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Author (up) Clark, T.B.; Peterson, B.V.; Whitcomb, R.F.; Henegar, R.B.; Hackett, K.J.; Tully, J.G. openurl 
  Title Spiroplasmas in the Tabanidae Type Journal Article
  Year 1984 Publication Israel Journal of Medical Sciences Abbreviated Journal Isr J Med Sci  
  Volume 20 Issue 10 Pages 1002-1005  
  Keywords Animals; Diptera/*microbiology/ultrastructure; Ecology; Female; Hemolymph/microbiology; Male; Maryland; North Carolina; Plants/microbiology; Spiroplasma/classification/*isolation & purification/physiology  
  Abstract Spiroplasmas were observed in seven species of the family Tabanidae (horse flies and deer flies). This is the fifth family of the order Diptera now known to harbor spiroplasmas. Noncultivable spiroplasmas were seen in the hemolymph of three species of the genus Tabanus, and cultivable forms were isolated from the guts of six species in three genera. Isolates from T. calens and T. sulcifrons were serologically similar and closely related to a spiroplasma in the lampyrid beetle, Ellychnia corrusca. These three isolates represent a new serogroup. Isolates from Hybomitra lasiophthalma were related to Group IV strains, while those from T. nigrovittatus and Chrysops sp. both represented new serogroups. At least some tabanids probably acquire spiroplasmas from contaminated flower surfaces. The possibility of vertebrate reservoirs for some tabanid spiroplasmas remains an open question.  
  Address  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0021-2180 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:6511308 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2687  
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Author (up) Clayton, H.M. openurl 
  Title Classification of collected trot, passage and piaffe based on temporal variables Type Journal Article
  Year 1997 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 23 Pages 54-57  
  Keywords Analysis of Variance; Animals; Discriminant Analysis; Gait/*physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Male; Running/physiology; Time Factors; Videotape Recording; Walking/physiology  
  Abstract The objective was to determine whether collected trot, passage and piaffe could be distinguished as separate gaits on the basis of temporal variables. Sagittal plane, 60 Hz videotapes of 10 finalists in the dressage competitions at the 1992 Olympic Games were analysed to measure the temporal variables in absolute terms and as percentages of stride duration. Classification was based on analysis of variance, a graphical method and discriminant analysis. Stride duration was sufficient to distinguish collected trot from passage and piaffe in all horses. The analysis of variance showed that the mean values of most variables differed significantly between passage and piaffe. When hindlimb stance percentage was plotted against diagonal advanced placement percentage, some overlap was found between all 3 movements indicating that individual horses could not be classified reliably in this manner. Using hindlimb stance percentage and diagonal advanced placement percentage as input in a discriminant analysis, 80% of the cases were classified correctly, but at least one horse was misclassified in each movement. When the absolute, rather than percentage, values of the 2 variables were used as input in the discriminant analysis, 90% of the cases were correctly classified and the only misclassifications were between passage and piaffe. However, the 2 horses in which piaffe was misclassified as passage were the gold and silver medallists. In general, higher placed horses tended toward longer diagonal advanced placements, especially in collected trot and passage, and shorter hindlimb stance percentages in passage and piaffe.  
  Address Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
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  Notes PMID:9354290 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3738  
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Author (up) Clayton, H.M. openurl 
  Title Comparison of the stride kinematics of the collected, medium, and extended walks in horses Type Journal Article
  Year 1995 Publication American Journal of Veterinary Research Abbreviated Journal Am J Vet Res  
  Volume 56 Issue 7 Pages 849-852  
  Keywords Analysis of Variance; Animals; *Gait; Horses/*physiology; *Locomotion; Motion Pictures; Physical Conditioning, Animal; Walking  
  Abstract Six horses, highly trained for dressage competition, were used to study the stride kinematics of the walk, and to compare the kinematics of the collected, medium, and extended walks. Horses were filmed in a sagittal plane at a rate of 150 frames/s; temporal, linear, and angular data were extracted from the films. Results of ANOVA and Duncan's multiple range test indicated that the speed of the collected walk (1.37 m/s) was significantly (P < 0.01) slower than that of the medium (1.73 m/s) and extended (1.82 m/s) walks, values for which were not significantly different from each other. The increase in speed was associated with a significant increase in stride length, from 157 cm in the collected walk to 193 cm in the extended walk. This was a result of an increase in the over-tracking distance, whereas there was no significant difference in the distance between lateral placements of the limbs. Stride duration decreased (P < 0.01) from the collected walk (1,159 ms) to the extended walk (1,064 ms). Angles of the metacarpal and metatarsal segments, measured on the palmar/ plantar aspect, were higher at impact and lower at lift off in the collected than in the extended walk (P < 0.01). This indicated greater range of angular motion of this segment during the stance phase in the extended walk. Only 1 of the 6 horses had a regular 4-beat rhythm of the footfalls, with equal time elapsing between the lateral and diagonal footfalls.  
