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	Author	Dyson, S.
	Title	The Ridden Horse Pain Ethogram			Type	Journal Article
	Year	2022	Publication	Equine Veterinary Education	Abbreviated Journal	Equine Vet Educ
	Volume	34	Issue	7	Pages	372-380
	Keywords	horse; lameness; canter; behaviour; saddle-fit; rider skill
	Abstract	Summary The Ridden Horse Pain Ethogram (RHpE) comprises 24 behaviours, the majority of which are at least 10 times more likely to be seen in lame horses compared with non-lame horses. The observation of >=8/24 behaviours is likely to reflect the presence of musculoskeletal pain, although some lame horses score <8/24 behaviours. A marked reduction in RHpE scores after resolution of lameness using diagnostic anaesthesia proves a causal relationship between pain and RHpE scores. Horses should be assessed for approximately 10?min in walk, trot (including 10?m diameter circles), canter and transitions. The validity of the RHpE has been verified for use in horses which perform dressage-type movements, and which have been trained to work with the front of the head in a vertical position. It has not, as yet, been used in horses while jumping, racehorses, western performance or endurance horses. The RHpE provides a valuable tool for riders, trainers, veterinarians and other equine professionals to recognise the presence of musculoskeletal pain, even if overt lameness cannot be recognised. Riders with a higher skill-level may improve gait quality, but cannot obscure behavioural signs of pain, although specific behaviours may change. Tight saddle tree points, the rider sitting on the caudal third of the saddle and rider weight may influence RHpE scores. Accurate application of the RHpE requires training and practice. The RHpE is a powerful tool for the assessment of ridden horses and the identification of likely musculoskeletal pain. Such pain merits further investigation and treatment, to improve equine welfare and performance. The RHpE provides an additional means of evaluating the response to diagnostic anaesthesia. It provides a mechanism for client education and a diplomatic way of communicating with clients about equine discomfort related to saddle-fit, rider size, their position in the saddle and ability to ride in balance.
	Address
	Corporate Author				Thesis
	Publisher	American Medical Association (AMA)	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0957-7734	ISBN		Medium
	Area		Expedition		Conference
	Notes	https://doi.org/10.1111/eve.13468			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6693
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	Author	Gleerup, K.B.; Lindegaard, C.
	Title	Recognition and quantification of pain in horses: A tutorial review			Type	Journal Article
	Year	2016	Publication	Equine Veterinary Education	Abbreviated Journal	Equine Vet Educ
	Volume	28	Issue	1	Pages	47-57
	Keywords	horse; pain evaluation; pain scale; pain behaviour; pain face
	Abstract	Summary Pain management is dependent on the quality of the pain evaluation. Ideally, pain evaluation is objective, pain-specific and easily incorporated into a busy equine clinic. This paper reviews the existing knowledge base regarding the identification and quantification of pain in horses. Behavioural indicators of pain in horses in the context of normal equine behaviour, as well as various physiological parameters potentially useful for pain evaluation, are discussed. Areas where knowledge is sparse are identified and a new equine pain scale based on results from all reviewed papers is proposed. Finally, the most important considerations in relation to the implementation of a pain scale in a hospital setting are discussed.
	Address
	Corporate Author				Thesis
	Publisher	American Medical Association (AMA)	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0957-7734	ISBN		Medium
	Area		Expedition		Conference
	Notes	https://doi.org/10.1111/eve.12383			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6705
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	Author	Malmgren, L.; Andresen, Ø.; Dalin, A.-M.
	Title	Effect of GnRH immunisation on hormonal levels, sexual behaviour, semen quality and testicular morphology in mature stallions			Type	Journal Article
	Year	2001	Publication	Equine Veterinary Journal	Abbreviated Journal	Equine vet. J.
