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	Author	Witter, M.S.; Swaddle, J.P.
	Title	Fluctuating Asymmetries, Competition and Dominance			Type	Journal Article
	Year	1994	Publication	Proceedings of the Royal Society B: Biological Sciences	Abbreviated Journal	Proc. Roy. Soc. Lond. B Biol. Sci.
	Volume	256	Issue	1347	Pages	299-303
	Keywords
	Abstract	Levels of fluctuating asymmetry (FA) in the primary feathers of European starlings, Sturnus vulgaris, have been shown to be sensitive to nutritional and energetic stress. Furthermore, between-individual variation in plumage FA has been found to be related to social dominance, even without social interactions during feather growth, with dominant birds exhibiting the highest levels of FA. Here we examine whether the relation between dominance and FA differs when birds are housed in social groups, under different degrees of competition for food, during moult. We reason that dominants should derive a greater benefit from their social status as competition for food increases. Our results support this proposition. The relation between dominance and FA differed significantly according to the degree of competition for food. However, in no cases did the dominants exhibit lower levels of FA than subdominants. When competition for food was low, dominants had higher levels of FA than subdominants. When competition for food was high, there was no systematic relation between dominance and FA. These results suggest that dominants may only derive a net benefit from their social status, under the circumstances of our experiment, during severe conditions of competition.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2203
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	Author	Pluhacek, J.; Bartos, L.; Culik, L.
	Title	High-ranking mares of captive plains zebra Equus burchelli have greater reproductive success than low-ranking mares			Type	Journal Article
	Year	2006	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	99	Issue	3-4	Pages	315-329
	Keywords	Equus burchelli; Social hierarchy; Reproductive success; Inter-birth interval; Zoo; Zebra
	Abstract	Plains zebra live in harems that include one to six adult mares. Between these mares is a strong order of social hierarchy. The social rank of an equid mare is typically correlated with her age. Further, high-ranking captive plains zebra mares produce more surviving offspring than low-ranking mares. The objectives of this study were to, first, examined the factors that influence social rank of captive plains zebra mares, and second, test if high-ranking mares conceive earlier and if they have shorter inter-birth intervals than low-ranking ones. We observed three herds of captive plains zebra (a total of 18 mares) at the Dvur Kralove Zoo, Czech Republic. During the 831 h of observation, we recorded 1713 aggressive interactions (biting and offensive kicking) between the mares. These data were used to determine, for each mare, the total number of mares that dominated her in each period of social stability. The GLMM model revealed that older mares were dominated by a lower number of mares than the younger mares. We also found that the probability that a mare would conceive declined with the increasing number of dominant mares. Further, we tested the relationship between the number of dominant mares and the inter-birth interval using 29 intervals for 15 mares. These inter-birth intervals were divided into two groups. When a stallion was continuously present in the herd, the intervals lasted from one birth to the next birth (natural intervals). When a herd was without a stallion, the intervals lasted from the release of the stallion into the herd to the birth of foal (stallion-influenced intervals). The analysis revealed that the inter-birth intervals decreased with an increasing number of dominant mares and the natural intervals decreased with an increasing number of offspring successfully reared by a mare. This finding is the first one in equids and contributes to the previous findings that suggest that social status influences reproductive success.
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	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2228
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	Author	Vieuille, C.; Berger, F.; Le Pape, G.; Bellanger, D.
	Title	Sow behaviour involved in the crushing of piglets in outdoor farrowing huts--a brief report			Type	Journal Article
	Year	2003	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	80	Issue	2	Pages	109-115
	Keywords	Pig-maternal behaviour; Crushing; Free-ranging; Welfare
	Abstract	This study focuses on maternal activities involved in the trapping of piglets by the sow's body in outdoor farrowing systems and examine the mother-piglet context leading either to the death of piglets or to their survival. The behaviour of six Large-WhitexLandrace sows and their litters was continuously video recorded at their first and second parity, during the 40 h following parturition. Crushing mainly occurred at evening and night, during the first 12 h of farrowing and involved changes between lying, sitting and standing positions, as well as between udder and side lying. No piglet died from savaging. Nevertheless, aggressive behaviours of sows were observed, particularly in their first maternal experience. The immediate context of trapping was related to the labour of the sow and to the feeding and resting of piglets. The immediate crushing context was related to active avoidance of restless piglets while lying down, as well as sitting and standing behaviours. These results are discussed in terms of differential reactions of the sow to suckling attempts of piglets.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2235
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	Author	Berger, A.; Scheibe, K.-M.; Eichhorn, K.; Scheibe, A.; Streich, J.
