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Sondergaard, E.; Ladewig, J. |
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Title |
Group housing exerts a positive effect on the behaviour of young horses during training |
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2004 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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87 |
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1-2 |
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105-118 |
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Horses; Housing; Human-animal relationship; Social environment; Training; Learning |
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In an experiment on the effects of social environment and training on the human-animal relationship, 20 horses were handled according to a defined schedule. Eight horses were housed singly and 12 horses were housed in four groups of 3 horses. Horses were handled three times per week in 10 min sessions from an age of 6 months until 2 years of age during two winter periods. A total of 50 and 70 sessions were given in the first and second period, respectively. Five randomly allocated people performed the training. The training scheme involved leading, tying up, touching, lifting feet, etc. in 43 stages. The horse had to fulfil the performance criteria of each stage in order to get to the next stage. In the first winter period, horses were led to the stable when they had “passed” a stage or after 10 min of training. In the second winter period, horses would start off at stage 1 again, and when they “passed” a stage they went on to the next stage within the same training session. Because of the change in training procedure results were analysed separately for the two winter periods. There was a significant difference between trainers in the number of times they allowed a horse to “pass” a stage within each winter period (χ32, P<0.05; χ32, P<0.001 for the first and the second winter period, respectively). Group housed horses “passed” more stages than single housed horses (17 versus 14; 27 versus 18 in the first and second winter period, respectively; P<0.05 for the interaction). Singly housed horses bit the trainer more frequently than did group housed horses (P<0.01). The responses of group housed horses to training clearly demonstrate the benefits of raising young horses in groups. |
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McDonnell, S.M.; Haviland, J.C.S. |
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Title |
Agonistic ethogram of the equid bachelor band |
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1995 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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43 |
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3 |
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147-188 |
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Equid; Horse; Bachelor; Agonistic; Social behavior; Ethogram |
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An ethogram of agonistic and related behaviors among equid bachelor band members was developed. Several key English-language studies on equids were reviewed to derive a preliminary inventory of specific behaviors to be included in the ethogram. A bachelor band of domestic pony stallions pastured together was observed for approximately 50 daylight hours to obtain detailed descriptions of each behavior, enable photographic and video documentation of behaviors, and identify any behaviors to be added to the preliminary inventory. An initial draft of the ethogram was sent to 65 equine researchers for review. Twenty-eight critical reviews were received and their suggestions considered for the final draft. A total of 49 elemental behaviors including five distinct vocalizations was included in the ethogram. Three complex behavioral sequences were also included. Most of the behaviors catalogued from the direct observation of pastured pony stallions were also found in the equid literature. For many, references to these behaviors specifically among males or bachelor band members were not found. The results offer a practical tool for quantitative research and other studies of equid inter-male behavior as well as for teaching of equid behavior, and should facilitate progress toward development of a complete ethogram for the horse and other equids. |
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refbase @ user @ ; Equine Behaviour @ team @ room B 3.029 |
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749 |
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van Dierendonck, M.C.; Bandi, N.; Batdorj, D.; Dugerlham, S.; Munkhtsog, B. |
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Behavioural observations of reintroduced Takhi or Przewalski horses (Equus ferus przewalskii) in Mongolia |
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1996 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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50 |
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2 |
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95-114 |
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Horse; Przewalski's horse; Reintroduction; Time budget; Social organisation; Basic reference dataset |
