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Author Søndergaard, E.; Jensen, M.B.; Nicol, C.J. url  doi
openurl 
  Title Motivation for social contact in horses measured by operant conditioning Type Journal Article
  Year Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume In Press, Corrected Proof Issue Pages (up)  
  Keywords Horse; Housing; Social behaviour; Operant conditioning; Motivation  
  Abstract Although horses are social animals they are often housed individually with limited social contact to other horses and this may compromise their welfare. The present study included eight young female horses and investigated the strength of motivation for access to full social contact, head contact and muzzle contact, respectively, to a familiar companion horse. Horses were housed individually next to their companion horse and separations between pens prevented physical contact. During daily test sessions horses were brought to a test area where they could access an arena allowing social contact. Arena access during 3 min was given after completion of a predetermined number of responses on a panel. Fixed ratios (FR) of 8, 16, 24, 32 and 40 responses per arena access were applied in a random order, one per daily test session, within each test week (Monday to Friday), and the number of rewards per daily test session was recorded. All horses could access all three types of social contact in a cross-over design, and an empty arena was used as control. Motivational strength was assessed using elasticity of demand functions, which were estimated based on the number of rewards earned and FR. Elasticities of demand for the three types of social contact were low (-0.20), and not significantly different, although increasing FR still resulted in a decrease in rewards obtained for all three types of social contact (P < 0.001). Across FR-levels horses earned more rewards for social contact than for an empty arena, as shown by much higher intercept values (2.51 vs. 0.99; P < 0.001). However, the elasticity of demand for infrequent access to an empty arena (-0.08) was lower than for social contact (P < 0.01) and not significantly different from zero (P = 0.07). Horses performed more social behaviour the lesser the restriction on social contact (full > head > muzzle). However, the finding that horses showed a similar and high motivation for all three types of social contact suggests that they are valued equally highly in a situation where the alternative is no social contact.  
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  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5388  
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Author Górecka-Bruzda, A.; Jastrzebska, E.; Sosnowska, Z.; Jaworski, Z.; Jezierski, T.; Chruszczewski, M.H. url  doi
openurl 
  Title Reactivity to humans and fearfulness tests: Field validation in Polish Cold Blood Horses Type Journal Article
  Year Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume In Press, Corrected Proof Issue Pages (up)  
  Keywords Cold blood horse; Temperament; Reactivity; Behavioural tests; Questionnaire; Validity  
  Abstract The aim of the present paper was to examine in field conditions the feasibility of the experimentally developed behavioural tests for assessment of two temperamental traits of cold blood horses: fearfulness and reactivity to humans. The predictive, convergent and discriminant validity of the tests was evaluated. Fifty-three Polish Cold Blood Horses (CBHs) were tested with the battery of behavioural tests involving passive and active contact with humans, handling, startling, novel object and surface tests. The behaviour of horses in real, on-farm conditions was evaluated with the caretaker's questionnaire (CQ). The tests demonstrated that a non-startling object and unforced human manipulations were willingly accepted by horses. In contrast, the horses were more reluctant to interact spontaneously with humans and showed longer persistence of a withdrawal reaction when presented with a startling stimulus or when restrained. CQ confirmed the common opinion of calmness, patience and good manageability of the CBHs. The results of behavioural tests were correlated with the scores derived from factor analysis of the CQ. Moreover, the majority of tests measures were correlated within reactivity to humans (median rs = 0.56) and fear tests (median rs = 0.41). Thus, predictive and convergent validity of the employed tests has been confirmed. However, reactivity to humans and fear measures were also correlated (median rs = 0.36), therefore discriminant validity of the tests used has been not confirmed. Therefore, we suggest that the parameters of tests designed to measure two separate traits: reactivity to humans and fearfulness, measured in fact avoidance of potential harm and withdrawal behaviour. Although selection for the work-appropriate behaviour is almost nonexistent nowadays, we conclude that Polish Cold Blood Horses are still predisposed for disciplines of equitation and driving, which demand an easy to handle horse. Nonetheless, we recommend behavioural testing of horses when approving for breeding. For this purpose, we can confirm usefulness of the experimentally developed tests in field assessment of general behavioural reactivity of horses.  
