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Author Rivera, E.; Benjamin, S.; Nielsen, B.; Shelle, J.; Zanella, A.J. url  doi
openurl 
  Title Behavioral and physiological responses of horses to initial training: the comparison between pastured versus stalled horses Type Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 78 Issue 2-4 Pages 235-252  
  Keywords Horses; Housing; Training; Behavior; Stress; Cortisol; Welfare  
  Abstract Horses kept in stalls are deprived of opportunities for social interactions, and the performance of natural behaviors is limited. Inadequate environmental conditions may compromise behavioral development. Initial training is a complex process and it is likely that the responses of horses may be affected by housing conditions. Sixteen 2-year-old Arabian horses were kept on pasture (P) (n=8) or in individual stalls (S) (n=8). Twelve horses (six P and six S) were subjected to a standardized training procedure, carried out by two trainers in a round pen, and 4 horses (two P and two S) were introduced to the round pen but were not trained (C; control). On sample collection day 0, 7, 21 and 28, behavior observations were carried out, blood samples were drawn and heart rates were monitored. Total training time for the stalled horses was significantly higher than total time for the pastured horses (S: 26.4+/-1.5 min; P: 19.7+/-1.1; P=0.032). The stalled group required more time to habituate to the activities occurring from the start of training to mounting (S: 11.4+/-0.96; P: 7.3+/-0.75 min; P=0.007). Frequency of unwanted behavior was higher in the stalled horses (S: 8.0+/-2.0; P: 2.2+/-1.0; P=0.020). Pastured horses tended to have higher basal heart rates on day 0 (S: 74.7+/-4.8; P: 81.8+/-5.3 bpm; P=0.0771). While the physiological data failed to identify differences between housing groups, the behavioral data suggest that pasture-kept horses adapt more easily to training than stalled horses.  
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  Call Number (up) refbase @ user @ Serial 717  
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Author Byrne, R.W. url  openurl
  Title Imitation of novel complex actions: What does the evidence from animals mean? Type Book Chapter
  Year 2002 Publication Advances in the Study of Behavior Abbreviated Journal Adv Stud Behav  
  Volume 31 Issue Pages 77-105  
  Keywords  
  Abstract Summary Underlying the various behaviors that are classified as imitation, there may be several distinct mechanisms, differing in adaptive function, cognitive basis, and computational power. Experiments reporting “true motor imitation” in animals do not as yet give evidence of production learning by imitation; instead, contextual imitation can explain their data, and this can be explained by a simple mechanism (response facilitation) which matches known neural findings. When imitation serves a function in social mimicry, which applies to a wide range of phenomena from neonatal imitation in humans and great apes to pair-bonding in some bird species, the fidelity of the behavioral match is crucial. Learning of novel behavior can potentially be achieved by matching the outcome of a model's action, and it is argued that vocal imitation by birds is a clear example of this method (which is sometimes called emulation). Alternatively, the behavior itself may be perceived in terms of actions that the observer can perform, and thus it may be copied. If the imitation is linear and stringlike (action level), following the surface form rather than the underlying plan, then its utility for learning new instrumental methods is limited. However, the underlying plan of hierarchically organized behavior is visible in output behavior, in subtle but detectable ways, and imitation could instead be based on this organization (program level), extracted automatically by string parsing. Currently, the most likely candidates for such capacities are all great apes. It is argued that this ability to perceive the underlying plan of action, in addition to allowing highly flexible imitation of novel instrumental methods, may have resulted in the competence to understand the intentions (theory of mind) of others.  
  Address  
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  Publisher Academic Press Place of Publication San Diego Editor Snowdon, C. T.; Roper, T. J.;Rosenblatt,J. S.  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 746  
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Author Dyer, F.C. doi  openurl
  Title Animal behaviour: when it pays to waggle Type
  Year 2002 Publication Nature Abbreviated Journal Nature  
  Volume 419 Issue 6910 Pages 885-886  
  Keywords *Animal Communication; Animals; Bees/*physiology; California; Dancing/physiology; Environment; Evolution; Female; Flowers/chemistry; *Food; Gravitation; Lighting; Motor Activity/*physiology; Odors; Seasons; Sunlight  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0028-0836 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:12410290 Approved no  
  Call Number (up) refbase @ user @ Serial 769  
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Author Jansen, T.; Forster, P.; Levine, M.A.; Oelke, H.; Hurles, M.; Renfrew, C.; Weber, J.; Olek, K. doi  openurl
  Title Mitochondrial DNA and the origins of the domestic horse Type Journal Article
  Year 2002 Publication Proceedings of the National Academy of Sciences of the United States of America Abbreviated Journal Proc. Natl. Acad. Sci. U.S.A.  
