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	Author	Hostetter, A.B.; Russell, J.L.; Freeman, H.; Hopkins, W.D.
	Title	Now you see me, now you don't: evidence that chimpanzees understand the role of the eyes in attention			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	1	Pages	55-62
	Keywords	Animal Husbandry/methods; Animals; Attention; Awareness; Female; Fixation, Ocular/physiology; Humans; Male; Pan troglodytes/psychology; Social Behavior; *Social Perception
	Abstract	Chimpanzees appear to understand something about the attentional states of others; in the present experiment, we investigated whether they understand that the attentional state of a human is based on eye gaze. In all, 116 adult chimpanzees were offered food by an experimenter who engaged in one of the four experimental manipulations: eyes closed, eyes open, hand over eyes, and hand over mouth. The communicative behavior of the chimpanzees was observed. More visible behaviors were produced when the experimenter's eyes were visible than when the experimenter's eyes were not visible. More vocalizations were produced when the experimenter's eyes were closed than when they were open, but there were no differences in other attention getting behaviors. There was no effect of age or rearing history. The results suggest that chimpanzees use the presence of the eyes as a cue that their visual gestures will be effective.
	Address	Department of Psychology, University of Wisconsin-Madison, 1202 W. Johnson Street, Madison, WI 53706, USA. abhostetter@wisc.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16847659			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2457
Permanent link to this record



	Author	Morton, D.B.
	Title	Self-consciousness and animal suffering			Type	Journal Article
	Year	2000	Publication	Biologist (London, England)	Abbreviated Journal	Biologist (London)
	Volume	47	Issue	2	Pages	77-80
	Keywords	Animal Population Groups/psychology; Animal Welfare/standards; Animals; Behavior, Animal; Consciousness; Dogs; Ego; Horses/psychology; Pain/psychology/*veterinary; Pan troglodytes/psychology; Parrots; Pongo pygmaeus/psychology; Self Concept
	Abstract	Animals with relatively highly developed brains are likely to experience some degree of self-awareness and the ability to think. As well as being interesting in its own right, self-consciousness matters from an ethical point of view, since it can give rise to forms of suffering above and beyond the immediate physical sensations of pain or distress. This article surveys the evidence for animal self-consciousness and its implications for animal welfare.
	Address	Division of Primary Care, Public and Occupational Health, School of Medicine, University of Birmingham, Edgbaston, Birmingham, B15 2TT, UK. d.b.morton@bham.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-3347	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11190233			Approved	no
	Call Number	refbase @ user @			Serial	618
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	Author	Russell, J.L.; Braccini, S.; Buehler, N.; Kachin, M.J.; Schapiro, S.J.; Hopkins, W.D.
	Title	Chimpanzee (Pan troglodytes) intentional communication is not contingent upon food			Type	Journal Article
	Year	2005	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	4	Pages	263-272
	Keywords	Animal Communication; Animals; Feeding Behavior; Female; Humans; Male; Pan troglodytes/*psychology
	Abstract	Studies of great apes have revealed that they use manual gestures and other signals to communicate about distal objects. There is also evidence that chimpanzees modify the types of communicative signals they use depending on the attentional state of a human communicative partner. The majority of previous studies have involved chimpanzees requesting food items from a human experimenter. Here, these same communicative behaviors are reported in chimpanzees requesting a tool from a human observer. In this study, captive chimpanzees were found to gesture, vocalize, and display more often when the experimenter had a tool than when she did not. It was also found that chimpanzees responded differentially based on the attentional state of a human experimenter, and when given the wrong tool persisted in their communicative efforts. Implications for the referential and intentional nature of chimpanzee communicative signaling are discussed.
	Address	Division of Psychobiology, Yerkes National Primate Research Center, Atlanta, GA 30322, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15742162			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2491
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	Author	Parr, L.A.