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  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5395  
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Author Birke, L.; Hockenhull, J.; Creighton, E.; Pinno, L.; Mee, J.; Mills, D. url  doi
openurl 
  Title Horses' responses to variation in human approach Type Journal Article
  Year Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume In Press, Corrected Proof Issue Pages (up)  
  Keywords Horse; Flight response; Human approach; Body posture; Approach speed; Natural horsemanship  
  Abstract The behaviour of humans around horses is thought to have a substantial impact on how people are perceived in subsequent interactions and many horse trainers give detailed advice on how handlers should behave when initially approaching a loose horse. Here we report on three studies designed to explore the effect of different human approach styles on the behaviour of naïve and experienced horses. In the first study, the change in flight distance (distance at which horses started to avoid an approaching human) of twelve semi-feral Dartmoor ponies, undergoing training to allow handling, was assessed. Over the 10 handling sessions median flight distance decreased significantly (p < 0.001) from 2.38 m to 0.00 m and there was a significant positive shift in the ponies' behaviour following the appearance of the researcher (p = 0.002). In a second study the effect of a direct (vigorous, swinging a lead rope and with eye contact) versus indirect (relaxed, no rope swinging and without eye contact) approach style was assessed on six adult experienced riding horses. The mean flight distance during a direct approach style (6.87 m) was significantly greater than that which occurred during an indirect approach style (2.32 m). Direction of approach was not found to significantly affect flight distance. In a third study, the effect of the rope was removed and a similar method to the second study applied to a group of naïve, feral ponies. The effect of different components of approach style, speed of approach, handler body posture and direction of gaze, which might contribute to observed differences in behavioural responses, were then examined systematically in this population. This revealed no significant difference in mean flight distance between the two approach styles (2.28 m indirect versus 2.37 m direct approach), but ponies were significantly more likely to move off in trot (p = 0.025) and to travel further (p = 0.001) when a direct approach was used. Speed of approach was the most salient factor, with a fast approach increasing both the tendency to move off in trot (p < 0.001) and distance travelled (p < 0.001). Body posture (relaxed or tense) had no effect, while flight distance was significantly greater when the person was looking away (p = 0.045). These results suggest horses may have an important egocentric spatial barrier, which perhaps relates to personal space and triggering of the flight response. Contrary to popular belief, body posture did not appear to be very important in the contexts examined unless accompanied by extraneous aids, while the speed of approach is particularly significant. These results are of important practical relevance in reducing the risk of injury, and the effective management of horses with minimal stress.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5401  
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Author Hartmann, E.; Søndergaard, E.; Keeling, L.J. doi  openurl
  Title Keeping horses in groups: A review Type Journal Article
  Year Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume Issue Pages (up)  
  Keywords Equine; Behaviour; Welfare; Housing; Social  
  Abstract Although husbandry conditions for horses have improved over the last decades, many horses are still kept singly with limited or no physical contact to other horses. This is surprising, given the fact that keeping horses in groups is recognised best to fulfil their physical and behavioural needs, especially their need for social contact with conspecifics, as well as to have a beneficial effect on horse–human interactions during training. Group housing of farm animals is widely applied in practice. As a consequence, scientists have investigated numerous aspects of group housing to help further improve animal welfare and human–animal interactions under these conditions. However, compared to this literature available in farm animals, and the plentiful studies conducted of feral horse populations, there is much less done when it comes to the management of horses kept in groups in the domestic environment. In particular, limited scientific information is available into the effect of group size and group composition on behaviour and methods of introducing new horses into established groups, even though problems related to social integration are repeatedly taken as arguments against keeping horses in groups. This review, therefore, aims to provide an overview of the current scientific knowledge regarding keeping horses in groups. Furthermore, it aims to give insight into whether or not some of the concerns related to keeping horses in groups are justified and to review scientifically based solutions that could be useful in practice to improve horse welfare and human safety.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5490  