	Volume	33	Issue	1	Pages	75-83
	Keywords	horse; GnRH; immunisation; stallion; testosterone; oestrone sulphate; sexual behaviour; semen quality
	Abstract	Summary The aim of this study was to investigate the effect of gonadotrophin-releasing hormone (GnRH) immunisation on mature stallions that had been used for breeding. Four Standardbred stallions were used in the study: 3 experimental animals and 1 control animal. Semen was collected regularly, i.e. twice/week, during the 4 months prior to the experimental period. The stallions were immunised against GnRH with a GnRH-BSA conjugate. Equimune was used as the adjuvant. The stallions were immunised on 5 occasions, 4 at 2 week intervals, and the fifth 4 weeks after the fourth. Blood samples were taken once a week for analysis of GnRH antibody titre and every third week for testosterone and oestrone sulphate analyses. Semen was collected once a week, and libido and sexual behaviour were observed. Ejaculate volume, sperm concentration, total number of sperm in the ejaculate, sperm motility and sperm morphology were evaluated. Testicular size was measured once a week. At the end of the study, the stallions were castrated, and a histological examination of the testes performed. All immunised stallions produced antibodies against GnRH, and plasma testosterone concentration decreased. However, the effect of immunisation varied between stallions. In 2 of the stallions, high levels of antibodies were found, while in the third, the level was moderate. Four weeks after the first immunisation, a decrease in libido was observed. Two months after the first immunisation, marked changes in semen quality were observed in the 2 stallions with high antibody titres. Fourteen weeks after the first immunisation, the total number of sperm/ejaculate had decreased from >8.6×109 to<2.7×109, sperm motility from >59 to<10% and the frequency of morphological normal spermatozoa had decreased from >60 to<14%. The dominating abnormalities were abnormal head shapes, proximal cytoplasmic droplets and detached heads. In the third stallion, only slight changes in semen quality were found. No changes were observed in the control stallion. Decreases in testicular size were noted in all of the experimental stallions. Pronounced histological alterations in the testes were observed in 2 of the stallions. It is concluded that the vaccine was effective in stimulating production of GnRH antibodies and in suppressing testicular function and androgen secretion. However, there was an individual variation in the responses among the stallions and, further, libido was not totally suppressed.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	2042-3306	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5312
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	Author	Goodwin, D.
	Title	The importance of ethology in understanding the behaviour of the horse			Type	Journal Article
	Year	1999	Publication	Equine Veterinary Journal	Abbreviated Journal	Equine Veterinary Journal
	Volume	31	Issue	S28	Pages	15-19
	Keywords	horse; behaviour; domestication; interspecific communication
	Abstract	Summary Domestication has provided the horse with food, shelter, veterinary care and protection, allowing individuals an increased chance of survival. However, the restriction of movement, limited breeding opportunities and a requirement to expend energy, for the benefit of another species, conflict with the evolutionary processes which shaped the behaviour of its predecessors. The behaviour of the horse is defined by its niche as a social prey species but many of the traits which ensured the survival of its ancestors are difficult to accommodate in the domestic environment. There has been a long association between horses and man and many features of equine behaviour suggest a predisposition to interspecific cooperation. However, the importance of dominance in human understanding of social systems has tended to overemphasise its importance in the human-horse relationship. The evolving horse-human relationship from predation to companionship, has resulted in serial conflicts of interest for equine and human participants. Only by understanding the nature and origin of these conflicts can ethologists encourage equine management practices which minimise deleterious effects on the behaviour of the horse.
	Address
	Corporate Author				Thesis
	Publisher	American Medical Association (AMA)	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0425-1644	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6714
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	Author	Chaudhuri, M.; Ginsberg, J.R.
	Title	*Urinary androgen concentrations and social status in two species of free ranging zebra (Equus burchelli* and E. grevyi).**			Type	Journal Article
	Year	1990	Publication	Journal of Reproduction and Fertility	Abbreviated Journal	J Reprod Fert
	Volume	88	Issue		Pages	127-133
	Keywords	zebra; testosterone; androgens; territoriality; social behaviour; dominance
	Abstract	In both species of zebra, breeding males had higher urinary androgen concentrations (ng androgens/mg Cr) than did non-breeding bachelor males (30.0 +/- 5.0 (N = 9) versus 11.4 +/- 2.8, (N = 7) in the plains zebra; 19.0 +/- 2.2 (N = 17) versus 10.7 +/- 1.2 (N = 14) in the Grevy's zebra). In the more stable family structure of the plains zebra (single male non-territorial groups) variations in androgen concentrations could not be ascribed to any measured variable. In the Grevy's zebra, androgen values were significantly lower in samples taken from territorial (breeding) males which had temporarily abandoned their territories (N = 4) and the urinary androgen concentration for a male on his territory was negatively correlated with the time since females last visited the territory.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes	from Professor Hans Klingels Equine Reference List			Approved	yes
	Call Number				Serial	987
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	Author	Bahloul, K.; Pereladova, O.B.; Soldatova, N.; Fisenko, G.; Sidorenko, E.; Sempere, A.J.