  Volume 99 Issue 16 Pages 10905-10910  
  Keywords Animals; Animals, Domestic/classification/*genetics; Base Sequence; DNA, Complementary; *DNA, Mitochondrial; *Evolution, Molecular; Horses/classification/*genetics; Molecular Sequence Data; Phylogeny  
  Abstract The place and date of the domestication of the horse has long been a matter for debate among archaeologists. To determine whether horses were domesticated from one or several ancestral horse populations, we sequenced the mitochondrial D-loop for 318 horses from 25 oriental and European breeds, including American mustangs. Adding these sequences to previously published data, the total comes to 652, the largest currently available database. From these sequences, a phylogenetic network was constructed that showed that most of the 93 different mitochondrial (mt)DNA types grouped into 17 distinct phylogenetic clusters. Several of the clusters correspond to breeds and/or geographic areas, notably cluster A2, which is specific to Przewalski's horses, cluster C1, which is distinctive for northern European ponies, and cluster D1, which is well represented in Iberian and northwest African breeds. A consideration of the horse mtDNA mutation rate together with the archaeological timeframe for domestication requires at least 77 successfully breeding mares recruited from the wild. The extensive genetic diversity of these 77 ancestral mares leads us to conclude that several distinct horse populations were involved in the domestication of the horse.  
  Address Biopsytec Analytik GmbH, Marie-Curie-Strasse 1, 53359 Rheinbach, Germany. jansen@biopsytec.com  
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  Series Volume Series Issue Edition  
  ISSN 0027-8424 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:12130666 Approved no  
  Call Number (up) refbase @ user @ Serial 772  
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Author Christensen, J.W.; Zharkikh, T.; Ladewig, J.; Yasinetskaya, N. url  doi
openurl 
  Title Social behaviour in stallion groups (Equus przewalskii and Equus caballus) kept under natural and domestic conditions Type Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 76 Issue 1 Pages 11-20  
  Keywords Domestic horse; Przewalski horse; Stallion group; Social behaviour; Equus caballus; Equus przewalskii  
  Abstract The aim of this study was to investigate social behaviour in differently reared stallions in their respective environments; one group of stallions was reared under typical domestic conditions whereas the other group was reared and lives under natural conditions. The domestic group consisted of 19, 2-year-old stallions (Equus caballus), which were all weaned at 4 months of age and experienced either individual or group housing facilities before being pastured with the other similarly aged stallions. The natural living and mixed age group of Przewalski stallions (E. przewalskii) consisted of 13 stallions, most of which were juveniles (n=11, <=4 years; n=2, >9 years). The domestic group was studied in a 4-ha enclosure at the Danish Institute of Agricultural Sciences and the Przewalski group under free-ranging conditions in a 75-ha enclosure in the Askania Nova Biosphere Reserve, Ukraine. Behavioural data was collected during 168 h of direct observation. The occurrence of 14 types of social interactions was recorded and group spacing behaviour was studied using nearest neighbour recordings. In spite of very different environments, reflecting domestic and natural rearing conditions, many similarities in behaviour was found. Play and play fight behaviour was very similar in the two stallion groups. Quantitative differences were found in social grooming since Przewalski stallions groomed more frequently (P=0.004), and in investigative behaviours, since domestic stallions showed more nasal (P=0.005) and body sniffing (P<0.001), whereas Przewalski stallions directed more sniffing towards the genital region (P<0.001). These differences may, however, be attributed to environmental factors and in the period of time the stallions were together prior to the study period. Quantitative differences appeared in some agonistic behaviours (kick threat, P<0.001; and kick, P<0.001), but data do not support earlier findings of Przewalski horses being significantly more aggressive than domestic horses. In general, Przewalski stallions engaged in more social interactions, and they showed less group spacing, i.e. maintained a significantly shorter distance between neighbours (P<0.001). The study indicates that also domestic horses, which have been reared under typical domestic conditions and allowed a period on pasture, show social behaviour, which is very similar to that shown by their non-domestic relatives.  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 776  
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Author Hemelrijk, C.K. doi  openurl
  Title Despotic societies, sexual attraction and the emergence of male 'tolerance': an agent-based model Type Journal Article
  Year 2002 Publication Behaviour Abbreviated Journal Behaviour  
  Volume 139 Issue 6 Pages 729-747  
  Keywords  
  Abstract During the period when females are sexually attractive – but only then – males of certain species of primates, such as chimpanzees, allow females access to resources. Because males are usually dominant over females, such male 'tolerance' is explained as a special, reproductive strategy to gain access to females. In this paper a simpler hypothesis is proposed on the basis of an individual-based model (called DomWorld): male 'tolerance' towards females arises in 'despotic' artificial societies as a kind of 'respectful timidity', because sexual attraction automatically increases female dominance over males as a side-effect. The model consists in a homogeneous, virtual world with agents that group and perform dominance-interactions in which the effects of victory and defeat are self-reinforcing. The artificial sexes differ in that VirtualMales have a higher intensity of aggression, they start with a greater capacity to win conflicts than VirtualFemales and they are especially attracted to the opposite sex during certain periods, whereas VirtualFemales are not. I shall explain how the introduction into DomWorld of the attraction of VirtualMales by VirtualFemales leads to female dominance, why it does so only in despotic, but not in egalitarian societies, and how it leads to other phenomena that are relevant to the study of primate behaviour.  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 864  