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Author Rocha, A.D. de L.; Menescal-de-Oliveira, L.; da Silva, L.F.S. url  doi
openurl 
  Title Effects of human contact and intra-specific social learning on tonic immobility in guinea pigs, Cavia porcellus Type Journal Article
  Year 2017 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume Issue Pages (up)  
  Keywords Cohabitation; Fear; Motor response; Defensive behaviour; Predator-prey  
  Abstract Abstract Social learning is the capacity of animals to acquire adaptive information from others. In the case of fear responses, animals can learn fearful or non-fearful responses by observing the behavior of conspecifics. Tonic immobility (TI) is an anti-predatory behavior elicited during intense fear situations. Studies have revealed that regular contact with humans can reduce TI responses in animals. In our study, we evaluated the effect of human contact on the TI responses in guinea pigs. We also evaluated the effect of cohabitation (non-fearful animals with fearful animals) on their TI responses. To achieve this, we measured the TI responses induced by postural inversion and restraint in guinea pigs as a result of different treatments. In our first experiment, we determined the effect of human contact on TI responses by establishing 3 treatment groups: no contact, handled, and habituated. In our second experiment, we addressed the effect of social learning on TI response by testing TI response in habituated, and unhabituated animals that had cohabitated for 10 days. In the first experiment, 10 days of either handling or habituation did not prevent TI in guinea pigs, but habituation did increase latency [F(2,119) = 14.19; p < 0.0001] and handling or habituation decrease duration [F(2,119) = 15.01; p < 0.0001] of the TI behavior in the guinea pigs. In the second experiment, the cohabitation of unhabituated and habituated animals reduced TI duration [F(2,93) = 5.058; p < 0.008]. These data suggest that both forms of human interaction can reduce experimenter fear in guinea pigs. It therefore seems that unhabituated guinea pigs learn not to fear the experimenter by cohabitating with habituated guinea pigs.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6133  
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Author Araba, B.D.; Crowell-Davis, S.L. doi  openurl
  Title Dominance relationships and aggression of foals (Equus caballus) Type Journal Article
  Year 1994 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 41 Issue 1-2 Pages (up) 1-25  
  Keywords aggression; dominance; horse; Equus caballus  
  Abstract Studied a herd of 15 Belgian brood-mares and 10 foals. Specific aspects of social structure studied were dominance-subordinance relationships, preferred associates, social spacing, aggression rates, the frequency of aggressions administered down the dominance hierarchy, and interactive play bouts. The rank order of the foals, both before and after weaning, was positively correlated with the rank order of their dams. There was also a significant relationship between a foal's rank and its total aggression or aggression rate per subordinate post-weaning. Higher ranking foals had higher rates of aggression. Over 80% of threats were directed down the dominance hierachy. The play-rank order of the foals, scored by the number of times foal left a play bout, was not significantly correlated with the rank order as scored by agonistic interactions. -from Authors  
  Address Dept Anatomy and Radiology, Univ of Georgia, Athens, GA, USA  
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  Series Volume Series Issue Edition  
  ISSN 01681591 (Issn) ISBN Medium  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 790  
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Author Houpt, K.A.; Fraser, A.F. url  doi
openurl 
  Title Przewalski horses Type Journal Article
  Year 1988 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 21 Issue 1-2 Pages (up) 1-3  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 2302  
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Author Caanitz, H.; O'Leary, L.; Houpt, K.; Petersson, K.; Hintz, H. url  openurl
  Title Effect of exercise on equine behavior Type Journal Article
  Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 31 Issue 1-2 Pages (up) 1-12  
