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Gomez Alvarez, C.B.; Rhodin, M.; Bobber, M.F.; Meyer, H.; Weishaupt, M.A.; Johnston, C.; Van Weeren, P.R. |
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Title |
The effect of head and neck position on the thoracolumbar kinematics in the unridden horse |
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Journal Article |
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Year |
2006 |
Publication |
Equine Veterinary Journal. Supplement |
Abbreviated Journal |
Equine Vet J Suppl |
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Volume |
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36 |
Pages |
445-451 |
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Animals; Biomechanics; Head/*physiology; Horses/*physiology; Lumbar Vertebrae/physiology; Male; Neck/*physiology; Physical Conditioning, Animal/physiology; Posture/*physiology; Sports; Thoracic Vertebrae/physiology; Weight-Bearing |
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REASONS FOR PERFORMING STUDY: In many equestrian activities a specific position of head and/or neck is required that is dissimilar to the natural position. There is controversy about the effects of these positions on locomotion pattern, but few quantitative data are available. OBJECTIVES: To quantify the effects of 5 different head and neck positions on thoracolumbar kinematics of the horse. METHODS: Kinematics of 7 high level dressage horses were measured walking and trotting on an instrumented treadmill with the head and neck in the following positions: HNP2 = neck raised, bridge of the nose in front of the vertical; HNP3 = as HNP2 with bridge of the nose behind the vertical; HNP4 = head and neck lowered, nose behind the vertical; HNP5 = head and neck in extreme high position; HNP6 = head and neck forward and downward. HNP1 was a speed-matched control (head and neck unrestrained). RESULTS: The head and neck positions affected only the flexion-extension motion. The positions in which the neck was extended (HNP2, 3, 5) increased extension in the anterior thoracic region, but increased flexion in the posterior thoracic and lumbar region. For HNP4 the pattern was the opposite. Positions 2, 3 and 5 reduced the flexion-extension range of motion (ROM) while HNP4 increased it. HNP5 was the only position that negatively affected intravertebral pattern symmetry and reduced hindlimb protraction. The stride length was significantly reduced at walk in positions 2, 3, 4 and 5. CONCLUSIONS: There is a significant influence of head/neck position on back kinematics. Elevated head and neck induce extension in the thoracic region and flexion in the lumbar region; besides reducing the sagittal range of motion. Lowered head and neck produces the opposite. A very high position of the head and neck seems to disturb normal kinematics. POTENTIAL RELEVANCE: This study provides quantitative data on the effect of head/neck positions on thoracolumbar motion and may help in discussions on the ethical acceptability of some training methods. |
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Department of Equine Sciences, Utrecht University, Yalelaan 12, 3584 CM Utrecht, The Netherlands |
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PMID:17402464 |
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Equine Behaviour @ team @ |
Serial |
3702 |
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Author |
Weishaupt, M.A.; Wiestner, T.; von Peinen, K.; Waldern, N.; Roepstorff, L.; van Weeren, R.; Meyer, H.; Johnston, C. |
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Title |
Effect of head and neck position on vertical ground reaction forces and interlimb coordination in the dressage horse ridden at walk and trot on a treadmill |
Type |
Journal Article |
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Year |
2006 |
Publication |
Equine Veterinary Journal. Supplement |
Abbreviated Journal |
Equine Vet J Suppl |
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Volume |
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Issue |
36 |
Pages |
387-392 |
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Keywords |
Animals; Biomechanics; Exercise Test/instrumentation/methods/*veterinary; Forelimb/physiology; Gait; Head/physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Male; Neck/physiology; Physical Conditioning, Animal/methods/*physiology; Posture; Statistics, Nonparametric; Walking/*physiology |
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REASONS FOR PERFORMING STUDY: Little is known in quantitative terms about the influence of different head-neck positions (HNPs) on the loading pattern of the locomotor apparatus. Therefore it is difficult to predict whether a specific riding technique is beneficial for the horse or if it may increase the risk for injury. OBJECTIVE: To improve the understanding of forelimb-hindlimb balance and its underlying temporal changes in relation to different head and neck positions. METHODS: Vertical ground reaction force and time parameters of each limb were measured in 7 high level dressage horses while being ridden at walk and trot on an instrumented treadmill in 6 predetermined HNPs: HNP1 – free, unrestrained with loose reins; HNP2 – neck raised, bridge of the nose in front of the vertical; HNP3 – neck raised, bridge of the nose behind the vertical; HNP4 – neck lowered and flexed, bridge of the nose considerably behind the vertical; HNP5 – neck extremely elevated and bridge of the nose considerably in front of the vertical; HNP6 – neck and head extended forward and downward. Positions were judged by a qualified dressage judge. HNPs were assessed by comparing the data to a velocity-matched reference HNP (HNP2). Differences were tested using paired t test or Wilcoxon signed rank test (P<0.05). RESULTS: At the walk, stride duration and overreach distance increased in HNP1, but decreased in HNP3 and HNP5. Stride impulse was shifted to the forehand in HNP1 and HNP6, but shifted to the hindquarters in HNP5. At the trot, stride duration increased in HNP4 and HNP5. Overreach distance was shorter in HNP4. Stride impulse shifted to the hindquarters in HNP5. In HNP1 peak forces decreased in the forelimbs; in HNP5 peak forces increased in fore- and hindlimbs. CONCLUSIONS: HNP5 had the biggest impact on limb timing and load distribution and behaved inversely to HNP1 and HNP6. Shortening of forelimb stance duration in HNP5 increased peak forces although the percentage of stride impulse carried by the forelimbs decreased. POTENTIAL RELEVANCE: An extremely high HNP affects functionality much more than an extremely low neck. |