	Title	Social organization and dispersion of introduced kulans (Equus hemionus kulan) and Przewalski horses (Equus przewalski) in the Bukhara Reserve, Uzbekistan			Type	Journal Article
	Year	2001	Publication	Journal of Arid Environments	Abbreviated Journal	J. Arid. Environ.
	Volume	47	Issue	3	Pages	309-323
	Keywords	Przewalski horses; kulans; Central Asia; home range; behaviour
	Abstract	Asiatic wild asses and Przewalski horses initially inhabited steppe, semi-desert and desert areas, but Przewalski horses became extinct in the wild, and kulans disappeared at the beginning of the 20th century, except for a small population in Turkmenistan. The Bukhara Breeding Centre (Uzbekistan) was created in 1976 for reintroduction and conservation of wild ungulate species. In 1977-1978, five kulans (two males and three females), from Barsa-Kelmes island on the Aral sea, were introduced into the reserve. The group increased to 25-30 animals in 1989-1990, when eight Przewalski horses from Moscow and St Petersburg zoos were introduced. We analysed the home ranges, preferred habitats and social interactions of these closely related species during 1995-1998 by seasonal and group composition. Horses and asses formed a reproductive group and a secondary non-reproductive group. The home range of the secondary group was larger than the reproductive group and seemed to be less dependent from the watering places. Przewalski horses were less adapted to semi-desert conditions (both water and vegetation needs) than kulan.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	777
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	Author	Baragli, P.; Vitale, V.; Paoletti, E.; Sighieri, C.; Reddon, A.R.
	Title	Detour behaviour in horses (Equus caballus)			Type	Journal Article
	Year	2011	Publication	Journal of Ethology	Abbreviated Journal	J. Ethol.
	Volume	29	Issue	2	Pages	227-234
	Keywords	Detour behaviour; Equus caballus; Horses; Lateralization; Spatial reasoning
	Abstract	The objective of this study was to investigate the ability of horses (Equus caballus) to detour around symmetric and asymmetric obstacles. Ten female Italian saddle horses were each used in three detour tasks. In the first task, the ability to detour around a symmetrical obstacle was evaluated; in the second and third tasks subjects were required to perform a detour around an asymmetrical obstacle with two different degrees of asymmetry. The direction chosen to move around the obstacle and time required to make the detour were recorded. The results suggest that horses have the spatial abilities required to perform detour tasks with both symmetric and asymmetric obstacles. The strategy used to perform the task varied between subjects. For five horses, lateralized behaviour was observed when detouring the obstacle; this was consistently in one direction (three on the left and two on the right). For these horses, no evidence of spatial learning or reasoning was found. The other five horses did not solve this task in a lateralized manner, and a trend towards decreasing lateralization was observed as asymmetry, and hence task difficulty, increased. These non-lateralized horses may have higher spatial reasoning abilities.
	Address
	Corporate Author				Thesis
	Publisher	Springer Japan	Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0289-0771	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5686
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	Author	Benz, B.; Münzing, C.; Krueger, K.; Winter, D.
	Title	Ethologische Untersuchung von Heuraufen in der Pferdehaltung [Ethological investigation of hayracks in equine husbandry]			Type	Journal Article
	Year	2014	Publication	Landtechnik	Abbreviated Journal	Landtechnik
	Volume	69	Issue	5	Pages	239-244
	Keywords	Pferdehaltung, Fressverhalten, Futterraufen [horse keeping, feeding behaviour, roughage racks]
	Abstract	Eine tiergerechte, physiologisch und anatomisch auf die Bedürfnisse der Pferde ausgerichtete Raufutterversorgung sollte die Kaubedürfnisse und die Beschäftigungszeiten von Pferden ausreichend berücksichtigen. Daher – und auch aufgrund des bestehenden Kostendrucks bei Raufutter – steigt das Interesse an Raufutterraufen, durch die möglicherweise die Futteraufnahmezeiten verlängert sowie Futterverluste minimiert werden können. Die vorliegende Untersuchung vergleicht das Fressverhalten und die Körperhaltung von acht Pferden beim Einsatz von drei unterschiedlichen Futterraufen mit der Bodenfütterung in Einzelboxenhaltung. Die Ergebnisse dieser Studie lassen den Schluss zu, dass der Einsatz von Raufutterraufen die Futteraufnahmezeiten verlängert und somit längere Beschäftigungszeiten für die Futteraufnahme gewährleistet werden. Außerdem konnte festgestellt werden, dass die Pferde das Raufutter bei einer der drei untersuchten Raufen überwiegend mit natürlicher Kopf- Hals-Haltung aufnahmen. [Regarding the species horse, an appropriate supply of roughage should take into account the need to chew as well as the need for occupation. In this context, and due to the current cost pressure for hay, the interest in roughage racks increases. It is assumed that roughage racks could help to extend the feeding time and reduce food losses. The present study places the emphasis on the observation of the feeding behaviour of eight horses in single horse boxes. Three different roughage racks are compared to traditional feeding on the floor. On the basis of the results it may be concluded that the use of roughage racks extends the feeding time and thus ensures longer occupation. In one of the three roughage racks investigated the horses mainly eat in a natural posture of their head and neck.]
