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	Author	Baragli,P.; Cozzi,A; Rugani, R.; Sighieria, C.; Regolin, L.
	Title	*Delayed search for non-social goals by Equids (Equus caballus* and Equus asinus)**			Type	Conference Article
	Year	2008	Publication	IESM 2008	Abbreviated Journal
	Volume		Issue		Pages
	Keywords	cognitive abilities, delayed-response, Equids, working memory.
	Abstract	Delayed-responses have been traditionally employed to investigate the temporal characteristics of animals“ ability to represent and recall objects that have disappeared. In the typical condition, the animal, usually a mammal, observes the experimenter hiding an interesting goal (e.g. some food) in a certain location. A delayed-response task (DRT) was administered to 4 female Esperia pony (2 years old) coming from a free-range breed (Frosinone, Italy) and to 7 female Amiata donkeys (4.2±2 years old) coming from a conservation stock (University of Pisa, Italy). The DRT's apparatus was located in a square fence. A single ”U-shaped“ screen (330x160x140 cm) made by wood shavings blocks was positioned in the centre of the fence. A gap (40x50 cm) on the ground was in the middle of the central side of the U-shaped-screen and served to make the food-attractor disappear. The food-attractor consisted in cereal flakes and fresh grass for ponies and cereal flakes for donkeys. A bucket full of food was placed on a dolly tied on a rope which could be pulled by an experimenter. In a preliminary training each animal was allowed to eat food from the bucket and, while the animal was eating, the dolly was gently pulled away from the animal, and beyond the screen through the gap. The subjects needed to move around of the screen in order to retrieve the food. As a reinforcement, they were allowed to eat some food from the bucket once behind the screen. From trial to trial, the bucket was presented farther and farther (starting with a distance of 1 m in front of the screen to reach 7 m). Therefore subjects were tested in the DRT requiring them to rejoin the bucket with the goal-food disappearing behind the screen as in the preliminary training but following a 10 s delay. For the DRT, the bucket was placed 7 m in front of the screen, 3 m away from the animal's starting area. Then the dolly was pulled away from the animal. Ten seconds after the disappearance of the dolly behind the screen the animal was released from the starting area. The DRT ended when the subject had reached the attractor behind the screen on 3 consecutive trials. Results showed that all animals were able to rejoin the food behind the screen after 10 s delay. The mean time of the delayed-response (mean±sd, in s) in the ponies (1st: 19.8±8; 2nd: 10.8±2.2; 3rd: 12.8±2.8) and in the donkeys (1st: 28.4±10; 2nd: 26.9±13; 3rd: 24.3±16.6) showed a trend to decrease from first trial to third. These preliminary results suggest that like other mammals our ponies and donkeys can maintain a working memory trace of the location where biologically attractive objects have been seen to disappear. In conclusion, this study paves the way to set up a viable model system for the investigation of the more sophisticated aspects of Equids” cognitive abilities such as working memory.
	Address	Department of Veterinary Anatomy, Biochemistry and Physiology, University of Pisa, Viale delle Piagge 2 – 56124, Pisa, Italy
	Corporate Author	Cozzi, A			Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference	IESM 2008
	Notes	Talk 15 min IESM 2008			Approved	yes
	Call Number	Equine Behaviour @ team @			Serial	4475
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	Author	Moehlman, P.D.
	Title	Endangered wild equids			Type	Journal Article
	Year	2005	Publication	Scientific American	Abbreviated Journal	Sci Am
	Volume	292	Issue	3	Pages	74-81
	Keywords	Africa; Americas; Animals; Animals, Wild/physiology; Asia; Conservation of Energy Resources; Environment; *Equidae/physiology; Food Chain; Humans; Male; Reproduction
	Abstract
	Address	IUCN-The World Conservation Union/Species Survival Commission Equid Specialist Group
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0036-8733	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15859216			Approved	no
	Call Number	refbase @ user @			Serial	551
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	Author	McDonnell, S.M.; Haviland, J.C.S.
	Title	Agonistic ethogram of the equid bachelor band			Type	Journal Article
	Year	1995	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	43	Issue	3	Pages	147-188
	Keywords	Equid; Horse; Bachelor; Agonistic; Social behavior; Ethogram
	Abstract	An ethogram of agonistic and related behaviors among equid bachelor band members was developed. Several key English-language studies on equids were reviewed to derive a preliminary inventory of specific behaviors to be included in the ethogram. A bachelor band of domestic pony stallions pastured together was observed for approximately 50 daylight hours to obtain detailed descriptions of each behavior, enable photographic and video documentation of behaviors, and identify any behaviors to be added to the preliminary inventory. An initial draft of the ethogram was sent to 65 equine researchers for review. Twenty-eight critical reviews were received and their suggestions considered for the final draft. A total of 49 elemental behaviors including five distinct vocalizations was included in the ethogram. Three complex behavioral sequences were also included. Most of the behaviors catalogued from the direct observation of pastured pony stallions were also found in the equid literature. For many, references to these behaviors specifically among males or bachelor band members were not found. The results offer a practical tool for quantitative research and other studies of equid inter-male behavior as well as for teaching of equid behavior, and should facilitate progress toward development of a complete ethogram for the horse and other equids.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @ ; Equine Behaviour @ team @ room B 3.029			Serial	749
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	Author	Haring, H.
