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Author |
Wolff, A.; Hausberger, M. |
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Title |
Learning and memorisation of two different tasks in horses: the effects of age, sex and sire |
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Journal Article |
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Year |
1996 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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46 |
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3-4 |
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137-143 |
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Horse; Learning; Memory; Instrumental task; Spatial task |
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Learning and memory abilities of 1-3 year old horses were assessed using instrumental and spatial tasks. No important differences were observed in the success of learning of the instrumental task (chest opening) according to sex or age. Younger females, however, seemed to learn more quickly. The offspring of a particular stallion were slower to learn than other horses. All horses memorised this task and opened the chest in a very short time in the second session. The animals that learned the task easily were not necessarily faster in the memorisation test. In the spatial task, learning ability did not seem to be related to age but more females than males were successful. The offspring of one stallion were more successful than other horses. Only 76% of the horses succeeded in the memorisation test, independently of age or sex. No correlation was found between the tasks in the latencies of either the learning or the memorisation tests for the same horses. The instrumental and spatial tasks may involve different processes. |
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refbase @ user @ |
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855 |
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Francis-Smith, K.; Wood-Gush, D.G.M. |
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Title |
Copropgagia as seen in thoroughbred foals |
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Journal Article |
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Year |
1977 |
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Equine veterinary journal |
Abbreviated Journal |
Equine Vet J |
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Volume |
9 |
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3 |
Pages |
155-157 |
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Keywords |
animal; article; coprophagy; defecation; eating; female; horse; horse disease; human; mastication |
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Four Thoroughbred foals were seen to quickly eat part of the faeces deposited by their own dams on some 40 per cent of the mare-defaecating occasions observed between the second and fifth week after birth. They did not do it before or after this period. This behaviour was thought to be a feeding pattern which formed a normal part of the foal's development. |
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from Professor Hans Klingels Equine Reference List |
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1090 |
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Author |
Ishida, N.; Hirano, T.; Mukoyama, H. |
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Title |
Detection of aberrant alleles in the D-loop region of equine mitochondrial DNA by single-strand conformation polymorphism (SSCP) analysis |
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Journal Article |
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Year |
1994 |
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Animal Genetics |
Abbreviated Journal |
Anim Genet |
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25 |
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4 |
Pages |
287 |
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Keywords |
*Alleles; Animals; Base Sequence; *DNA, Mitochondrial; DNA, Single-Stranded/genetics; Female; Gene Frequency; Genomic Imprinting; Horses/*genetics; Male; Molecular Sequence Data; Pedigree; *Polymorphism, Genetic |
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Laboratory of Molecular and Cellular Biology, Japan Racing Association, Tokyo |
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0268-9146 |
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PMID:7985852 |
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Equine Behaviour @ team @ |
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2213 |
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McDonnell, S.M.; Henry,M. Bristol, F. |
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Title |
Spontaneous erection and masturbation in equids Proc 35th |
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1991 |
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J. Reprod. Fert. Suppl |
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44 |
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664-665 |
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Horse; Masturbation; Erection; Aversive conditioning; Sexual behavior; Ejaculation; Semen |
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This paper is part of the special issue entitled “Proceedings of the 4th International Symposium on Stallion Reproduction”, Guest Edited by Dr. Edward Squires. |
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1378 |
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Merkies, K.; McKechnie, M.J.; Zakrajsek, E. |
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Title |
Behavioural and physiological responses of therapy horses to mentally traumatized humans |
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Journal Article |
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2018 |
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Applied Animal Behaviour Science |
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Equine-assisted therapy; Ptsd; Horse; Behaviour; Cortisol; Heart rate |
