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	Author	Burke, D.; Cieplucha, C.; Cass, J.; Russell, F.; Fry, G.
	Title	Win-shift and win-stay learning in the short-beaked echidna (Tachyglossus aculeatus)			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	2	Pages	79-84
	Keywords	Animals; Echidna/psychology; Ecology; Female; Learning; Memory; Predatory Behavior; Reinforcement (Psychology)
	Abstract	Numerous previous investigators have explained species differences in spatial memory performance in terms of differences in foraging ecology. In three experiments we attempted to extend these findings by examining the extent to which the spatial memory performance of echidnas (or “spiny anteaters”) can be understood in terms of the spatio-temporal distribution of their prey (ants and termites). This is a species and a foraging situation that have not been examined in this way before. Echidnas were better able to learn to avoid a previously rewarding location (to “win-shift”) than to learn to return to a previously rewarding location (to “win-stay”), at short retention intervals, but were unable to learn either of these strategies at retention intervals of 90 min. The short retention interval results support the ecological hypothesis, but the long retention interval results do not.
	Address	Department of Psychology, University of Wollongong, Wollongong, NSW 2522, Australia. darren_burke@uow.edu.au
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12150039			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2605
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	Author	Emery, N.J.; Dally, J.M.; Clayton, N.S.
	Title	Western scrub-jays ( Aphelocoma californica) use cognitive strategies to protect their caches from thieving conspecifics			Type	Journal Article
	Year	2004	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	7	Issue	1	Pages	37-43
	Keywords	Animals; Birds/physiology; Feeding Behavior/physiology; Female; Food; Male; Memory
	Abstract	Food caching birds hide food and recover the caches when supplies are less abundant. There is, however, a risk to this strategy because the caches are susceptible to pilfering by others. Corvids use a number of different strategies to reduce possible cache theft. Scrub-jays with previous experience of pilfering other's caches cached worms in two visuospatially distinct caching trays either in private or in the presence of a conspecific. When these storers had cached in private, they subsequently observed both trays out of reach of a conspecific. When these storers had cached in the presence of a conspecific, they subsequently watched the observer pilfering from one of the trays while the other tray was placed in full view, but out of reach. The storers were then allowed to recover the remaining caches 3 h later. Jays cached more worms when they were observed during caching. At the time of recovery, they re-cached more than if they had cached in private, selectively re-caching outside of the trays in sites unbeknown to potential thieves. In addition, after a single pilfering trial, the jays switched their recovery strategy from predominantly checking their caches (i.e. returning to a cache site to see whether the food remained there) to predominantly eating them. Re-caching remained constant across the three trials. These results suggest that scrub-jays use flexible, cognitive caching and recovery strategies to aid in reducing potential future pilfering of caches by conspecifics.
	Address	Sub-department of Animal Behaviour, University of Cambridge, High Street, CB3 8AA Madingley, Cambs, UK. nje23@cam.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12827547			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2566
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	Author	Shettleworth, S.J.
	Title	Varieties of learning and memory in animals			Type	Journal Article
	Year	1993	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	19	Issue	1	Pages	5-14
	Keywords	Animals; Association Learning; Birds; Conditioning, Classical; Evolution; Imprinting (Psychology); Learning; Memory; Social Environment; Species Specificity; Taste
	Abstract	It is often assumed that there is more than one kind of learning--or more than one memory system--each of which is specialized for a different function. Yet, the criteria by which the varieties of learning and memory should be distinguished are seldom clear. Learning and memory phenomena can differ from one another across species or situations (and thus be specialized) in a number of different ways. What is needed is a consistent theoretical approach to the whole range of learning phenomena, and one is explored here. Parallels and contrasts in the study of sensory systems illustrate one way to integrate the study of general mechanisms with an appreciation of species-specific adaptations.
	Address	Department of Psychology, University of Toronto, Ontario, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:8418217			Approved	no
	Call Number	refbase @ user @			Serial	380
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	Author	Zentall, T.R.
	Title	Timing, memory for intervals, and memory for untimed stimuli: the role of instructional ambiguity			Type	Journal Article
	Year	2005	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	70	Issue	3	Pages	209-222
	Keywords	Animals; Attention; Columbidae; Discrimination Learning; Memory, Short-Term; Practice (Psychology); Reinforcement Schedule; Retention (Psychology); *Time Perception
	Abstract	Theories of animal timing have had to account for findings that the memory for the duration of a timed interval appears to be dramatically shorted within a short time of its termination. This finding has led to the subjective shortening hypothesis and it has been proposed to account for the poor memory that animals appear to have for the initial portion of a timed interval when a gap is inserted in the to-be-timed signal. It has also been proposed to account for the poor memory for a relatively long interval that has been discriminated from a shorter interval. I suggest here a simpler account in which ambiguity between the gap or retention interval and the intertrial interval results in resetting the clock, rather than forgetting the interval. The ambiguity hypothesis, together with a signal salience mechanism that determines how quickly the clock is reset at the start of the intertrial interval can account for the results of the reported timing experiments that have used the peak procedure. Furthermore, instructional ambiguity rather than memory loss may account for the results of many animal memory experiments that do not involve memory for time.