	Title	Development, level and prospects of the german horse breeding			Type	Journal Article
	Year	2005	Publication	Zuechtungskunde	Abbreviated Journal	Zuechtungskunde
	Volume	77	Issue	6	Pages	490-495
	Keywords	Breeding program; Breeding/Member association; Change of structure; Estimation of breeding values; Fédération Equestre Nationale (FN)/German Equestrian Federation; Step of selection; Equidae; Equus caballus
	Abstract	The economic impact of the horses of the Federal Republic of Germany has gone up, the statistic numerals verify obviously that Germany took pride of place in Europe in terms of numbers of riders as well as numbers of horses. Successes of German branded horses let their breeders reach the summit worldwide. The carefully agreed breeding programme connects practical cognitions with those of science and permits the leading breeding areas unobstructed space to set their own priorities. Globalisation and rised demand of customers forces breeding associations towards a far-reaching reorganisation because just large powerful institutions can meet these requirements. An end of this process, which scarcely has just begun, cannot yet be conceivable seen. – Eugen Ulmer KG, Stuttgart.
	Address	Deutsche Reiterliche Vereinigung e. V., 48229 Warendorf, Germany
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	00445401 (Issn)	ISBN		Medium
	Area		Expedition		Conference
	Notes	Cited By (since 1996): 1; Export Date: 21 April 2007; Source: Scopus; Language of Original Document: German; Correspondence Address: Haring, H.; Deutsche Reiterliche Vereinigung e. V. 48229 Warendorf, Germany; email: Hharing@fn-dokr.de			Approved	no
	Call Number	refbase @ user @			Serial	791
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	Author	Ryder, O.A.
	Title	Przewalski's horse – putting the wild horse back in the wild			Type	Journal Article
	Year	1988	Publication	Oryx	Abbreviated Journal	Oryx
	Volume	22	Issue		Pages	154-157
	Keywords	Equidae genetics.
	Abstract	The Asian wild horse, or Przewalski's horse (Equus przewalskii), is believed to currently survive only in captivity. There are more than 660 individuals in over 70 zoological collections, and animals are available for reintroduction. The Przewalski's horse had been bred in captivity for 12 generations, and inbreeding has occurred. Genetic variability has been lost, and released animals will require acclimatization on or near release sites. In China, a program is under way for acclimatization, breeding, and release of wild horses, and plans are being made for a similar program in Mongolia. (LCA)
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
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	Area		Expedition		Conference
	Notes	from Professor Hans Klingels Equine Reference List			Approved	yes
	Call Number				Serial	1537
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	Author	Thackeray, J.F.
	Title	Zebras from wonderwerk cave, northern Cape province, South Africa: attempts to distinguish Equus burchelli and E. quagga			Type	Journal Article
	Year	1988	Publication	South African journal of science	Abbreviated Journal	Suid- Afrikaanse Tydsskrif vir Wetenskap
	Volume	84	Issue		Pages	99-101
	Keywords	Cape Province; Teeth; Statistical analysis; Equidae; Hippomorpha; South Africa; Southern Africa; Perissodactyla; Mammalia; Vertebrata
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0038-2353	ISBN		Medium
	Area		Expedition		Conference
	Notes	from Professor Hans Klingels Equine Reference List			Approved	yes
	Call Number				Serial	1644
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	Author	Burden, F.; Trawford, A.
	Title	Equine interspecies aggression Comment on			Type
	Year	2006	Publication	The Veterinary record	Abbreviated Journal	Vet. Rec.
	Volume	159	Issue	25	Pages	859-860
	Keywords	Aggression; Animals; Behavior, Animal; Cats; Dogs; Equidae
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0042-4900	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17172484			Approved	no
	Call Number				Serial	1777
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	Author	Fischhoff, I.R.; Sundaresan, S.R.; Cordingley, J.; Larkin, H.M.; Sellier, M.-J.; Rubenstein, D.I.
	Title	Social relationships and reproductive state influence leadership roles in movements of plains zebra, Equus burchellii			Type	Journal Article
	Year	2007	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	73	Issue	5	Pages	825-831
	Keywords	collective movements; drinking; equid; Equus burchellii; foraging; group dynamics; motivation; plains zebra; reproductive status; social relationships
	Abstract	In animal groups, collective movements emerge from individual interactions. Biologists seek to identify how characteristics of actors in these groups, and their relationships, influence the decision-making process. We distinguished two basic factors determining leadership in group choices: identity and state. We hypothesized that identity is more important to leadership in groups with stable relationships, which permit the development of habitual roles. In groups with fluid membership, particular individuals or subgroups are less likely to emerge as consistent leaders. Instead, we predicted that movement initiation in unstable groups depends on individual state at the time of the decision. We characterized how identity and reproductive state influenced leadership patterns in the movements of plains zebra. As in many other mammals, lactation in this species significantly alters water and energy needs. We investigated leadership in tightly knit harems and loosely bonded herds of multiple harems. Harem females tended to have habitual roles in the initiation of harem movement. In herds, however, we found no consistent leaders among harems. At both levels of social organization, lactation was a key determinant of leadership. In harems, lactating females were more likely to initiate movement than nonlactating females. In turn, harems containing lactating females were more likely to lead herd movements. Thus, we conclude that social relationships and reproductive state together shape the interactions that produce group behaviours. One benefit to lactating females of leading herd movements is preferential access to scarce water. Thus, leadership roles in group decisions may have fitness consequences.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ room B 3.092			Serial	1825
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	Author	Cameron, E.Z.; Linklater, W.L.; Stafford, K.J.; Minot, E.O.