	Title	Diurnal and ultradian rhythms of behaviour in a mare group of Przewalski horse (Equus ferus przewalskii), measured through one year under semi-reserve conditions			Type	Journal Article
	Year	1999	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	64	Issue	1	Pages	1-17
	Keywords	Horse; Przewalski horse; Circadian rhythm; Ultradian rhythm; Feeding and nutrition; Telemetry; Stress
	Abstract	Investigations were conducted on four horses from a group of 12 Przewalski mares raised in different zoos and kept in a 44-ha enclosure under semi-natural conditions. Activity and feeding were continuously measured every second and were saved every 15 min by the storage-telemetry system ETHOSYS, from June 1995 to July 1996. Body mass of the horses was regularly recorded. Daily and monthly mean values, power spectra and DFC (as a measure for stability of rhythms synchronised with circadiurnal period) for activity and feeding were calculated. The general pattern of activity and feeding over the year was closely related to sunrise and sunset. Feeding accounted for 40% of total activity in summer and 62% in spring (all-year average being 52%). The level of activity was lowest in winter; whereas feeding was lowest in summer. The time budget for feeding reflected both feeding conditions and the annual pattern of body condition. Greatest activity occurred during daylight hours. Only on hot summer days, activity at night was higher than during daylight hours. Spectral analysis of activity and feeding in Przewalski horse showed a time pattern which was characterised by 24-h rhythmicity, but also by ultradian components with period lengths between 4.8 and 12 h, i.e., an activity pattern of up to five strong bouts per day. Annual variation in the pattern of power spectra was not high during the year. Results are discussed in connection with horse feeding strategy. Analysing the time structure of long-term and continuously measured activity and feeding could be a useful method to follow the general living conditions, especially the nutritional situation and to detect stressful conditions.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2236
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	Author	Scheibe, K.M.; Schleusner, T.; Berger, A.; Eichhorn, K.; Langbein, J.; Dal Zotto, L.; Streich, W.J.
	Title	ETHOSYS (R)--new system for recording and analysis of behaviour of free-ranging domestic animals and wildlife			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	55	Issue	3-4	Pages	195-211
	Keywords	Telemetry; Diurnal patterns; Activity; Rhythms; Feeding and nutrition
	Abstract	A storage telemetry system has been developed to monitor domestic animals and wildlife, and has been tested under variable conditions on sheep, Przewalski horse and mouflon. It can be used for automatic recording of different patterns of behaviour, such as activity and feeding, and is based on advanced analysis of sensor-emitted signals. The system is made up of collars (ETHOREC) with sensors and electronic devices for behaviour recording, a central station (ETHOLINK) and software for data transmission and processing (ETHODAT). All components of the ETHOREC recording device are integrated in the collar. Long-time recording of behaviour through up to four different channels and in numerous animals at one and the same time are necessary elements to facilitate biorhythmic analysis of animals under free-ranging conditions. The results obtained from this telemetry system were compared with visual observations on six sheep and four Przewalski horses. Parallel recordings were taken from four sheep, using a recorder for jaw movements. Locomotor activity usually was rated somewhat higher by observers, whereas feed uptake was rated lower. Higher feed uptake values were measured by means of the jaw movement recorder, although deviations thus measured varied less than those noticed by visual observations. All measured series exhibited significant correlations with control values. The system, consequently, was found to be more suitable for determination of diurnal patterns, change over time and relative comparison between behaviour levels than it actually was for measurement of absolute duration of a given behaviour.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2237
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	Author	Boyd, L.
	Title	The 24-h time budget of a takh harem stallion (Equus ferus przewalskii) pre- and post-reintroduction			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	4	Pages	291-299
	Keywords	Takhi; Przewalski's horse; Asian wild horse; Mongolian wild horse; Time budget; Reintroduction
	Abstract	Focal animal sampling was used to determine the 24-h time budget of a takh harem stallion (Equus ferus przewalskii) during the 2 weeks prior to, and the two weeks following, reintroduction into the Hustain Nuruu Steppe Reserve, Mongolia. Both before and after release, the stallion spent approximately 47% of his time grazing, 6% standing, and 5% in recumbent rest. The biggest changes to the time budget after release were a 4-fold increase in the amount of time spent moving, and a 50% decrease in the amount of time spent resting in a standing position. During the middle of the day when the temperatures were hottest, the stallion exhibited less grazing and more standing resting behaviour than in the morning or evening hours. Recumbent rest invariably occurred in the hours before dawn.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2245
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	Author	Christensen, J.W.; Keeling, L.J.; Nielsen, B.L.