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The benefits to humans of equine-assisted therapy (EAT) have been well-researched, however few studies have analyzed the effects on the horse. Understanding how differing mental states of humans affect the behaviour and response of the horse can assist in providing optimal outcomes for both horse and human. Four humans clinically diagnosed and under care of a psychotherapist for Post-Traumatic Stress Disorder (PTSD) were matched physically to four neurotypical control humans and individually subjected to each of 17 therapy horses loose in a round pen. A professional acting coach instructed the control humans in replicating the physical movements of their paired PTSD individual. Both horses and humans were equipped with a heart rate (HR) monitor recording HR every 5secs. Saliva samples were collected from each horse 30 min before and 30 min after each trial to analyze cortisol concentrations. Each trial consisted of 5 min of baseline observation of the horse alone in the round pen after which the human entered the round pen for 2 min, followed by an additional 5 min of the horse alone. Behavioural observations indicative of stress in the horse (gait, head height, ear orientation, body orientation, distance from the human, latency of approach to the human, vocalizations, and chewing) were retrospectively collected from video recordings of each trial and analyzed using a repeated measures GLIMMIX with Tukey's multiple comparisons for differences between treatments and time periods. Horses moved slower (p < 0.0001), carried their head lower (p < 0.0001), vocalized less (p < 0.0001), and chewed less (p < 0.0001) when any human was present with them in the round pen. Horse HR increased in the presence of the PTSD humans, even after the PTSD human left the pen (p < 0.0001). Since two of the PTSD/control human pairs were experienced with horses and two were not, a post-hoc analysis showed that horses approached quicker (p < 0.016) and stood closer (p < 0.0082) to humans who were experienced with horses. Horse HR was lower when with inexperienced humans (p < 0.0001) whereas inexperienced human HR was higher (p < 0.0001). Horse salivary cortisol did not differ between exposure to PTSD and control humans (p > 0.32). Overall, behavioural and physiological responses of horses to humans are more pronounced based on human experience with horses than whether the human is diagnosed with a mental disorder. This may be a reflection of a directness of movement associated with humans who are experienced with horses that makes the horse more attentive. It appears that horses respond more to physical cues from the human rather than emotional cues. This knowledge is important in tailoring therapy programs and justifying horse responses when interacting with a patient in a therapy setting. |
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0168-1591 |
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Equine Behaviour @ team @ |
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6385 |
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Author |
Dougherty, D.M.; Lewis, P. |
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Title |
Stimulus generalization, discrimination learning, and peak shift in horses |
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Journal Article |
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Year |
1991 |
Publication |
Journal of the experimental analysis of behavior |
Abbreviated Journal |
J Exp Anal Behav |
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56 |
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1 |
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97-104 |
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Animals; *Appetitive Behavior; Attention; *Conditioning, Operant; *Discrimination Learning; Female; *Generalization, Stimulus; Horses/*psychology; Male; *Pattern Recognition, Visual; Size Perception |
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Using horses, we investigated three aspects of the stimulus control of lever-pressing behavior: stimulus generalization, discrimination learning, and peak shift. Nine solid black circles, ranging in size from 0.5 in. to 4.5 in. (1.3 cm to 11.4 cm) served as stimuli. Each horse was shaped, using successive approximations, to press a rat lever with its lip in the presence of a positive stimulus, the 2.5-in. (6.4-cm) circle. Shaping proceeded quickly and was comparable to that of other laboratory organisms. After responding was maintained on a variable-interval 30-s schedule, stimulus generalization gradients were collected from 2 horses prior to discrimination training. During discrimination training, grain followed lever presses in the presence of a positive stimulus (a 2.5-in circle) and never followed lever presses in the presence of a negative stimulus (a 1.5-in. [3.8-cm] circle). Three horses met a criterion of zero responses to the negative stimulus in fewer than 15 sessions. Horses given stimulus generalization testing prior to discrimination training produced symmetrical gradients; horses given discrimination training prior to generalization testing produced asymmetrical gradients. The peak of these gradients shifted away from the negative stimulus. These results are consistent with discrimination, stimulus generalization, and peak-shift phenomena observed in other organisms. |
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Department of Psychology, Ohio University, Athens 45701 |
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0022-5002 |
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PMID:1940765 |
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1764 |
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Rapin, V.; Poncet, P.A.; Burger, D.; Mermod, C.; Richard, M.A. |
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Title |
[Measurement of the attention time in the horse] |
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Journal Article |
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Year |
2007 |
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Schweizer Archiv fur Tierheilkunde |
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Schweiz Arch Tierheilkd |
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149 |
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2 |
Pages |
77-83 |
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Age Factors; Animals; *Attention/physiology; Conditioning, Operant; Cross-Over Studies; Female; Horses/*physiology; Learning/*physiology; Male; Memory/*physiology; Time Factors |
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Abstract |
A study carried out on 49 horses showed that it is possible to measure the attention time by operant conditioning. After teaching horses an instrumental task using a signal, we were then able to test their attention time by asking them to prolong it increasingly while setting success and failure criteria. Two tests were performed 3 weeks apart. The 2nd test was feasible without relearning, a proof of memory, and was repeatable, a proof of consistency in the attention time. A significant difference was observed between the 3 age groups. Young horses often performed very well during the 1st test but their attention dropped in the 2nd test while older horses were more stable with respect to attention and even increased it slightly. The study shows that there are individual differences but it was not possible to prove a significant influence of breed, gender and paternal influence. Consequently, learning appears to be one of the most interesting approaches for evaluating the attention of horses and for observing their behaviour. |
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Laboratoire d'Ecoethologie, Universite de Neuchatel, Suisse. veronique.rapin@unine.ch |
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French |
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Mesure de la duree d'attention chez le cheval |