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Author Goodwin, D.; Davidson, H.P.B.; Harris, P. openurl 
  Title Foraging enrichment for stabled horses: effects on behaviour and selection Type Journal Article
  Year 2002 Publication Equine Veterinary Journal Abbreviated Journal Equine Vet J  
  Volume 34 Issue 7 Pages 686-691  
  Keywords Animal Feed; Animal Husbandry/*methods; Animal Welfare; Animals; *Behavior, Animal; Female; Horse Diseases/*psychology; Horses/*physiology; Housing, Animal; Male; *Poaceae; *Stereotyped Behavior; Videotape Recording  
  Abstract The restricted access to pasture experienced by many competition horses has been linked to the exhibition of stereotypic and redirected behaviour patterns. It has been suggested that racehorses provided with more than one source of forage are less likely to perform these patterns; however, the reasons for this are currently unclear. To investigate this in 4 replicated trials, up to 12 horses were introduced into each of 2 identical stables containing a single forage, or 6 forages for 5 min. To detect novelty effects, in the first and third trials the single forage was hay. In the second and fourth, it was the preferred forage from the preceding trial. Trials were videotaped and 12 mutually exclusive behaviour patterns compared. When hay was presented as the single forage (Trials 1 and 3), all recorded behaviour patterns were significantly different between stables; e.g. during Trial 3 in the 'Single' stable, horses looked over the stable door more frequently (P<0.001), moved for longer (P<0.001), foraged on straw bedding longer (P<0.001), and exhibited behaviour indicative of motivation to search for alternative resources (P<0.001) more frequently. When a previously preferred forage was presented as the single forage (Trials 2 and 4) behaviour was also significantly different between stables, e.g in Trial 4 horses looked out over the stable door more frequently (P<0.005) and foraged for longer in their straw bedding (P<0.005). Further study is required to determine whether these effects persist over longer periods. However, these trials indicate that enrichment of the stable environment through provision of multiple forages may have welfare benefits for horses, in reducing straw consumption and facilitating the expression of highly motivated foraging behaviour.  
  Address Anthrozoology Institute, University of Southampton, UK  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 0425-1644 ISBN Medium  
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  Notes PMID:12455839 Approved no  
  Call Number (up) refbase @ user @ Serial 1910  
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Author Isenbugel, E. openurl 
  Title [From wild horse to riding horse] Type Journal Article
  Year 2002 Publication Schweizer Archiv fur Tierheilkunde Abbreviated Journal Schweiz Arch Tierheilkd  
  Volume 144 Issue 7 Pages 323-329  
  Keywords Animal Husbandry/*history; Animals; Animals, Domestic; Animals, Wild; *Bonding, Human-Pet; Breeding/history; Evolution; Female; History, 15th Century; History, 16th Century; History, 17th Century; History, 18th Century; History, 19th Century; History, 20th Century; History, Ancient; History, Medieval; *Horses/physiology/psychology; Humans; Male; Paintings; Predatory Behavior; Sculpture; Sports/history  
  Abstract Over 45 million years of evolution the horse developed to a highly specialized animal in anatomy, physiology and behavior. No other animal had influenced the economic and cultural history of men to such extent. Hunting prey since the ice age, domesticated 4000 B.C. and used for thousands of years as unique animal all over the world has attained a new role today as partner in sport, as companion animal and even as cotherapeutic. The well known behavioral demands in use and keeping are still often not fulfilled.  
  Address Zoologischer Garten Zurich  
  Corporate Author Thesis  
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  Language German Summary Language Original Title Vom Wildpferd zum Reitpferd  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0036-7281 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:12174680 Approved no  
  Call Number (up) refbase @ user @ Serial 1913  
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Author Pollmann, U. openurl 
  Title [Keeping of horses in circus and show businesses] Type Journal Article
  Year 2002 Publication DTW. Deutsche Tierarztliche Wochenschrift Abbreviated Journal Dtsch Tierarztl Wochenschr  
  Volume 109 Issue 3 Pages 126-129  
  Keywords Animal Husbandry/*methods; *Animal Welfare; Animals; Behavior, Animal; *Horses; *Housing, Animal; Humans; Reward  
  Abstract The conditions under which horses are kept and the performance of acts in the circus ring may give rise to animal protection-relevant aspects for circus and show horses. A number of intolerable conditions under which horses are kept and procedures adopted for the work with circus and show horses are described. In addition, attention is drawn to monitoring methods capable of exposing the deplorable shortcomings of these businesses.  
  Address Fachbereich Ethologie und Tierschutz des Chemischen und Veterinaruntersuchungsamtes Freiburg. Ursula.Pollmann@cvuafr.bwl.de  
  Corporate Author Thesis  
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  Language German Summary Language Original Title Pferdehaltung in Zirkus- und Schaustellerbetrieben  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0341-6593 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:11963363 Approved no  
  Call Number (up) refbase @ user @ Serial 1914  
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Author Houpt, K.A.; Rudman, R. url  doi
openurl 
  Title Foreword to special issue on equine behavior Type Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 78 Issue 2-4 Pages 83-85  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 1986  
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