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Equine Hospital, University of Zurich, CH-8057 Zurich, Switzerland |
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PMID:17402453 |
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Equine Behaviour @ team @ |
Serial |
3704 |
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George, I.; Cousillas, H.; Richard, J.-P.; Hausberger, M. |
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Title |
Song perception in the European starling: hemispheric specialisation and individual variations |
Type |
Journal Article |
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Year |
2002 |
Publication |
Comptes Rendus Biologies |
Abbreviated Journal |
Compt. Rend. Biol. |
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Volume |
325 |
Issue |
3 |
Pages |
197-204 |
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Keywords |
lateralisation; perception; birdsong; starling; electrophysiology; individual variations; latéralisation; perception; chant; étourneaux; électrophysiologie; variations individuelles |
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Abstract |
Hemispheric specialisation for speech in humans has been well documented. The lateralisation for song production observed in songbirds is reminiscent of this hemispheric dominance. In order to investigate whether song perception is also lateralised, we made multiunit recordings of the neuronal activity in the field L of starlings during the presentation of species-specific and artificial non-specific sounds. We observed a systematic stronger activation in one hemisphere than in the other one during the playback of species-specific sounds, with inter-subject variability in the predominant hemisphere for song perception. Such an asymmetry was not observed for artificial non-specific sounds. Thus, our results suggest that, at least at the individual level, the two hemispheres of the starlings' brain perceive and process conspecific signals differently. |
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Equine Behaviour @ team @ |
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4636 |
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Author |
Seyfarth, R.M.; Cheney, D.L. |
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Title |
The acoustic features of vervet monkey grunts |
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Journal Article |
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Year |
1984 |
Publication |
The Journal of the Acoustical Society of America |
Abbreviated Journal |
J Acoust Soc Am |
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Volume |
75 |
Issue |
5 |
Pages |
1623-1628 |
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Keywords |
*Acoustics; Animals; Auditory Perception; Cercopithecus/*physiology; Cercopithecus aethiops/*physiology; Cues; Dominance-Subordination; Female; Male; Social Behavior; Sound Spectrography; *Vocalization, Animal |
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Abstract |
East African vervet monkeys give short (125 ms), harsh-sounding grunts to each other in a variety of social situations: when approaching a dominant or subordinate member of their group, when moving into a new area of their range, or upon seeing another group. Although all these vocalizations sound similar to humans, field playback experiments have shown that the monkeys distinguish at least four different calls. Acoustic analysis reveals that grunts have an aperiodic F0, at roughly 240 Hz. Most grunts exhibit a spectral peak close to this irregular F0. Grunts may also contain a second, rising or falling frequency peak, between 550 and 900 Hz. The location and changes in these two frequency peaks are the cues most likely to be used by vervets when distinguishing different grunt types. |
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0001-4966 |
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Notes |
PMID:6736426 |
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no |
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Call Number |
refbase @ user @ |
Serial |
703 |
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Author |
Ratzlaff, M.H.; Wilson, P.D.; Hyde, M.L.; Balch, O.K.; Grant, B.D. |
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Title |
Relationship between locomotor forces, hoof position and joint motion during the support phase of the stride of galloping horses |
Type |
Journal Article |
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Year |
1993 |
Publication |
Acta Anatomica |
Abbreviated Journal |
Acta Anat (Basel) |
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Volume |
146 |
Issue |
2-3 |
Pages |
200-204 |
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Keywords |
Animals; Equipment Design; Hoof and Claw/*physiology; Horses/*physiology; Joints/*physiology; *Locomotion; Motor Activity/*physiology; Physiology/instrumentation; *Posture; Shoes; Transducers |
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Abstract |
Three methods were used simultaneously to determine the relationships between the vertical forces exerted on the hooves and the positions of the limbs and hooves at the times of peak vertical forces from 2 horses galloping on a track straightaway. Vertical forces were recorded from an instrumented shoe, fetlock joint motion was measured with an electrogoniometer and the angles of the carpus, fetlock and hoof were determined from slow-motion films. At hoof contact, the mean angles of the carpus and fetlock were 181-182 degrees and 199-206 degrees, respectively. Peak vertical forces on the heel occurred at or near maximum extension of the carpal and fetlock joints. Peak forces on the toe occurred during flexion of the fetlock joint and at mean hoof angles of 28-31 degrees from the horizontal. The mean angles of the hoof from the horizontal at the time of heel contact were 6-7 degrees. Hoof lift occurred at mean carpal angles of 173-174 degrees and mean fetlock angles of 199-200 degrees. |