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During 1992 and 1993, 14 reintroduced Przewalski Horses or Takhi (Equus ferus przewalskii) were studied in the Hustain Nuruu Mountain Steppe reserve in Mongolia. Most of the individuals did not know each other before reintroduction. These Takhi were the first of five groups due to be released in the reserve after an acclimatisation period of at least 1 year. During acclimatisation the Takhi, lived visually and acoustically separately, in fenced enclosures of approximately 45 ha each. The observations, mostly scan-sampling, were carried out in each season. The observation bouts were divided over six periods and over two harem herds. Two of the periods were in the same consecutive seasons, so comparison over the years was possible. Social integration within the Takhi herds was very high from the beginning, as described by the spatial relation and synchronisation data. Between 50 and 89% of the observation time, the behaviour of all herd members was synchronised. The amount of time spent grazing by the Takhi (30-68% of the daylight period) was similar to that of feral horses and Takhi in captivity and semi-reserves. The Takhi tended to rest in the morning and have a bimodal period of grazing at dawn and in the afternoon. The Takhi displayed clear habitat preferences for certain activities. They had a strong preference to rest at the highest point in their enclosure. They fed preferably on two or three different vegetation types (with five types available in each enclosure). The amount of time spent grazing during the non-growing seasons (49 +/- 15%) indicates that the feeding value and availability of food were sufficient. Health changes were detected adequately using condition scoring sheets. No supplementary food or water was supplied during the harsh winters. Moreover, low mortality rates and high reproductive success show that the mountain steppe is a habitat which is potentially suitable for establishing a healthy Takhi population. Takhi is the first species to return to its native habitat after living only in zoos for so many generations. |
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762 |
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Feh, C. |
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Social behaviour and relationships of Prezewalski horses in Dutch semi-reserves |
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1988 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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71-87 |
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A short-term study was made of 2 groups of Przewalski horses, a bachelor group of 4 juvenile stallions in Ooij Polder and a harem group of 1 stallion and 4 mares. All social interactions were recorded and the nearest and farthest neighbour was noted. Correspondence analysis was used to determine what parameters determined the relationships among the horses. There was a linear hierarchy among the bachelor stallions. The dominant stallion of the group was also the oldest. The hierarchy was not linear in the harem group, and the 3-year-old stallion was subordinate to the 5-year-old mares. He was also most likely to be farthest from other horses. The mares of the same age, who had also arrived in the park at the same time, tended to be one another's nearest neighbours. The frequency of aggression is higher among Przewalski horses than among domestic horses of similar ages. Correspondence analysis revealed that head-threats and other forms of aggression accounted for more of the variance in the data than any other behaviour, but submission, play and social interactions also contributed. |
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764 |
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Keiper, R.R. |
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Social interactions of the Przewalski horse (Equus przewalskii Poliakov, 1881) herd at the Munich Zoo |
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1988 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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89-97 |
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Data were collected on 972 aggressions and 233 acts of mutual grooming in a herd of 9 Przewalski horses in the Hellabrunn Tierpark in Munich, West Germany. The herd was composed of 1 adult stallion, 5 adult mares and 3 foals. A distinct linear dominance hierarchy was present in the herd, with the stallion being the top-ranking animal. Age was significantly correlated with rank. Almost 40% of all aggressions consisted of herding actions by the stallion. Threats to bite (20% of all aggressions) and threats to kick (11.4%) were next in frequency of occurrence. Most mutual grooms (71%) involved grooming the front part of the body. Although mutual grooming may be used to appease higher-ranked animals, most grooming bouts were between related horses. Foals initiate 47.6% of all allogrooming. Mutual grooming may reduce weaning conflict between a mare and her foal or may result in female coalitions that defend against predators or aggression by the herd stallion. |
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775 |
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Christensen, J.W.; Zharkikh, T.; Ladewig, J.; Yasinetskaya, N. |
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Social behaviour in stallion groups (Equus przewalskii and Equus caballus) kept under natural and domestic conditions |
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2002 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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76 |
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1 |
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11-20 |
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Domestic horse; Przewalski horse; Stallion group; Social behaviour; Equus caballus; Equus przewalskii |