	Address	Department of Psychology, University of Kentucky, 202B Kastle Hall, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16095851			Approved	no
	Call Number	refbase @ user @			Serial	222
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	Author	Treichler, F.R.
	Title	Successive reversal of concurrent discriminations by macaques (Macaca mulatta): proactive interference effects			Type	Journal Article
	Year	2005	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	2	Pages	75-83
	Keywords	Animals; Choice Behavior; Discrimination Learning; Female; Macaca mulatta/psychology; Memory; Proactive Inhibition; Random Allocation; *Reversal Learning
	Abstract	Rhesus monkeys received concurrent within-session training on eight, two-choice object pairs and then underwent successive reversals of these problems. Initially, reversals required about six times more training than acquisition with no improvement over seven successive reversals. Surprisingly, performance on these eight problems was unimpaired if they were embedded in different eight-problem tasks, thereby indicating a release from proactive interference. When the original eight problems again underwent successive reversal, no improvement was seen over seven reversals, although there was significantly less error-per-reversal than in the initial test. Subsequently, monkeys appeared to be developing a learning set for successive reversal because performance on successive reversal of eight novel problems was not different from that seen with the old familiar task. Set acquisition was confirmed when proficient reversal was eventually achieved on both old and new concurrent tasks. Thus, “concurrent reversal set” did develop, but it required arduous training to overcome proactive interference effects on memory. The ubiquitous influence of measurement context on organization of monkey memory was noted.
	Address	Kent State University, Kent, Ohio 44242, USA. rtreichl@kent.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15365875			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2512
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	Author	Schwartz, B.L.; Colon, M.R.; Sanchez, I.C.; Rodriguez, I.A.; Evans, S.
	Title	Single-trial learning of “what” and “who” information in a gorilla (Gorilla gorilla gorilla): implications for episodic memory			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	2	Pages	85-90
	Keywords	Animals; Cognition; Gorilla gorilla/psychology; Learning; Male; *Memory; Perception; Reinforcement Schedule
	Abstract	Single-trial learning and long-term memory of “what” and “who” information were examined in an adult gorilla (Gorilla gorilla gorilla). We presented the gorilla with a to-be-remembered food item at the time of study. In Experiment 1, following a retention interval of either approximately 7 min or 24 h, the gorilla responded with one of five cards, each corresponding to a particular food. The gorilla was accurate on 70% of the short retention-interval trials and on 82% of the long retention-interval trials. In Experiment 2, the food stimulus was provided by one of two experimenters, each of whom was represented by a card. The gorilla identified the food (55% of the time) and the experimenter (82% of the time) on the short retention-interval trials. On the long retention-interval trials, the gorilla was accurate for the food (73%) and for the person (87%). The results are interpreted in light of theories of episodic memory.
	Address	Department of Psychology, Florida International University, University Park, Miami, FL 33199, USA. schwartb@fiu.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12150040			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2604
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	Author	Hampton, R.R.; Zivin, A.; Murray, E.A.
	Title	Rhesus monkeys (Macaca mulatta) discriminate between knowing and not knowing and collect information as needed before acting			Type	Journal Article
	Year	2004	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	7	Issue	4	Pages	239-246
	Keywords	Animals; Association Learning; Awareness; Choice Behavior; Concept Formation; Discrimination Learning; Female; Macaca mulatta/psychology; Male; *Memory, Short-Term; Observation
	Abstract	Humans use memory awareness to determine whether relevant knowledge is available before acting, as when we determine whether we know a phone number before dialing. Such metacognition, or thinking about thinking, can improve selection of appropriate behavior. We investigated whether rhesus monkeys ( Macaca mulatta) are capable of a simple form of metacognitive access to the contents of short-term memory. Monkeys chose among four opaque tubes, one of which concealed food. The tube containing the reward varied randomly from trial to trial. On half the trials the monkeys observed the experimenter baiting the tube, whereas on the remaining trials their view of the baiting was blocked. On each trial, monkeys were allowed a single chance to select the tube containing the reward. During the choice period the monkeys had the opportunity to look down the length of each tube, to determine if it contained food. When they knew the location of the reward, most monkeys chose without looking. In contrast, when ignorant, monkeys often made the effort required to look, thereby learning the location of the reward before choosing. Looking improved accuracy on trials on which monkeys had not observed the baiting. The difference in looking behavior between trials on which the monkeys knew, and trials on which they were ignorant, suggests that rhesus monkeys discriminate between knowing and not knowing. This result extends similar observations made of children and apes to a species of Old World monkey, suggesting that the underlying cognitive capacities may be widely distributed among primates.