  Address Department of Veterinary Anatomy, Western College of Veterinary Medicine, Canada  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0002-9645 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:7574149 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3743  
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Author (up) Clayton, H.M. openurl 
  Title Comparison of the stride kinematics of the collected, working, medium and extended trot in horses Type Journal Article
  Year 1994 Publication Equine Veterinary Journal Abbreviated Journal Equine Vet J  
  Volume 26 Issue 3 Pages 230-234  
  Keywords Analysis of Variance; Animals; Biomechanics; Female; Forelimb/anatomy & histology/physiology; Gait/*physiology; Hindlimb/anatomy & histology/physiology; Horses/anatomy & histology/*physiology; Locomotion/physiology; Male; Motion Pictures  
  Abstract Highly-trained dressage horses were studied to test the hypothesis that stride length is altered independently of stride duration in the transitions between the collected, working, medium and extended trot. Six well-trained dressage horses were filmed at a frame rate of 150 frames/s performing the collected, working, medium and extended trots in a sand arena. Temporal, linear and angular data were extracted from the films, with 4 strides being analysed for each horse and gait type. There were no significant asymmetries between the left and rights limbs or diagonals when data from the whole group were pooled, but 3 horses showed asymmetries in one or more variables (P < 0.01). Analysis of variance and post-hoc tests indicated that the speed increased significantly (P < 0.01) from the collected (3.20 m/s) to the working (3.61 m/s) to the medium (4.47 m/s) to the extended (4.93 m/s) trot. The increases in speed were associated with a significant increase in stride length from 250 cm in the collected trot, to 273 cm in the working trot, 326 cm in the medium trot and 355 cm in the extended trot (P < 0.01). The lengthening of the stride was a result of increases between each gait type in the over-reach distance, whereas the diagonal distance was significantly longer in the extended than the collected trot only (P < 0.01). The stride duration tended to decrease as speed increased, and the difference became significant between the collected and extended trots (P < 0.01).  
  Address Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0425-1644 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:8542844 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3746  
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Author (up) Clayton, H.M. openurl 
  Title The extended canter: a comparison of some kinematic variables in horses trained for dressage and for racing Type Journal Article
  Year 1993 Publication Acta Anatomica Abbreviated Journal Acta Anat (Basel)  
  Volume 146 Issue 2-3 Pages 183-187  
  Keywords Animal Husbandry; Animals; *Gait; Horses/*physiology; *Physical Conditioning, Animal; *Sports  
  Abstract This study was designed to test the hypothesis that there is no significant difference in selected temporal and linear stride variables of the extended canter in horses bred and trained for dressage or racing. Nine advanced-level dressage horses and 7 Thoroughbred racehorses were filmed at a frame rate of 200 Hz at an extended canter on a sand track. Two strides were recorded per trial, and each horse performed 6 or 7 trials. Temporal and linear data were determined from the films, and descriptive statistics (mean, SD) were calculated. Strides were selected for analysis on the basis of having a velocity in the range of 6.0-7.0 m/s, and multivariate analysis of variance was used to detect significant differences in the stride kinematics of horses trained for the two sports (p < or = 0.01). The average velocity of the dressage horses was 6.37 m/s, compared with 6.40 m/s for the racehorses. There were no significant differences between the two groups in velocity, stride duration, stride length or the distances between limb placements. The stance durations of all four limbs and the overlaps between them were longer, whereas the duration of the suspension phase was shorter in the dressage horses than in the racehorses (p < or = 0.01). The time between impacts of the diagonal limb pair was close to zero in both groups, with individual horses showing some variability in the order of placement of the diagonal limb pair. However, the sequence of footfalls was not significantly different between the two groups (p < or = 0.01).  