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The aim of this study was to investigate social behaviour in differently reared stallions in their respective environments; one group of stallions was reared under typical domestic conditions whereas the other group was reared and lives under natural conditions. The domestic group consisted of 19, 2-year-old stallions (Equus caballus), which were all weaned at 4 months of age and experienced either individual or group housing facilities before being pastured with the other similarly aged stallions. The natural living and mixed age group of Przewalski stallions (E. przewalskii) consisted of 13 stallions, most of which were juveniles (n=11, <=4 years; n=2, >9 years). The domestic group was studied in a 4-ha enclosure at the Danish Institute of Agricultural Sciences and the Przewalski group under free-ranging conditions in a 75-ha enclosure in the Askania Nova Biosphere Reserve, Ukraine. Behavioural data was collected during 168 h of direct observation. The occurrence of 14 types of social interactions was recorded and group spacing behaviour was studied using nearest neighbour recordings. In spite of very different environments, reflecting domestic and natural rearing conditions, many similarities in behaviour was found. Play and play fight behaviour was very similar in the two stallion groups. Quantitative differences were found in social grooming since Przewalski stallions groomed more frequently (P=0.004), and in investigative behaviours, since domestic stallions showed more nasal (P=0.005) and body sniffing (P<0.001), whereas Przewalski stallions directed more sniffing towards the genital region (P<0.001). These differences may, however, be attributed to environmental factors and in the period of time the stallions were together prior to the study period. Quantitative differences appeared in some agonistic behaviours (kick threat, P<0.001; and kick, P<0.001), but data do not support earlier findings of Przewalski horses being significantly more aggressive than domestic horses. In general, Przewalski stallions engaged in more social interactions, and they showed less group spacing, i.e. maintained a significantly shorter distance between neighbours (P<0.001). The study indicates that also domestic horses, which have been reared under typical domestic conditions and allowed a period on pasture, show social behaviour, which is very similar to that shown by their non-domestic relatives. |
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776 |
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Keiper, R.; Receveur, H. |
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Social interactions of free-ranging Przewalski horses in semi-reserves in the Netherlands |
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1992 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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33 |
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4 |
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303-318 |
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Social interactions were recorded in two bands of free-ranging Przewalski horses living on large (greater than 30 ha) pastures in the Netherlands. The average number of aggressive interactions per hour was 8.86 at Lelystad and 10.36 at Noorderheide. The most common aggressive interactions were lower intensity, lower cost displacements (17.2% of all aggressive acts at Lelystad, 13.2% at Noorderheide), threats to bite (42.3% and 40.7%, respectively) and threats to kick (15.4% and 23.9%, respectively). Analysis of aggression revealed that a clear, linear dominance hierarchy was present in each band. For each band there was a positive and highly significant correlation between the age of a horse and its rank in the hierarchy. In each band, the stallion was not the highest ranked horse. Non-agonistic behaviors exceeded the number of agonistic interactions (1253 vs. 558 for Lelystad; 1257 vs. 995 at Noorderheide). There was a negative correlation between the rank of a horse in the dominance hierarchy and the number of non-agonistic behaviors displayed. The group displaying the highest number of non-agonistic interactions were foals (48.9% of total non-agonistic behaviors at Lelystad; 51.1% at Noorderheide). The non-agonistic/agonistic ratio was greater than 1 for yearlings and the band stallion, as was also the case for foals, but was less than 1 for males. |
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778 |
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Hogan, E.S.; Houpt, K.A.; Sweeney, K. |
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The effect of enclosure size on social interactions and daily activity patterns of the captive Asiatic wild horse (Equus przewalskii) |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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147-168 |