	Title	Perceptual biases for multimodal cues in chimpanzee (Pan troglodytes) affect recognition			Type	Journal Article
	Year	2004	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	7	Issue	3	Pages	171-178
	Keywords	Acoustic Stimulation; Animal Communication; Animals; Auditory Perception/physiology; Cues; Discrimination Learning/physiology; Facial Expression; Female; Male; Pan troglodytes/psychology; Perceptual Masking/physiology; Photic Stimulation; Recognition (Psychology)/physiology; Visual Perception/physiology; Vocalization, Animal
	Abstract	The ability of organisms to discriminate social signals, such as affective displays, using different sensory modalities is important for social communication. However, a major problem for understanding the evolution and integration of multimodal signals is determining how humans and animals attend to different sensory modalities, and these different modalities contribute to the perception and categorization of social signals. Using a matching-to-sample procedure, chimpanzees discriminated videos of conspecifics' facial expressions that contained only auditory or only visual cues by selecting one of two facial expression photographs that matched the expression category represented by the sample. Other videos were edited to contain incongruent sensory cues, i.e., visual features of one expression but auditory features of another. In these cases, subjects were free to select the expression that matched either the auditory or visual modality, whichever was more salient for that expression type. Results showed that chimpanzees were able to discriminate facial expressions using only auditory or visual cues, and when these modalities were mixed. However, in these latter trials, depending on the expression category, clear preferences for either the visual or auditory modality emerged. Pant-hoots and play faces were discriminated preferentially using the auditory modality, while screams were discriminated preferentially using the visual modality. Therefore, depending on the type of expressive display, the auditory and visual modalities were differentially salient in ways that appear consistent with the ethological importance of that display's social function.
	Address	Division of Psychobiology, Yerkes National Primate Research Center, Emory University, 954 Gatewood Road, GA 30329, Atlanta, USA. parr@rmy.emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14997361			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2544
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	Author	Collier-Baker, E.; Davis, J.M.; Nielsen, M.; Suddendorf, T.
	Title	Do chimpanzees (Pan troglodytes) understand single invisible displacement?			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	1	Pages	55-61
	Keywords	Animals; Behavior, Animal; Cognition; Male; Pan troglodytes/psychology; Space Perception; Spatial Behavior; Task Performance and Analysis; *Visual Perception
	Abstract	Previous research suggests that chimpanzees understand single invisible displacement. However, this Piagetian task may be solvable through the use of simple search strategies rather than through mentally representing the past trajectory of an object. Four control conditions were thus administered to two chimpanzees in order to separate associative search strategies from performance based on mental representation. Strategies involving experimenter cue-use, search at the last or first box visited by the displacement device, and search at boxes adjacent to the displacement device were systematically controlled for. Chimpanzees showed no indications of utilizing these simple strategies, suggesting that their capacity to mentally represent single invisible displacements is comparable to that of 18-24-month-old children.
	Address	Early Cognitive Development Unit, School of Psychology, University of Queensland, Brisbane, Queensland 4072, Australia. e.collier-baker@psy.uq.edu.au
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16163481			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2482
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	Author	Nielsen, M.; Collier-Baker, E.; Davis, J.M.; Suddendorf, T.
	Title	Imitation recognition in a captive chimpanzee (Pan troglodytes)			Type	Journal Article
	Year	2005	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	1	Pages	31-36
	Keywords	Animals; Awareness; Humans; Imitative Behavior; Male; Motor Activity; Movement; Pan troglodytes/psychology; *Recognition (Psychology)
	Abstract	This study investigated the ability of a captive chimpanzee (Pan troglodytes) to recognise when he is being imitated. In the experimental condition of test 1a, an experimenter imitated the postures and behaviours of the chimpanzee as they were being displayed. In three control conditions the same experimenter exhibited (1) actions that were contingent on, but different from, the actions of the chimpanzee, (2) actions that were not contingent on, and different from, the actions of the chimpanzee, or (3) no action at all. The chimpanzee showed more “testing” sequences (i.e., systematically varying his actions while oriented to the imitating experimenter) and more repetitive behaviour when he was being imitated, than when he was not. This finding was replicated 4 months later in test 1b. When the experimenter repeated the same actions she displayed in the experimental condition of test 1a back to the chimpanzee in test 2, these actions now did not elicit those same testing sequences or repetitive behaviours. However, a live imitation condition did. Together these results provide the first evidence of imitation recognition in a nonhuman animal.
	Address	Early Cognitive Development Unit, School of Psychology, University of Queensland, QLD, Brisbane, 4072, Australia. nielsen@psy.uq.edu.au
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15322942			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2515
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	Author	de Waal, F.B.M.
	Title	How animals do business			Type	Journal Article
	Year	2005	Publication	Scientific American	Abbreviated Journal	Sci Am
	Volume	292	Issue	4	Pages	54-61
	Keywords	Animals; Attitude; Behavior, Animal; Cebus; Cooperative Behavior; Economics; Emotions; Fishes; Food; Humans; Pan troglodytes; Papio; Social Behavior
	Abstract
	Address	Emory University, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0036-8733	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15915815			Approved	no
	Call Number	refbase @ user @			Serial	166
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	Author	O'Connell, S.; Dunbar, R.I.M.