	Title	Aging and improving reproductive success in horses: declining residual reproductive value or just older and wiser?			Type	Journal Article
	Year	2000	Publication	Behavioral Ecology and Sociobiology	Abbreviated Journal	Behav. Ecol. Sociobiol.
	Volume	47	Issue	4	Pages	243-249
	Keywords	Maternal investment – Equidae – Equus caballus
	Abstract	In many mammalian species, female success in raising offspring improves as they age. The residual reproductive value hypothesis predicts that each individual offspring will be more valuable to the mother as she ages because there is less conflict between the current and potential future offspring. Therefore, as mothers age, their investment into individual offspring should increase. Empirical evidence for an influence of declining residual reproductive value on maternal investment is unconvincing. Older mothers may not invest more, but may be more successful due to greater experience, allowing them to target their investment more appropriately (targeted reproductive effort hypothesis). Most studies do not preclude either hypothesis. Mare age significantly influenced maternal investment in feral horses living on the North Island of New Zealand. Older mares, that were more successful at raising foals, were more protective for the first 20 days of life, but less diligent thereafter. Total maternal input by older mothers did not seem to be any greater, but was better targeted at the most critical period for foal survival and a similar pattern was observed in mares that had lost a foal in the previous year. In addition, older mothers were more likely to foal in consecutive years, supporting the hypothesis that they are investing less than younger mares in individual offspring. Therefore, older mothers seem to become more successful by targeting their investment better due to experience, not by investing more in their offspring.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2019
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	Author	Linklater, W.L.
	Title	Adaptive explanation in socio-ecology: lessons from the Equidae			Type	Journal Article
	Year	2000	Publication	Biological Reviews of the Cambridge Philosophical Society	Abbreviated Journal	Biol. Rev.
	Volume	75	Issue	1	Pages	1-20
	Keywords	Adaptation, Physiological; Animals; Ecology; Equidae/physiology; Female; Male; Phylogeny
	Abstract	Socio-ecological explanations for intra- and interspecific variation in the social and spatial organization of animals predominate in the scientific literature. The socio-ecological model, developed first for the Bovidae and Cervidae, is commonly applied more widely to other groups including the Equidae. Intraspecific comparisons are particularly valuable because they allow the role of environment and demography on social and spatial organization to be understood while controlling for phylogeny or morphology which confound interspecific comparisons. Feral horse (Equus caballus Linnaeus 1758) populations with different demography inhabit a range of environments throughout the world. I use 56 reports to obtain 23 measures or characteristics of the behaviour and the social and spatial organization of 19 feral horse populations in which the environment, demography, management, research effort and sample size are also described. Comparison shows that different populations had remarkably similar social and spatial organization and that group sizes and composition, and home range sizes varied as much within as between populations. I assess the few exceptions to uniformity and conclude that they are due to the attributes of the studies themselves, particularly to poor definition of terms and inadequate empiricism, rather than to the environment or demography per se. Interspecific comparisons show that equid species adhere to their different social and spatial organizations despite similarities in their environments and even when species are sympatric. Furthermore, equid male territoriality has been ill-defined in previous studies, observations presented as evidence of territoriality are also found in non-territorial equids, and populations of supposedly territorial species demonstrate female defence polygyny. Thus, territoriality may not be a useful categorization in the Equidae. Moreover, although equid socio-ecologists have relied on the socio-ecological model derived from the extremely diverse Bovidae and Cervidae for explanations of variation in equine society, the homomorphic, but large and polygynous, and monogeneric Equidae do not support previous socio-ecological explanations for relationships between body size, mating system and sexual dimorphism in ungulates. Consequently, in spite of the efforts of numerous authors during the past two decades, functional explanations of apparent differences in feral horse and equid social and spatial organization and behaviour based on assumptions of their current utility in the environmental or demographic context remain unconvincing. Nevertheless, differences in social cohesion between species that are insensitive to intra- and interspecific variation in habitat and predation pressure warrant explanation. Thus, I propose alternative avenues of inquiry including testing for species-specific differences in inter-individual aggression and investigating the role of phylogenetic constraints in equine society. The Equidae are evidence of the relative importance of phylogeny and biological structure, and unimportance of the present-day environment, in animal behaviour and social and spatial organization.
	Address	Institute of Natural Resources, Massey University, Palmerston North, New Zealand
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1464-7931	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10740891			Approved	no
	Call Number				Serial	2024
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