previously been dismissed in all but a tiny minority of insect societies. “Team” is a term not generally used in studies of vertebrates. We propose a new rigorous definition of a team that may be applied to both vertebrate and invertebrate societies. We reconsider what it means to work as a team or group and suggest that there are many more teams in insect societies than previously thought. A team task requires different subtasks to be performed concurrently for successful completion. There is a division of labor within a team. Contrary to previous reviews of teams in social insects, we do not constrain teams to consist of members of different castes and argue that team members may be interchangeable. Consequently, we suggest that a team is simply the set of individuals that performs a team task. We contrast teams with groups and suggest that a group task requires the simultaneous performance and cooperation of two or more individuals for successful completion. In a group, there is no division of labor--each individual performs the same task. We also contrast vertebrate and invertebrate teams and find that vertebrate teams tend to be associated with hunting and are based on individual recognition. Invertebrate teams occur in societies characterized by a great deal of redundancy, and we predict that teams in insect societies are more likely to be found in large polymorphic (“complex”) societies than in small monomorphic (“simple”) societies.
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	Notes	10.1093/beheco/12.5.534			Approved	no
	Call Number				Serial	2070
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	Author	Anderson, M.K.; Friend, T.H.; Evans, J.W.; Bushong, D.M.
	Title	Behavioral assessment of horses in therapeutic riding programs			Type	Journal Article
	Year	1999	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	63	Issue	1	Pages	11-24
	Keywords	Horses; Therapeutic riding; Temperament; Cortisol; Catecholamines
	Abstract	A behavioral assessment of horses who were being used and not used in therapeutic riding programs was conducted to help determine useful methods of selecting horses for use in therapeutic riding programs. A total of 103 horses (76 horses from five therapeutic riding centers and 27 non-therapeutic riding horses from four sites) were used. Temperament survey for each horse were completed by three riding instructors at each therapeutic riding center or by the individual most knowledgeable about the horse at the other sites. Twenty personality traits from the survey were used to quantify temperament. Concentrations of plasma cortisol, norepinephrine and epinephrine were also measured in each horse. A reactivity test was then conducted which involved introducing three novel stimuli: a walking and vocalizing toy pig placed on a cardboard surface in front of the horse for 20 s; popping a balloon near the horse's flank area; and suddenly opening an umbrella and holding it open in front of the horse for 20 s. Reactions (expressions, vocalizations and movement) to each of the stimuli were scored and used to calculate an average reactivity score for each horse. The therapeutic riding instructors did not often agree on the temperament of their center's horses. The personality trait ratings made by the therapeutic riding instructors at each center were on average significantly correlated (P<0.01, r>0.52) for only 37.8% of the horses for any two instructors and 7.8% for three instructors. No significant correlations were found between temperament, reactivity, and the hormone concentrations (r<0.19), but regression analysis indicated a possibility of predicting temperament from the reactivity score and hormone concentrations (P<0.08). There was also a tendency for relationships between extremes in temperament (desirable vs. undesirable) and the hormone concentrations (P<0.09), and between extremes in reactivity (low vs. high) and the hormone concentrations (P=0.08). The difference in ratings among riding instructors indicates a need for more collaboration between instructors when evaluating horse temperament. This study also indicates that it was very difficult to objectively determine the suitability of horses for therapeutic riding programs regarding their temperament and reactivity, probably because other traits (e.g., smoothness of gait) are also considered very important.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4812
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	Author	Andrew, R.J.; Osorio, D.; Budaev, S.
	Title	Light during embryonic development modulates patterns of lateralization strongly and similarly in both zebrafish and chick			Type	Journal Article
	Year	2009	Publication	Philosophical Transactions of the Royal Society B: Biological Sciences	Abbreviated Journal	Phil. Trans. Biol. Sci.
	Volume	364	Issue	1519	Pages	983-989
	Keywords
	Abstract	Some aspects of lateralization are widespread. This is clear for the association between left-eye (LE) use and readiness to respond intensely to releasing stimuli presented by others, which has been found in representatives of all major groups of tetrapods and in fishes. In the chick, this behavioural asymmetry is linked developmentally to greater ability to sustain response against distracting stimuli with right-eye (RE) use, in that both reverse with the reversal of the normal RE exposure to light. In the zebrafish, the same two asymmetries (normally) have similar associations with the LE and the RE, and both also reverse together (owing to epithalamic reversal). Here, we show that light exposure early in development is needed in zebrafish to generate both asymmetries. Dark development largely abolishes both the enhanced abilities, confirming their linkage. Resemblance to the chick is increased by the survival in the chick, after dark development, of higher ability to assess familiarity of complex stimuli when using the LE. A somewhat similar ability survives in dark-developed zebrafish. Here, LE use causes lesser reliance on a single recent experience than on longer term past experience in the assessment of novelty. Such resemblances between a fish and a bird suggest that we should look not only for resemblances between different groups of vertebrates in the most common overall pattern of lateralization, but also for possible resemblances in the nature of inter-individual variation and in the way in which it is generated during development.
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	Corporate Author				Thesis
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5370
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	Author	ANSELL, WFH
	Title	The breeding of some larger mammals in northern Rhodesia			Type	Journal Article
	Year	1960	Publication	Proceedings of the Zoological Society of London	Abbreviated Journal	Proc. Zool. Soc. London.
	Volume	134	Issue		Pages
	Keywords
	Abstract	Details are given of a standard manner of recording breeding data of large mammals by game department officers, which is considered a great improvement on previous casual reporting. Information on the breeding in Northern Rhodesia of several ungulates and large Carnivora is summarised, and comparisons made with published data from other territories. It is stressed that much more study is needed, but certain points emerge from consideration of present information. Many species have their young at any time of year, and only a few have a really sharply defined birth season. The wart hog (Phacochoerus aethiopicus) seems markedly seasonal in the south of its range, but not in the northern parts. There is at present no data on the fluctuations in frequency of births among non-seasonal breeders. In the seasonally breeding species the actual birth months vary somewhat from north to south, and in the wildebeest (Connochaetes taurinus) there may be variation in the calving time due apparently to purely localdifferences of environment. The wild dog (Lycaon pictus) appears to be monoestrus, but data on the larger felids, though incomplete, seems to indicate that young may be born at any time of the year. Limiting factors in the study of breeding in African large mammals are pointed out, but it is suggested that further work on the lines of that carried out in Northern. Rhodesia would be profitable.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
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	Notes	from Prof. Hans Klingels Equine Reference List			Approved	no
	Call Number	refbase @ user @			Serial	643
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