	Title	The perception of causality in chimpanzees (Pan spp.)			Type	Journal Article
	Year	2005	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	1	Pages	60-66
	Keywords	Animals; Association Learning; Awareness; Concept Formation; Female; Habituation, Psychophysiologic; Male; Pan paniscus/psychology; Pan troglodytes/*psychology; Perception
	Abstract	Chimpanzees (Pan spp.) were tested on a habituation/dishabituation paradigm that was originally developed to test for comprehension of causality in very young human infants. Three versions of the test were used: a food item being moved by a hand, a human pushing another human off a chair to obtain a food item, and a film clip of natural chimpanzee behaviour (capturing and eating a monkey). Chimpanzees exhibited similar results to those obtained with human infants, with significantly elevated levels of looking on the dishabituation trials. Since the level of response was significantly greater on natural/unnatural sequences than on unnatural/natural sequences, we conclude that the chimpanzees were not responding just to novelty but rather to events that infringed their sense of natural causation.
	Address	Evolutionary Psychology Research Group, School of Biological Sciences, Biosciences Building, University of Liverpool, Liverpool, L69 7ZB, UK
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15322943			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2514
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	Author	Dunbar, R.I.M.; McAdam, M.R.; O'connell, S.
	Title	Mental rehearsal in great apes (Pan troglodytes and Pongo pygmaeus) and children			Type	Journal Article
	Year	2005	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	69	Issue	3	Pages	323-330
	Keywords	Algorithms; Animals; Child; Child, Preschool; Food; Frontal Lobe/anatomy & histology/physiology; Humans; Imagination; Pan troglodytes; Pongo pygmaeus; Problem Solving/physiology; Psychomotor Performance/physiology; Reward
	Abstract	The ability to rehearse possible future courses of action in the mind is an important feature of advanced social cognition in humans, and the “social brain” hypothesis implies that it might also be a feature of primate social cognition. We tested two chimpanzees, six orangutans and 63 children aged 3-7 years on a set of four puzzle boxes, half of which were presented with an opportunity to observe the box before being allowed to open it (“prior view”), the others being given without an opportunity to examine the boxes before handling them (“no prior view”). When learning effects are partialled out, puzzle boxes in the “prior view” condition were opened significantly faster than boxes given in the “no prior view” condition by the children, but not by either of the great apes. The three species differ significantly in the speed with which they opened boxes in the “no prior view” condition. The three species' performance on this task was a function of relative frontal lobe volume, suggesting that it may be possible to identify quantitative neuropsychological differences between species.
	Address	Evolutionary Psychology Research Group, School of Biological Sciences, University of Liverpool, Biosciences Building, Crown Street, Liverpool L69 7ZB, UK. rimd@liv.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15896530			Approved	no
	Call Number				Serial	2097
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	Author	Imura, T.; Tomonaga, M.
	Title	Perception of depth from shading in infant chimpanzees ( Pan troglodytes)			Type	Journal Article
	Year	2003	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	6	Issue	4	Pages	253-258
	Keywords	Age Factors; Animals; Contrast Sensitivity/physiology; Depth Perception/physiology; Discrimination Learning/physiology; Female; Male; Pan troglodytes/growth & development/physiology/*psychology; Pattern Recognition, Visual/physiology
	Abstract	We investigated the ability to perceive depth from shading, one of the pictorial depth cues, in three chimpanzee infants aged 4-10 months old, using a preferential reaching task commonly used to study pictorial depth perception in human infants. The chimpanzee infants reached significantly more to three-dimensional toys than to pictures thereof and more to the three-dimensional convex than to the concave. Furthermore, two of the three infants reached significantly more to the photographic convex than to the photographic concave. These infants also looked longer at the photographic convex than the concave. Our results suggest that chimpanzees perceive, at least as early as the latter half of the first year of life, pictorial depth defined by shading information. Photographic convexes contain richer information about pictorial depth (e.g., attached shadow, cast shadow, highlighted area, and global difference in brightness) than simple computer-graphic graded patterns. These cues together might facilitate the infants' perception of depth from shading.
	Address	Graduate School of Humanities, Kwansei Gakuin University, Uegahara, Nishinomiya, Hyogo 662-8501, Japan
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14610661			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2550
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