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5840
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	Author	Górecka-Bruzda, A.; Chruszczewski, M.H.; Jezierski, T.; Murphy, J.
	Title	Behaviour-related traits of Polish sport horse stallions participating in 100-day performance tests			Type	Journal Article
	Year		Publication	Livestock Science	Abbreviated Journal	Livest Sci
	Volume	In Press, Corrected Proof	Issue		Pages
	Keywords	Horse; 100-day performance tests; Behaviour; Temperament; Factor Analysis
	Abstract	Sport horse studbooks frequently use performance tests to identify, evaluate and approve stallions for future breeding programmes. The aim of this study was to analyse behaviour-related traits of Polish Warmblood stallions participating in studbook 100-day performance tests. Both relatedness and differences of traits and their contribution to overall variability were analysed. Three-year-old stallions (n = 374) participated in 100-day performance test programmes at two Polish training stations from 2004 to 2008. Traits including character, temperament and trainability (trainer rated), free-jumping, jumping under the rider, walk, trot and canter (independently rated by both trainer and the selection jury) and jumping ability, dressage-ability and rideability (rated by the independent test rider) were subjected to Factor Analysis (FA) with varimax rotation. The FA produced three factors [Cooperation (Cp), Jumping Potential (JP) and Dressage Potential (DP)] with eigenvalues > 1, which accounted for 64.6% of the total variance of the traits examined. After rotation, the factors represented Cp: 23.2.0%, JP: 21.5% and DP: 19.8% of variance respectively. The traits of character, temperament, trainability, jumping under the rider, rideability, dressage and jumping abilities had high loadings on the second factor (Cp) and thereafter, the assessment of all behaviour-related traits contributed to the general evaluation with approximately one-fifth of total variability. The findings may indicate that successful sport careers for any horse may be much more closely associated with willingness to cooperate with the rider and riding and handling behaviour, than to the horse's actual physical performance potential. It would appear that only so-called sport-suitable performance of the horse is being selected for in performance tests (during jumping and dressage suitability testing). This approach may be short-sighted and inadequate with serious implications for horse under-performance in leisure activities notwithstanding general horse welfare within the disciplines of equitation.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1871-1413	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5281
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	Author	Cooper, J.J.; Albentosa, M.J.
	Title	Behavioural adaptation in the domestic horse: potential role of apparently abnormal responses including stereotypic behaviour			Type	Journal Article
	Year	2005	Publication	Livestock Production Science	Abbreviated Journal	Livest. Prod. Sci.
	Volume	92	Issue	2	Pages	177-182
	Keywords	Behavioural adaptation; Horse; Stereotypic behaviour
	Abstract	Classically, biologists have considered adaptation of behavioural characteristics in terms of long-term functional benefits to the individual, such as survival or reproductive fitness. In captive species, including the domestic horse, this level of explanation is limited, as for the most part, horses are housed in conditions that differ markedly from those in which they evolved. In addition, an individual horse's reproductive fitness is largely determined by man rather than its own behavioural strategies. Perhaps for reasons of this kind, explanations of behavioural adaptation to environmental challenges by domestic animals, including the capacity to learn new responses to these challenges, tend to concentrate on the proximate causes of behaviour. However, understanding the original function of these adaptive responses can help us explain why animals perform apparently novel or functionless activities in certain housing conditions and may help us to appreciate what the animal welfare implications might be. This paper reviews the behavioural adaptation of the domestic horse to captivity and discusses how apparently abnormal behaviour may not only provide a useful practical indicator of specific environmental deficiencies but may also serve the animal as an adaptive response to these deficiencies in an “abnormal” environment.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0301-6226	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4829
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