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Department of Veterinary and Comparative Anatomy, Pharmacology and Physiology, College of Veterinary Medicine, Washington State University, Pullman 99164-6520 |
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ISSN |
0001-5180 |
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Notes |
PMID:8470468 |
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no |
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Call Number |
refbase @ user @ |
Serial |
1945 |
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Author |
Colahan, P.; Lindsey, E.; Nunier, C. |
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Title |
Determination of the center of pressure of the hoofs of the forelimbs of horses standing on a flat level surface |
Type |
Journal Article |
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Year |
1993 |
Publication |
Acta Anatomica |
Abbreviated Journal |
Acta Anat (Basel) |
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Volume |
146 |
Issue |
2-3 |
Pages |
175-178 |
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Keywords |
Animals; Forelimb/*physiology; Hoof and Claw/*physiology; Horses/*physiology; *Posture; Pressure |
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The pressure exerted on a flat level surface by recently trimmed, unshod hoofs of the front limbs of 23 sound, adult horses was measured using pressure-sensitive film and a specially built cassette. The horses were tranquilized and stood with one foot on the 2.9-cm-thick cassette and the other on a block of equal height. The hoofs were observed for motion during the measurement, and the developed film was examined for improper alignment of the film or slipping of the hoof. The center of pressure was located using the method of weighted proportions of Barrey. This static measurement system with a long measurement time and the number of measurements reduced the influence of variables inherent in the horses' behavior and the measuring system. The calculated point was recorded as falling medial to, lateral to or on a line bisecting the central sulcus of the frog. In the dorsal to palmar orientation the point was classified with reference to a line drawn halfway between the most dorsal and the most palmar mark on the film. Forty-six percent of the calculated centers of pressure were located in the medial heel area. Binomial analysis for large samples indicates that this was a significant variation from a random distribution. Seventy-six percent of the centers were located in or on the borders of the medial heel. |
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College of Veterinary Medicine, University of Florida, Gainesville 32608-0136 |
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0001-5180 |
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PMID:8470462 |
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no |
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refbase @ user @ |
Serial |
1946 |
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Author |
Clayton, H.M. |
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Title |
The extended canter: a comparison of some kinematic variables in horses trained for dressage and for racing |
Type |
Journal Article |
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Year |
1993 |
Publication |
Acta Anatomica |
Abbreviated Journal |
Acta Anat (Basel) |
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Volume |
146 |
Issue |
2-3 |
Pages |
183-187 |
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Animal Husbandry; Animals; *Gait; Horses/*physiology; *Physical Conditioning, Animal; *Sports |
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Abstract |
This study was designed to test the hypothesis that there is no significant difference in selected temporal and linear stride variables of the extended canter in horses bred and trained for dressage or racing. Nine advanced-level dressage horses and 7 Thoroughbred racehorses were filmed at a frame rate of 200 Hz at an extended canter on a sand track. Two strides were recorded per trial, and each horse performed 6 or 7 trials. Temporal and linear data were determined from the films, and descriptive statistics (mean, SD) were calculated. Strides were selected for analysis on the basis of having a velocity in the range of 6.0-7.0 m/s, and multivariate analysis of variance was used to detect significant differences in the stride kinematics of horses trained for the two sports (p < or = 0.01). The average velocity of the dressage horses was 6.37 m/s, compared with 6.40 m/s for the racehorses. There were no significant differences between the two groups in velocity, stride duration, stride length or the distances between limb placements. The stance durations of all four limbs and the overlaps between them were longer, whereas the duration of the suspension phase was shorter in the dressage horses than in the racehorses (p < or = 0.01). The time between impacts of the diagonal limb pair was close to zero in both groups, with individual horses showing some variability in the order of placement of the diagonal limb pair. However, the sequence of footfalls was not significantly different between the two groups (p < or = 0.01). |
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Department of Veterinary Anatomy, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada |
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0001-5180 |
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PMID:8470464 |
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no |
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Call Number |
Equine Behaviour @ team @ |
Serial |
3751 |
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Author |
Argue, C.K.; Clayton, H.M. |
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Title |
A preliminary study of transitions between the walk and trot in dressage horses |
Type |
Journal Article |