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0036-7281 |
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PMID:17343134 |
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1770 |
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van Heel, M.C.V.; Kroekenstoel, A.M.; van Dierendonck, M.C.; van Weeren, P.R.; Back, W. |
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Title |
Uneven feet in a foal may develop as a consequence of lateral grazing behaviour induced by conformational traits |
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Journal Article |
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2006 |
Publication |
Equine veterinary journal |
Abbreviated Journal |
Equine. Vet. J. |
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38 |
Issue |
7 |
Pages |
646-651 |
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Aging/*physiology; Animals; Animals, Newborn/anatomy & histology/growth & development/physiology; Feeding Behavior/*physiology; Female; Forelimb/*anatomy & histology/*physiology; *Horses/anatomy & histology/growth & development/physiology; Male |
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REASONS FOR PERFORMING STUDY: Conformational traits are important in breeding, since they may be indicative for performance ability and susceptibility to injuries. OBJECTIVES: To study whether certain desired conformational traits of foals are related to lateralised behaviour while foraging and to the development of uneven feet. METHODS: Twenty-four Warmblood foals, born and raised at the same location, were studied for a year. Foraging behaviour was observed by means of weekly 10 min scan-sampling for 8 h. A preference test (PT) was developed to serve as a standardised tool to determine laterality. The foals were evaluated at age 3, 15, 27 and 55 weeks. The PT and distal limb conformation were used to study the relation between overall body conformation, laterality and the development of uneven feet. Pressure measurements were used to determine the loading patterns under the feet. RESULTS: About 50% of the foals developed a significant preference to protract the same limb systematically while grazing, which resulted in uneven feet and subsequently uneven loading patterns. Foals with relatively long limbs and small heads were predisposed to develop laterality and, consequently unevenness. CONCLUSIONS: Conformational traits may stimulate the development of laterality and therefore indirectly cause uneven feet. |
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Derona Equine Performance Laboratory, Department of Equine Sciences, Faculty of Veterinary Medicine, Utrecht University, Yalelaan 12, NL-3584 CM Utrecht, The Netherlands |
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0425-1644 |
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PMID:17228580 |
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1774 |
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Author |
Giles, N.; Tupper, J. |
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Title |
Equine interspecies aggression |
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2006 |
Publication |
The Veterinary record |
Abbreviated Journal |
Vet. Rec. |
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159 |
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22 |
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756 |
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Aggression/*physiology; Animals; Horses/*physiology; Sheep/*physiology; Social Dominance |
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0042-4900 |
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PMID:17127768 |
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1779 |
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Author |
Arnold, W.; Ruf, T.; Kuntz, R. |
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Title |
Seasonal adjustment of energy budget in a large wild mammal, the Przewalski horse (Equus ferus przewalskii) II. Energy expenditure |
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Journal Article |
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Year |
2006 |
Publication |
The Journal of experimental biology |
Abbreviated Journal |
J Exp Biol |
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209 |
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Pt 22 |
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4566-4573 |
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Animals; Animals, Wild/*physiology; Body Temperature; Body Temperature Regulation; Eating; *Energy Metabolism; Female; Heart Rate; Horses/*physiology; Male; Motor Activity; Pregnancy; Reproduction; *Seasons |
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Many large mammals show pronounced seasonal fluctuations of metabolic rate (MR). It has been argued, based on studies in ruminants, that this variation merely results from different levels of locomotor activity (LA), and heat increment of feeding (HI). However, a recent study in red deer (Cervus elaphus) identified a previously unknown mechanism in ungulates--nocturnal hypometabolism--that contributed significantly to reduced energy expenditure, mainly during late winter. The relative contribution of these different mechanisms to seasonal adjustments of MR is still unknown, however. Therefore, in the study presented here we quantified for the first time the independent contribution of thermoregulation, LA and HI to heart rate (f(H)) as a measure of MR in a free-roaming large ungulate, the Przewalski horse or Takhi (Equus ferus przewalskii Poljakow). f(H) varied periodically throughout the year with a twofold increase from a mean of 44 beats min(-1) during December and January to a spring peak of 89 beats min(-1) at the beginning of May. LA increased from 23% per day during December and January to a mean level of 53% per day during May, and declined again thereafter. Daily mean subcutaneous body temperature (T(s)) declined continuously during winter and reached a nadir at the beginning of April (annual range was 5.8 degrees C), well after the annual low of air temperature and LA. Lower T(s) during winter contributed considerably to the reduction in f(H). In addition to thermoregulation, f(H) was affected by reproduction, LA, HI and unexplained seasonal variation, presumably reflecting to some degree changes in organ mass. The observed phase relations of seasonal changes indicate that energy expenditure was not a consequence of energy uptake but is under endogenous control, preparing the organism well in advance of seasonal energetic demands. |
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Research Institute of Wildlife Ecology, University of Veterinary Medicine, Vienna, Savoyenstrasse 1, 1160 Vienna, Austria. walter.arnold@vu-wien.ac.at |
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0022-0949 |
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PMID:17079726 |
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1782 |
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