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Author Weeks, J.W.; Crowell-Davis, S.L.; Heusner, G. url  openurl
  Title Preliminary study of the development of the Flehmen response in Equus caballus Type (up) Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 78 Issue 2-4 Pages 329-335  
  Keywords Horse behavior; Flehmen; Foal development  
  Abstract The flehmen response is commonly seen in most ungulates as well as in several other species (e.g. felids). The behavior is most often thought to be part of the sexual behavioral repertoire of males. One reigning hypothesis suggests that this behavior allows the male to determine the estrous state of a female through the chemosensory functions of the vomeronasal organ. However, females and young of both sexes also exhibit this behavior. Horse foals most frequently show the flehmen response during their first month of life with colts showing the behavior more often than fillies. This study tested the flehmen response on male and female foals throughout their pre-pubertal period. Foals were separately presented estrous and non-estrous urine weekly during the first month of life and then monthly until they were approximately 7 months of age. No significant differences were found between male and female foals for the following variables: latency to flehmen, duration of flehmen, frequency of flehmen and sniffs.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 615  
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Author Fischer, J.; Hammerschmidt, K.; Cheney, D.L.; Seyfarth, R.M. openurl 
  Title Acoustic features of male baboon loud calls: influences of context, age, and individuality Type (up) Journal Article
  Year 2002 Publication The Journal of the Acoustical Society of America Abbreviated Journal J Acoust Soc Am  
  Volume 111 Issue 3 Pages 1465-1474  
  Keywords Age Factors; Animal Communication; Animals; Individuality; Male; *Papio; *Social Environment; *Sound Spectrography; *Vocalization, Animal  
  Abstract The acoustic structure of loud calls (“wahoos”) recorded from free-ranging male baboons (Papio cynocephalus ursinus) in the Moremi Game Reserve, Botswana, was examined for differences between and within contexts, using calls given in response to predators (alarm wahoos), during male contests (contest wahoos), and when a male had become separated from the group (contact wahoos). Calls were recorded from adolescent, subadult, and adult males. In addition, male alarm calls were compared with those recorded from females. Despite their superficial acoustic similarity, the analysis revealed a number of significant differences between alarm, contest, and contact wahoos. Contest wahoos are given at a much higher rate, exhibit lower frequency characteristics, have a longer “hoo” duration, and a relatively louder “hoo” portion than alarm wahoos. Contact wahoos are acoustically similar to contest wahoos, but are given at a much lower rate. Both alarm and contest wahoos also exhibit significant differences among individuals. Some of the acoustic features that vary in relation to age and sex presumably reflect differences in body size, whereas others are possibly related to male stamina and endurance. The finding that calls serving markedly different functions constitute variants of the same general call type suggests that the vocal production in nonhuman primates is evolutionarily constrained.  
  Address Department of Psychology, University of Pennsylvania, Philadelphia 19104, USA. fischer@eva.mpg.de  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 0001-4966 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:11931324 Approved no  
  Call Number refbase @ user @ Serial 691  
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Author Seyfarth, R.M.; Cheney, D.L. doi  openurl
  Title What are big brains for? Type (up) Journal Article
  Year 2002 Publication Proceedings of the National Academy of Sciences of the United States of America Abbreviated Journal Proc. Natl. Acad. Sci. U.S.A.  