	Address	Section on the Neurobiology of Learning and Memory, Laboratory of Neuropsychology, National Institute of Mental Health, NIH, Bethesda, MD 20892-4415, USA. robert@ln.nimh.nih.gov
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15105996			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2525
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	Author	Urcuioli, P.J.; Zentall, T.R.
	Title	Retrospective coding in pigeons' delayed matching-to-sample			Type	Journal Article
	Year	1986	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	12	Issue	1	Pages	69-77
	Keywords	Animals; Color Perception; Columbidae; Discrimination Learning; Form Perception; Memory; Mental Recall; Orientation; Pattern Recognition, Visual; Retention (Psychology)
	Abstract	In this study we examined how coding processes in pigeons' delayed matching-to-sample were affected by the stimuli to be remembered. In Experiment 1, two groups of pigeons initially learned 0-delay matching-to-sample with identical comparison stimuli (vertical and horizontal lines) but with different sample stimuli (red and green hues or vertical and horizontal lines). Longer delays were then introduced between sample offset and comparison onset to assess whether pigeons were prospectively coding the same events (viz., the correct line comparisons) or retrospectively coding different events (viz., their respective sample stimuli). The hue-sample group matched more accurately and showed a slower rate of forgetting than the line-sample group. In Experiment 2, pigeons were trained with either hues or lines as both sample and comparison stimuli, or with hue samples and line comparisons or vice versa. Subsequent delay tests revealed that the hue-sample groups remembered more accurately and generally showed slower rates of forgetting than the line-sample groups. Comparison dimension had little or no effect on performance. Together, these data suggest that pigeons retrospectively code the samples in delayed matching-to-sample.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3701260			Approved	no
	Call Number	refbase @ user @			Serial	263
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	Author	Treichler, F.R.; Van Tilburg, D.
	Title	Premise-pair training for valid tests of serial list organization in macaques			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	2	Pages	97-105
	Keywords	Animals; Discrimination Learning; Female; Macaca/psychology; *Memory
	Abstract	This study evaluated the role of several different training procedures on (1) efficiency of acquisition and (2) organizational characteristics of memory for lists that could be serially ordered. Five macaque monkeys were trained via two-choice object discriminations in a formboard apparatus on several five-item-series tasks that provided different levels of intrasession conditionality. Although ease of acquisition differed for subsets of the constituent pairs, concurrent inclusion of the four premise pairs that defined a list required equivalent amounts of training on every task. All training procedures yielded similar retention-test performances and showed common organizational properties (on both error and latency measures) consistent with the view that lists were retained as internally represented ordered series. Test outcomes emphasized the need for integrated exposition of all concurrent conditional relationships to allow appropriate tests of serial organization. However, if given such training, the monkeys revealed integrated serial memory even though they had never seen many of the possible novel combinations of list items. In overview, their performances offered further definition of the procedures required for valid assessment of inferential properties in comparative cognition.
	Address	Department of Psychology, Kent State University, Kent, OH 44242-0001, USA. rtreichl@kent.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12150042			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2602
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	Author	Zentall, T.R.
	Title	Mental time travel in animals: a challenging question			Type	Journal Article
	Year	2006	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	72	Issue	2	Pages	173-183
	Keywords	Animals; Behavior, Animal; Columbidae; Concept Formation; Conditioning, Operant; Imagination; Memory; Mental Recall; Planning Techniques; Rats; Time Perception; Transfer (Psychology)
	Abstract	Humans have the ability to mentally recreate past events (using episodic memory) and imagine future events (by planning). The best evidence for such mental time travel is personal and thus subjective. For this reason, it is particularly difficult to study such behavior in animals. There is some indirect evidence, however, that animals have both episodic memory and the ability to plan for the future. When unexpectedly asked to do so, animals can report about their recent past experiences (episodic memory) and they also appear to be able to use the anticipation of a future event as the basis for a present action (planning). Thus, the ability to imagine past and future events may not be uniquely human.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16466863			Approved	no
	Call Number	refbase @ user @			Serial	218
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