	Title	Responses of horses to novel visual, olfactory and auditory stimuli			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	93	Issue	1-2	Pages	53-65
	Keywords	Horses; Novelty responses; Fear; Behaviour; Heart rate
	Abstract	Responses of horses towards fear-eliciting stimuli can have important consequences for both human and horse safety. This experiment was designed to investigate behavioural and heart rate (HR) responses of horses to novel visual, auditory and olfactory stimuli. Twenty-four 2-year-old, previously unhandled, stallions were habituated to receive a food reward from a container in a test arena. Each horse was exposed to three 2 min tests in a balanced design where in addition to the feed container, either a traffic cone (visual test), white noise (auditory test) or eucalyptus oil applied to the inside of the container (olfactory test) were used as the novel stimuli. Compared to the control, less time was spent eating during all tests. There was no difference in locomotion activity in the different test situations, but presentation of the novel visual and auditory stimuli elicited significantly increased HR responses in the horses, compared to their response to the arena without novel stimuli (control), whereas there was no increase in HR response to the olfactory stimulus. However, during the olfactory test, the horses had an increased number of eating bouts and became more vigilant towards their surroundings, whereas during the visual and auditory tests, more time was spent alert towards the stimulus. The horses also took significantly more steps backwards in response to the auditory test. The heart rate responses correlated between tests and reflect a non-differentiated activation of the sympathetic nervous system, while the behavioural responses were linked to the type of stimulus.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2256
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	Author	Christensen, J.W.; Ladewig, J.; Sondergaard, E.; Malmkvist, J.
	Title	Effects of individual versus group stabling on social behaviour in domestic stallions			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	75	Issue	3	Pages	233-248
	Keywords	Horse; Individual stabling; Group stabling; Social behaviour
	Abstract	Domestic horses (Equus caballus) are typically kept in individual housing systems, in which they are deprived of physical contact. In order to study the effects of social restrictions on behaviour in young horses, nineteen 2-year-old stallions were housed either singly (n=7), or in groups of three (n=12) for 9 months. Subsequently, the stallions were released into two separate 2 ha enclosures according to treatment, and recordings were made on social interactions and nearest neighbours during a 6-week-period, 28 h per week. Previously group stabled stallions frequently had a former group mate as their nearest neighbour (P=0.001), whereas previously singly stabled stallions did not associate more with their former box neighbours, to whom physical contact was limited by bars during the previous treatment. The nearest neighbour was more frequently recorded to be within one horselength of singly stabled than of group stabled stallions (P=0.005). More aggressive behaviour was recorded in the group of previously singly stabled stallions, i.e. bite threats (P=0.032), whereas group stabled stallions tended to make more use of subtle agonistic interactions (displacements, submissive behaviour). Singly stabled stallions also responded to the 9 months of social deprivation by significantly increasing the level of social grooming (P<0.001) and play behaviour (P<0.001), when subsequently interacting freely with other horses. The increased occurrence may relate to a build-up of motivation (a rebound effect), as well as to external factors, such as playful pasture companions and the increased space allowance of the pasture. It is concluded that 2-year-old domestic stallions are sensitive to social deprivation and that stabling has long-term effects, lasting 6 weeks at least, on the social behaviour in stallions.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2257
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	Author	Barber, J.A.; Crowell-Davis, S.L.
	Title	Maternal behavior of Belgian (Equus caballus) mares			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	41	Issue	3-4	Pages	161-189
	Keywords	Belgian; Draft horse; Ontogeny; Maternal aggression; Maternal behavior; Nursing behavior; Recumbency response
	Abstract	The relationship between ten Belgian mares and their offspring was studied from the first day of foal life to 17 weeks of age. Mares and foals spent more time at greater distances from each other as foals matured. Mares exhibited the recumbency response, being in closer proximity to their foals when foals were recumbent than when they were upright. Foals initiated the majority of nursing bouts. Frequency and duration of nursing bouts and percentage of time resting recumbently declined as foals matured. Foals also terminated the their foals, and they were most likely to do so in the first month of foal life. Maternal initiation of nursing. There was usually no discernible foal response to maternal aggression. Little difference between maternal behavior directed towards colts and fillies was found for all aspects of the study. Maternal behavior in the Belgian draft horse was similar to that reported for other equid breeds.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2268
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	Author	Crowell-Davis, S.L.
	Title	Daytime rest behavior of the Welsh pony (Equus caballus) mare and foal			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	40	Issue	3-4	Pages	197-210
	Keywords	Horse; Behavior; Rest; Sleep; Pony; Foal
	Abstract	Upright and recumbent rest of 15 Welsh pony foals and their mothers was studied over a 2 year period. During their first week of life, the foals spent 32% of the time in recumbent rest. Subsequently, the percentage of time spent in recumbent rest decreased, but was still greater than for the foal's mother by Week 21, when the foals spent 6.5% of their time in recumbent rest. Adults spent little time in recumbent rest. Foals rested upright only 3.5% of the time during their first week of life. Mares rested upright more than foals did to Week 13, at which time peak values for time spent in upright rest occurred for both mares (32.5%) and foals (23%). Subsequently, mares and foals spent equal, but decreasing, amounts of time resting upright. The total time spent resting by the foals decreased gradually, and was characterized by a transition from recumbent rest to upright rest. Foals were more likely to be resting, either recumbent or upright, if their mother was resting upright. During the late spring, summer, and early autumn, mares and foals were most likely to be resting upright between 09:00 and 17:00 h.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2269
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