  Volume 99 Issue 7 Pages 4141-4142  
  Keywords Animals; Brain/*anatomy & histology; *Intelligence; Learning; Primates/*anatomy & histology/*psychology; Social Behavior  
  Abstract  
  Address Department of Psychology, University of Pennsylvania, Philadelphia, PA 19104, USA. seyfarth@psych.upenn.edu  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0027-8424 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:11929989 Approved no  
  Call Number refbase @ user @ Serial 692  
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Author Rivera, E.; Benjamin, S.; Nielsen, B.; Shelle, J.; Zanella, A.J. url  doi
openurl 
  Title Behavioral and physiological responses of horses to initial training: the comparison between pastured versus stalled horses Type (up) Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 78 Issue 2-4 Pages 235-252  
  Keywords Horses; Housing; Training; Behavior; Stress; Cortisol; Welfare  
  Abstract Horses kept in stalls are deprived of opportunities for social interactions, and the performance of natural behaviors is limited. Inadequate environmental conditions may compromise behavioral development. Initial training is a complex process and it is likely that the responses of horses may be affected by housing conditions. Sixteen 2-year-old Arabian horses were kept on pasture (P) (n=8) or in individual stalls (S) (n=8). Twelve horses (six P and six S) were subjected to a standardized training procedure, carried out by two trainers in a round pen, and 4 horses (two P and two S) were introduced to the round pen but were not trained (C; control). On sample collection day 0, 7, 21 and 28, behavior observations were carried out, blood samples were drawn and heart rates were monitored. Total training time for the stalled horses was significantly higher than total time for the pastured horses (S: 26.4+/-1.5 min; P: 19.7+/-1.1; P=0.032). The stalled group required more time to habituate to the activities occurring from the start of training to mounting (S: 11.4+/-0.96; P: 7.3+/-0.75 min; P=0.007). Frequency of unwanted behavior was higher in the stalled horses (S: 8.0+/-2.0; P: 2.2+/-1.0; P=0.020). Pastured horses tended to have higher basal heart rates on day 0 (S: 74.7+/-4.8; P: 81.8+/-5.3 bpm; P=0.0771). While the physiological data failed to identify differences between housing groups, the behavioral data suggest that pasture-kept horses adapt more easily to training than stalled horses.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 717  
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Author Jansen, T.; Forster, P.; Levine, M.A.; Oelke, H.; Hurles, M.; Renfrew, C.; Weber, J.; Olek, K. doi  openurl
  Title Mitochondrial DNA and the origins of the domestic horse Type (up) Journal Article
  Year 2002 Publication Proceedings of the National Academy of Sciences of the United States of America Abbreviated Journal Proc. Natl. Acad. Sci. U.S.A.  
  Volume 99 Issue 16 Pages 10905-10910  
  Keywords Animals; Animals, Domestic/classification/*genetics; Base Sequence; DNA, Complementary; *DNA, Mitochondrial; *Evolution, Molecular; Horses/classification/*genetics; Molecular Sequence Data; Phylogeny  
  Abstract The place and date of the domestication of the horse has long been a matter for debate among archaeologists. To determine whether horses were domesticated from one or several ancestral horse populations, we sequenced the mitochondrial D-loop for 318 horses from 25 oriental and European breeds, including American mustangs. Adding these sequences to previously published data, the total comes to 652, the largest currently available database. From these sequences, a phylogenetic network was constructed that showed that most of the 93 different mitochondrial (mt)DNA types grouped into 17 distinct phylogenetic clusters. Several of the clusters correspond to breeds and/or geographic areas, notably cluster A2, which is specific to Przewalski's horses, cluster C1, which is distinctive for northern European ponies, and cluster D1, which is well represented in Iberian and northwest African breeds. A consideration of the horse mtDNA mutation rate together with the archaeological timeframe for domestication requires at least 77 successfully breeding mares recruited from the wild. The extensive genetic diversity of these 77 ancestral mares leads us to conclude that several distinct horse populations were involved in the domestication of the horse.  