  Address Department of Veterinary Anatomy, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
  Corporate Author Thesis  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0001-5180 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:8470464 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3751  
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Author (up) Clayton, H.M. openurl 
  Title Development of conditioning programs for dressage horses based on time-motion analysis of competitions Type Journal Article
  Year 1993 Publication Journal of Applied Physiology (Bethesda, Md. : 1985) Abbreviated Journal J Appl Physiol  
  Volume 74 Issue 5 Pages 2325-2329  
  Keywords Animals; Horses/*physiology; *Physical Conditioning, Animal; *Sports; Statistics; Time Factors; *Time and Motion Studies; Videotape Recording  
  Abstract The time-motion characteristics of Canadian basic- and medium-level dressage competitions are described, and the results are applied in formulating sport-specific conditioning programs. One competition was analyzed at the six levels from basic 1 to medium 3. Each test was divided into a series of sequences based on the type and speed of activity. The durations of the sequences were measured from videotapes. The basic-level tests had fewer sequences, and they were shorter in distance and duration than the medium tests (P < 0.10), but the average speed did not differ between the two levels. It is recommended that horses competing at the basic levels be conditioned using 5-min exercise periods, with short (10-s) bursts of lengthened trot and canter included at basic 2 and above. In preparation for medium-level competitions, the duration of the work periods increases to 7 min, 10- to 12-s bursts of medium or extended trot and canter are included, and transitions are performed frequently to simulate the energy expenditure in overcoming inertia.  
  Address Department of Veterinary Anatomy, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 8750-7587 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:8335563 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3750  
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Author (up) Clayton, H.M.; Lanovaz, J.L.; Schamhardt, H.C.; van Wessum, R. openurl 
  Title The effects of a rider's mass on ground reaction forces and fetlock kinematics at the trot Type Journal Article
  Year 1999 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume 30 Issue Pages 218-221  
  Keywords Animals; Body Weight; Computer Simulation; Gait/*physiology; Horses/*physiology; Physical Conditioning, Animal/*physiology; Stress, Mechanical; Weight-Bearing/*physiology  
  Abstract Ground reaction force (GRF) measurements are often normalised to body mass to facilitate inter-individual comparisons. The objective of this study was to explore the effect of a rider on the GRFs and fetlock joint kinematics of trotting horses. The subjects were 5 dressage-trained horses and 3 experienced dressage riders. Ground reaction force measurements and sagittal view videotapes were recorded as the horses trotted at the same velocity in hand (3.49 +/- 0.52 m/s) and with a rider (3.49 +/- 0.46 m/s). Data were time-normalised to stance duration. Ground reaction force measurements were expressed in absolute terms and normalised to the system mass (horse or horse plus rider). All the horses showed changes in the same direction when comparing the ridden condition with the in-hand condition. There was an increase in the absolute peak vertical GRFs of the fore- and hindlimbs with a rider. However, the mass-normalised peak vertical GRFs were lower for the ridden condition, with the peak occurring later in the forelimbs and earlier in the hindlimbs compared with the inhand condition. Maximal fetlock angle and its time of occurrence were similar for the 2 conditions, but the fore fetlock joint was more extended during the later part of the stance phase in ridden horses. The presence of a rider appeared to affect the GRFs and fetlock joint kinematics differently in the fore- and hindlimbs, and the ridden horse did not seem to be equivalent to a proportionately larger horse. This should be considered when normalising for body mass in studies comparing horses in hand and ridden horses.  