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  Abstract The effect of short periods of strenuous exertion, in this case treadmill exercise, on the subsequent behavior of Standardbred horses was examined. Six horses were exercised on a high-speed treadmill 4 or 5 days per week, for 3-4 miles (approximately 1.8 m s-1 for 3 min, 5 m s-1 for 12 min, 9 m s-1 for 3 min, 3 m s-1 for 3 min, 1.8 m s-1 for 3 min). The behavior of the horses was observed in the horse's home stall immediately after exercise and 2-7 h after exercise. Focal animal sampling for a total of 150 h revealed that the horses spent significantly more time drinking and less time resting after exercise than they did on control (non-exercise or rest days). The greatest influence on behavior was seen immediately after exercise. The horses spent 13.2+/-2.7 s per 15 min drinking after exercise and 7.2+/-2.3 s per 15 min drinking on non-exercise days. They spent 7.3+/-1.5 min h-1 stand resting after exercise and 9.7+/-2.1 min h-1 on non-exercise days. These changes in behavior may be related to the physiological changes that accompany exercise. Eating, walking, elimination and self-grooming were not significantly influenced by exercise. In a second experiment the activities of two groups of six Standardbred mares were compared. One group was exercised on the treadmill and the other was not. The exercised horses spent more time drinking and lying, but urinated less than the non-exercised group.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 1989  
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Author Berger, A.; Scheibe, K.-M.; Eichhorn, K.; Scheibe, A.; Streich, J. url  doi
openurl 
  Title Diurnal and ultradian rhythms of behaviour in a mare group of Przewalski horse (Equus ferus przewalskii), measured through one year under semi-reserve conditions Type Journal Article
  Year 1999 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 64 Issue 1 Pages (up) 1-17  
  Keywords Horse; Przewalski horse; Circadian rhythm; Ultradian rhythm; Feeding and nutrition; Telemetry; Stress  
  Abstract Investigations were conducted on four horses from a group of 12 Przewalski mares raised in different zoos and kept in a 44-ha enclosure under semi-natural conditions. Activity and feeding were continuously measured every second and were saved every 15 min by the storage-telemetry system ETHOSYS, from June 1995 to July 1996. Body mass of the horses was regularly recorded. Daily and monthly mean values, power spectra and DFC (as a measure for stability of rhythms synchronised with circadiurnal period) for activity and feeding were calculated. The general pattern of activity and feeding over the year was closely related to sunrise and sunset. Feeding accounted for 40% of total activity in summer and 62% in spring (all-year average being 52%). The level of activity was lowest in winter; whereas feeding was lowest in summer. The time budget for feeding reflected both feeding conditions and the annual pattern of body condition. Greatest activity occurred during daylight hours. Only on hot summer days, activity at night was higher than during daylight hours. Spectral analysis of activity and feeding in Przewalski horse showed a time pattern which was characterised by 24-h rhythmicity, but also by ultradian components with period lengths between 4.8 and 12 h, i.e., an activity pattern of up to five strong bouts per day. Annual variation in the pattern of power spectra was not high during the year. Results are discussed in connection with horse feeding strategy. Analysing the time structure of long-term and continuously measured activity and feeding could be a useful method to follow the general living conditions, especially the nutritional situation and to detect stressful conditions.  
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  Call Number Equine Behaviour @ team @ Serial 2236  
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Author Pickerel, T.M.; Crowell-Davis, S.L.; Caudle, A.B.; Estep, D.Q. url  openurl
  Title Sexual preference of mares (Equus caballus) for individual stallions Type Journal Article
  Year 1993 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 38 Issue 1 Pages (up) 1-13  
  Keywords Horse; Sexual behavior; Sexual preference; Vocalization  
  Abstract Eight mares were tested to determine if they remained near one of two stallions longer than would be expected if association was random. Six stallions were paired in 30 combinations and each mare was tested 30 times. The mares (Equus caballus) demonstrated a definite preference for individual stallions throughout the breeding season. This preference was influenced by the estrous state of the mare. During estrus, mares' preferences for stallions were positively correlated with the rate at which a given stallion vocalized. During diestrus, mares spent significantly less time in the proximity of stallions and did not exhibit any preference for individual stallions.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 2270  
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