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Two herds of Przewalski horses at the Minnesota Zoological Garden were observed during 1980 in each of 2 enclosures that differed in size. The larger enclosure was a 3.4-ha pasture; the smaller enclosure was a 17 x 30-m grass-less pen. One herd was composed of a stallion, 3 adult mares and 2 foals. The other consisted of a stallion and 2 mares. All occurrences of aggression, mutual grooming and snapping were recorded, and 5-min scan-samples of the activity state of each horse were taken. The time budgets, frequency of aggression and frequency of mutual grooming differed significantly with enclosure size for both herds. More time was spent pacing and milling in the smaller enclosure, and the frequency of aggressions and of mutual grooming was also higher. Only the foals exhibited snapping; frequency of snapping did not vary with enclosure size. More time was spent feeding in the larger enclosure. Provision of hay in the smaller enclosure eliminated the differences in time spent feeding. A second study was conducted during the spring of 1984 in an intermediate-sized enclosure, 0.4 ha, a sub-division of the pasture on which the horses were kept in 1980. One herd consisted of a stallion, 2 mares and 2 yearlings; the other consisted of a stallion, 3 mares and a foal. One of the stallions and all of the mares were those studied in 1980, but that stallion and one of the mares were in different herds than they had been in 1980. The frequency of aggression was similar to that observed in 1980. |
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Tilson, R.L.; Sweeny, K.A.; Binczik, G.A.; Reindl, N.J. |
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Buddies and bullies: Social structure of a bachelor group of Przewalski horses |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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169-185 |
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The Minnesota Zoological Garden established a bachelor (i.e. all-male) group of 8 Przewalski horses (Equus przewalskii) in 1984. Possible methods for mixing males emerged through preliminary introductions and review of the relevant literature. These included: (1) grouping colts (<=3 years old) after removing them from their natal bands; (2) introducing adult stallions to colts in spacious, controlled environments to reduce the risk of combat-related injury or death associated with largely unpredictable adult temperament; (3) introducing more than 1 animal at a time to established groups. Based on the frequency and orientation of agonistic interactions observed among the bachelors, a linear dominance hierarchy was discerned among the 4 adult stallions; all 4 colts were determined to be of equally low or seemingly non-ranking status. Social rank among the adults was most influenced by individual temperament, and contention for rank was limited almost exclusively to the 3 lower-ranking stallions; the alpha male participated in 73% of all agonistic interactions observed, but was identified as the recipient in less than 3% of these interactions. Intra-band associations were detected in the bachelor group. The most prevalent of these was among the 4 colts, found together in over 70% of the observations. Other affinities commonly observed were between the colts (as a sub-group) and different stallions. The dominant stallion, the colts' most frequent companion, actively discouraged the 3 lower-ranking stallions from associating with the colts. Further, the colts were subjected to significantly fewer agonistic behaviors than the non-alpha stallions (per colt=1.05+/-0.24 h-1; per non-alpha stallion=2.01+/-0.04 h-1). These findings support the hypothesis developed by other workers for ungulates that young bachelors are sheltered from aggression by association with an alpha male. With one exception, adult associations were rarely noted. Although combat-related injuries (superficial wounds and temporary lameness) were common among the bachelors, few required medical attention. No permanent disabilities or deaths have occurred since the group's inception. The existence of social organization and intra-band associations in the bachelor group support the suggestion that such groups were once a natural part of this species' social milieu. Bachelor groups are therefore recommended as an ethologically sound, space- and cost-effective means of managing “surplus” (i.e. non-harem) male Przewalski horses. |
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Araba, B.D.; Crowell-Davis, S.L. |
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Dominance relationships and aggression of foals (Equus caballus) |
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1994 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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41 |
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1-2 |
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1-25 |
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aggression; dominance; horse; Equus caballus |
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Studied a herd of 15 Belgian brood-mares and 10 foals. Specific aspects of social structure studied were dominance-subordinance relationships, preferred associates, social spacing, aggression rates, the frequency of aggressions administered down the dominance hierarchy, and interactive play bouts. The rank order of the foals, both before and after weaning, was positively correlated with the rank order of their dams. There was also a significant relationship between a foal's rank and its total aggression or aggression rate per subordinate post-weaning. Higher ranking foals had higher rates of aggression. Over 80% of threats were directed down the dominance hierachy. The play-rank order of the foals, scored by the number of times foal left a play bout, was not significantly correlated with the rank order as scored by agonistic interactions. -from Authors |
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Dept Anatomy and Radiology, Univ of Georgia, Athens, GA, USA |
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ISSN |
01681591 (Issn) |
ISBN |
|
Medium |
|
|
| |
Area |
|
Expedition |
|
Conference |
|
|
| |
Notes |
|
Approved |
no |
|
| |
Call Number |
refbase @ user @ |
Serial |
790 |
|
| Permanent link to this record |