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Year |
1993 |
Publication |
Acta Anatomica |
Abbreviated Journal |
Acta Anat (Basel) |
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Volume |
146 |
Issue |
2-3 |
Pages |
179-182 |
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Keywords |
Animals; Forelimb/physiology; Gait/*physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/physiology; *Physical Conditioning, Animal |
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The object of this study was to determine the limb support sequence during the transitions from walk to trot and from trot to walk in dressage horses under saddle and to test the null hypothesis that the limb support sequence during the transitions is not related to the level of training. Sixteen dressage horses training at novice to FEI Grand Prix level were videotaped performing an average of 9 transitions each from walk to trot and from trot to walk. The 30-Hz videotapes were viewed in slow motion, and based on the limb support sequence the transitions were categorized into two types. In type 1 transitions there were no intermediate steps between the walk and trot sequences. Type 2 transitions were characterized by intermediate steps, including a single support phase. The Kendall rank-order correlation coefficient showed that a higher level of training was positively associated with an increased percentage of type 1 transitions for both walk-to-trot transitions (p < or = 0.05) and trot-to-walk transitions (p < or = 0.01). No significant preference for initiating or completing the trot on the left or right diagonal was found using the binomial test for individual horses and the Wilcoxon signed-ranks test for the group. |
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Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada |
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0001-5180 |
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PMID:8470463 |
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Equine Behaviour @ team @ |
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3752 |
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Author |
Levin, L.E.; Grillet, M.E. |
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[Diversified leadership: a social solution of problems in schools of fish] |
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Journal Article |
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1988 |
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Acta Cientifica Venezolana |
Abbreviated Journal |
Acta Cient Venez |
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39 |
Issue |
2 |
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175-180 |
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Animals; Fishes; *Leadership; Reversal Learning/*physiology; *Social Behavior |
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Spanish |
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Liderazgo diversificado: una solucion social de problemas en el cardumen |
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0001-5504 |
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PMID:3251383 |
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2045 |
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Author |
Palagi, E.; Cordoni, G.; Borgognini Tarli, S. |
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Title |
Possible roles of consolation in captive chimpanzees (Pan troglodytes) |
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Journal Article |
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Year |
2006 |
Publication |
American Journal of Physical Anthropology |
Abbreviated Journal |
Am J Phys Anthropol |
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Volume |
129 |
Issue |
1 |
Pages |
105-111 |
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Keywords |
Animals; Animals, Zoo/*physiology; Empathy; Female; Male; Pan troglodytes/*physiology; *Social Behavior; Stress/physiopathology/veterinary |
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Abstract |
Empathy is a necessary prerequisite for the occurrence of consolation. The term “consolation” contains a hypothesis about function, which is distress alleviation. The present study aims to confirm the occurrence of consolation in captive chimpanzees via the post-conflict/matched-control method (PC-MC) and to suggest its possible roles. We collected 273 PC-MC pairs in the group of Pan troglodytes housed in the ZooParc de Beauval (France). We confirmed the presence of consolatory contacts (mean level of consolation, 49.5% +/- 22.3% SEM) in the colony. Consolation rates were significantly higher than reconciliation levels (mean level of reconciliation, 28.9% +/- 16.8% SEM). The level of consolation was greater in the absence of reconciliation than in the presence of it, suggesting that consolation might be an alternative behavior. As friendship and relatedness did not influence the occurrence of consolation, they did not seem to be the best prerequisites for this behavioral mechanism, at least in this chimpanzee colony. Affinitive contacts with third parties were significantly more frequent when the victim called attention to itself during severe aggressions by screaming. These high-pitched sounds seem to be useful in eliciting aid from conspecifics, as occurs in young humans. The occurrence of consolation reduced the likelihood of further attacks among group-members. From this perspective, both victims and consolers most likely gain potential advantages by interacting with each other when aggression is particularly severe, reconciliation is not immediate, and consequently social stress reaches high levels. |
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Centro Interdipartimentale Museo di Storia Naturale e del Territorio, Universita di Pisa, 56010 Calci, Italy. betta.palagi@museo.unipi.it |
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0002-9483 |
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PMID:16229027 |
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Call Number |
refbase @ user @ |
Serial |
2871 |
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| Permanent link to this record |