  Address Biopsytec Analytik GmbH, Marie-Curie-Strasse 1, 53359 Rheinbach, Germany. jansen@biopsytec.com  
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  ISSN 0027-8424 ISBN Medium  
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  Notes PMID:12130666 Approved no  
  Call Number refbase @ user @ Serial 772  
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Author Christensen, J.W.; Zharkikh, T.; Ladewig, J.; Yasinetskaya, N. url  doi
openurl 
  Title Social behaviour in stallion groups (Equus przewalskii and Equus caballus) kept under natural and domestic conditions Type (up) Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 76 Issue 1 Pages 11-20  
  Keywords Domestic horse; Przewalski horse; Stallion group; Social behaviour; Equus caballus; Equus przewalskii  
  Abstract The aim of this study was to investigate social behaviour in differently reared stallions in their respective environments; one group of stallions was reared under typical domestic conditions whereas the other group was reared and lives under natural conditions. The domestic group consisted of 19, 2-year-old stallions (Equus caballus), which were all weaned at 4 months of age and experienced either individual or group housing facilities before being pastured with the other similarly aged stallions. The natural living and mixed age group of Przewalski stallions (E. przewalskii) consisted of 13 stallions, most of which were juveniles (n=11, <=4 years; n=2, >9 years). The domestic group was studied in a 4-ha enclosure at the Danish Institute of Agricultural Sciences and the Przewalski group under free-ranging conditions in a 75-ha enclosure in the Askania Nova Biosphere Reserve, Ukraine. Behavioural data was collected during 168 h of direct observation. The occurrence of 14 types of social interactions was recorded and group spacing behaviour was studied using nearest neighbour recordings. In spite of very different environments, reflecting domestic and natural rearing conditions, many similarities in behaviour was found. Play and play fight behaviour was very similar in the two stallion groups. Quantitative differences were found in social grooming since Przewalski stallions groomed more frequently (P=0.004), and in investigative behaviours, since domestic stallions showed more nasal (P=0.005) and body sniffing (P<0.001), whereas Przewalski stallions directed more sniffing towards the genital region (P<0.001). These differences may, however, be attributed to environmental factors and in the period of time the stallions were together prior to the study period. Quantitative differences appeared in some agonistic behaviours (kick threat, P<0.001; and kick, P<0.001), but data do not support earlier findings of Przewalski horses being significantly more aggressive than domestic horses. In general, Przewalski stallions engaged in more social interactions, and they showed less group spacing, i.e. maintained a significantly shorter distance between neighbours (P<0.001). The study indicates that also domestic horses, which have been reared under typical domestic conditions and allowed a period on pasture, show social behaviour, which is very similar to that shown by their non-domestic relatives.  
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  Call Number refbase @ user @ Serial 776  
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Author Hemelrijk, C.K. doi  openurl
  Title Despotic societies, sexual attraction and the emergence of male 'tolerance': an agent-based model Type (up) Journal Article
  Year 2002 Publication Behaviour Abbreviated Journal Behaviour  
  Volume 139 Issue 6 Pages 729-747  
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  Abstract During the period when females are sexually attractive – but only then – males of certain species of primates, such as chimpanzees, allow females access to resources. Because males are usually dominant over females, such male 'tolerance' is explained as a special, reproductive strategy to gain access to females. In this paper a simpler hypothesis is proposed on the basis of an individual-based model (called DomWorld): male 'tolerance' towards females arises in 'despotic' artificial societies as a kind of 'respectful timidity', because sexual attraction automatically increases female dominance over males as a side-effect. The model consists in a homogeneous, virtual world with agents that group and perform dominance-interactions in which the effects of victory and defeat are self-reinforcing. The artificial sexes differ in that VirtualMales have a higher intensity of aggression, they start with a greater capacity to win conflicts than VirtualFemales and they are especially attracted to the opposite sex during certain periods, whereas VirtualFemales are not. I shall explain how the introduction into DomWorld of the attraction of VirtualMales by VirtualFemales leads to female dominance, why it does so only in despotic, but not in egalitarian societies, and how it leads to other phenomena that are relevant to the study of primate behaviour.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 864  
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Author Witte, K.; Ryan, M.J. url  openurl