  Address Department of Large Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing 48824-1314, USA  
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  Notes PMID:10659255 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3733  
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Author (up) Clement, T.S.; Feltus, J.R.; Kaiser, D.H.; Zentall, T.R. openurl 
  Title “Work ethic” in pigeons: reward value is directly related to the effort or time required to obtain the reward Type Journal Article
  Year 2000 Publication Psychonomic bulletin & review Abbreviated Journal Psychon Bull Rev  
  Volume 7 Issue 1 Pages 100-106  
  Keywords Animals; Behavior, Animal/*physiology; Cognition/physiology; Columbidae; Discrimination Learning/physiology; Female; Male; Reinforcement (Psychology); *Reward; Time Factors  
  Abstract Stimuli associated with less effort or with shorter delays to reinforcement are generally preferred over those associated with greater effort or longer delays to reinforcement. However, the opposite appears to be true of stimuli that follow greater effort or longer delays. In training, a simple simultaneous discrimination followed a single peck to an initial stimulus (S+FR1 S-FR1) and a different simple simultaneous discrimination followed 20 pecks to the initial stimulus (S+FR20 S-FR20). On test trials, pigeons preferred S+FR20 over S+FR1 and S-FR20 over S-FR1. These data support the view that the state of the animal immediately prior to presentation of the discrimination affects the value of the reinforcement that follows it. This contrast effect is analogous to effects that when they occur in humans have been attributed to more complex cognitive and social factors.  
  Address University of Kentucky, Lexington, USA  
  Corporate Author Thesis  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1069-9384 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:10780022 Approved no  
  Call Number refbase @ user @ Serial 248  
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Author (up) Clutton-Brock, T.H.; Russell, A.F.; Sharpe, L.L.; Brotherton, P.N.; McIlrath, G.M.; White, S.; Cameron, E.Z. doi  openurl
  Title Effects of helpers on juvenile development and survival in meerkats Type Journal Article
  Year 2001 Publication Science (New York, N.Y.) Abbreviated Journal Science  
  Volume 293 Issue 5539 Pages 2446-2449  
  Keywords Animals; Breeding; Carnivora/growth & development/*physiology; *Cooperative Behavior; Feeding Behavior; Female; Male; Survival Rate; *Weight Gain  
  Abstract Although breeding success is known to increase with group size in several cooperative mammals, the mechanisms underlying these relationships are uncertain. We show that in wild groups of cooperative meerkats, Suricata suricatta, reductions in the ratio of helpers to pups depress the daily weight gain and growth of pups and the daily weight gain of helpers. Increases in the daily weight gain of pups are associated with heavier weights at independence and at 1 year of age, as well as with improved foraging success as juveniles and higher survival rates through the first year of life. These results suggest that the effects of helpers on the fitness of pups extend beyond weaning and that helpers may gain direct as well as indirect benefits by feeding pups.  
  Address Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK. thcb@hermes.cam.ac.uk  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0036-8075 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:11577235 Approved no  
  Call Number refbase @ user @ Serial 414  
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Author (up) Colahan, P.; Lindsey, E.; Nunier, C. openurl 
  Title Determination of the center of pressure of the hoofs of the forelimbs of horses standing on a flat level surface Type Journal Article
  Year 1993 Publication Acta Anatomica Abbreviated Journal Acta Anat (Basel)  
  Volume 146 Issue 2-3 Pages 175-178  
  Keywords Animals; Forelimb/*physiology; Hoof and Claw/*physiology; Horses/*physiology; *Posture; Pressure  
  Abstract The pressure exerted on a flat level surface by recently trimmed, unshod hoofs of the front limbs of 23 sound, adult horses was measured using pressure-sensitive film and a specially built cassette. The horses were tranquilized and stood with one foot on the 2.9-cm-thick cassette and the other on a block of equal height. The hoofs were observed for motion during the measurement, and the developed film was examined for improper alignment of the film or slipping of the hoof. The center of pressure was located using the method of weighted proportions of Barrey. This static measurement system with a long measurement time and the number of measurements reduced the influence of variables inherent in the horses' behavior and the measuring system. The calculated point was recorded as falling medial to, lateral to or on a line bisecting the central sulcus of the frog. In the dorsal to palmar orientation the point was classified with reference to a line drawn halfway between the most dorsal and the most palmar mark on the film. Forty-six percent of the calculated centers of pressure were located in the medial heel area. Binomial analysis for large samples indicates that this was a significant variation from a random distribution. Seventy-six percent of the centers were located in or on the borders of the medial heel.  
  Address College of Veterinary Medicine, University of Florida, Gainesville 32608-0136  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0001-5180 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:8470462 Approved no  
  Call Number refbase @ user @ Serial 1946  
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