  Title Mate choice copying in the sailfin molly, Poecilia latipinna, in the wild Type (up) Journal Article
  Year 2002 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 63 Issue 5 Pages 943-949  
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  Notes Approved no  
  Call Number Serial 1809  
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Author Powers, P.; Harrison, A. openurl 
  Title Effects of the rider on the linear kinematics of jumping horses Type (up) Journal Article
  Year 2002 Publication Sports Biomechanics / International Society of Biomechanics in Sports Abbreviated Journal Sports Biomech  
  Volume 1 Issue 2 Pages 135-146  
  Keywords Animals; Behavior, Animal/*physiology; Biomechanics; Communication; Exertion/*physiology; Gait/*physiology; Horses/*physiology; Humans; Locomotion/*physiology; Posture/*physiology; Task Performance and Analysis; Video Recording; Weight-Bearing/*physiology  
  Abstract This study examined the effects of the rider on the linear projectile kinematics of show-jumping horses. SVHS video recordings (50 Hz) of eight horses jumping a vertical fence 1 m high were used for the study. Horses jumped the fence under two conditions: loose (no rider or tack) and ridden. Recordings were digitised using Peak Motus. After digitising the sequences, each rider's digitised data were removed from the ridden horse data so that three conditions were examined: loose, ridden (including the rider's data) and riderless (rider's data removed). Repeated measures ANOVA revealed significant differences between ridden and loose conditions for CG height at take-off (p < 0.001), CG distance to the fence at take-off (p = 0.001), maximum CG during the suspension phase (p < 0.001), CG position over the centre of the fence (p < 0.001), CG height at landing (p < 0.001), and vertical velocity at take-off (p < 0.001). The results indicated that the rider's effect on jumping horses was primarily due to behavioural changes in the horses motion (resulting from the rider's instruction), rather than inertial effects (due to the positioning of the rider on the horse). These findings have implications for the coaching of riders and horses.  
  Address Department of PE and Sports Sciences, University of Limerick, Limerick, Ireland  
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  ISSN 1476-3141 ISBN Medium  
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  Notes PMID:14658371 Approved no  
  Call Number Serial 1904  
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Author Goodwin, D.; Davidson, H.P.B.; Harris, P. openurl 
  Title Foraging enrichment for stabled horses: effects on behaviour and selection Type (up) Journal Article
  Year 2002 Publication Equine Veterinary Journal Abbreviated Journal Equine Vet J  
  Volume 34 Issue 7 Pages 686-691  
  Keywords Animal Feed; Animal Husbandry/*methods; Animal Welfare; Animals; *Behavior, Animal; Female; Horse Diseases/*psychology; Horses/*physiology; Housing, Animal; Male; *Poaceae; *Stereotyped Behavior; Videotape Recording  
  Abstract The restricted access to pasture experienced by many competition horses has been linked to the exhibition of stereotypic and redirected behaviour patterns. It has been suggested that racehorses provided with more than one source of forage are less likely to perform these patterns; however, the reasons for this are currently unclear. To investigate this in 4 replicated trials, up to 12 horses were introduced into each of 2 identical stables containing a single forage, or 6 forages for 5 min. To detect novelty effects, in the first and third trials the single forage was hay. In the second and fourth, it was the preferred forage from the preceding trial. Trials were videotaped and 12 mutually exclusive behaviour patterns compared. When hay was presented as the single forage (Trials 1 and 3), all recorded behaviour patterns were significantly different between stables; e.g. during Trial 3 in the 'Single' stable, horses looked over the stable door more frequently (P<0.001), moved for longer (P<0.001), foraged on straw bedding longer (P<0.001), and exhibited behaviour indicative of motivation to search for alternative resources (P<0.001) more frequently. When a previously preferred forage was presented as the single forage (Trials 2 and 4) behaviour was also significantly different between stables, e.g in Trial 4 horses looked out over the stable door more frequently (P<0.005) and foraged for longer in their straw bedding (P<0.005). Further study is required to determine whether these effects persist over longer periods. However, these trials indicate that enrichment of the stable environment through provision of multiple forages may have welfare benefits for horses, in reducing straw consumption and facilitating the expression of highly motivated foraging behaviour.  
  Address Anthrozoology Institute, University of Southampton, UK  
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  ISSN 0425-1644 ISBN Medium  
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  Notes PMID:12455839 Approved no  
  Call Number refbase @ user @ Serial 1910  
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