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	Author	Goodwin, D.
	Title	The importance of ethology in understanding the behaviour of the horse			Type	Journal Article
	Year	1999	Publication	Equine Veterinary Journal	Abbreviated Journal	Equine Veterinary Journal
	Volume	31	Issue	S28	Pages	15-19
	Keywords	horse; behaviour; domestication; interspecific communication
	Abstract	Summary Domestication has provided the horse with food, shelter, veterinary care and protection, allowing individuals an increased chance of survival. However, the restriction of movement, limited breeding opportunities and a requirement to expend energy, for the benefit of another species, conflict with the evolutionary processes which shaped the behaviour of its predecessors. The behaviour of the horse is defined by its niche as a social prey species but many of the traits which ensured the survival of its ancestors are difficult to accommodate in the domestic environment. There has been a long association between horses and man and many features of equine behaviour suggest a predisposition to interspecific cooperation. However, the importance of dominance in human understanding of social systems has tended to overemphasise its importance in the human-horse relationship. The evolving horse-human relationship from predation to companionship, has resulted in serial conflicts of interest for equine and human participants. Only by understanding the nature and origin of these conflicts can ethologists encourage equine management practices which minimise deleterious effects on the behaviour of the horse.
	Address
	Corporate Author				Thesis
	Publisher	American Medical Association (AMA)	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0425-1644	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6714
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	Author	McGreevy, P.; Yeates, J.
	Title	Horses (Equus caballus)			Type	Book Chapter
	Year	2018	Publication	Companion Animal Care and Welfare	Abbreviated Journal
	Volume		Issue		Pages
	Keywords	animal company; behavioural signs; diseases; domestic horses; euthanasia; human interaction; nutritional requirements
	Abstract	Summary Domestic horses are equid members of the class Mammalia, order Perissodactyla, and family Equidae. Horses are obligate herbivores, with nutritional requirements as listed in a table. Adequate space is necessary for exercise, exploration, flight, sharing resources, play, and rolling. Company is essential for all horses, including stallions. Company provides opportunities for mutual grooming and play and allows horses to stand head-to-tail to remove flies. Unhandled horses may respond to humans as they would to predators, whereas handled horses' responses depend on their previous interactions with humans. Horses can suffer from several diseases as listed in another table. The best method of euthanasia of horses is usually sedation followed by either cranial shooting or the injection of an overdose of pentobarbitone into the jugular vein. Behavioural signs of distress can include increased locomotory activity, vigilance behaviours, neighing, snorting, pawing, nibbling walls and buckets, defaecation, rearing, kicking stable walls or doors, and high-stepping 'prancing'.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title	Wiley Online Books	Abbreviated Series Title	Companion Animal Care and Welfare
	Series Volume		Series Issue		Edition
	ISSN		ISBN	9781119333708	Medium
	Area		Expedition		Conference
	Notes	doi:10.1002/9781119333708.ch13			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6506
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	Author	Skandakumar, S.; Stodulski, G.; Hau, J.
	Title	Salivary IgA: a Possible Stress Marker In Dogs			Type	Abstract
	Year	1995	Publication	Animal Welfare	Abbreviated Journal
	Volume	4	Issue	4	Pages	339-350
	Keywords	Animal Welfare; Behaviour; Cortisol; Dog; Salivary Iga (S-Iga); Stress; Well-Being
	Abstract	Stress in humans has been reported to be associated with a decrease in the salivary immunoglobulin A (s-IgA) levels enabling the possible use of s-IgA to assess stress. Prolonged stress, if reliably assessed in a non-invasive manner, may be used to assess animal welfare. This study analysed groups of dogs undergoing physical and temperamental training and s-IgA levels were measured by rocket immunoelectrophoresis in prospective samples. Behavioural assessment was carried out and cortisol levels in saliva were measured by ELISA. A significant negative correlation (P < 0.007) between the logarithmic cortisol concentrations and s-IgA levels in saliva was recorded. The behavioural assessment of the dogs agreed well with the biochemical markers. It is concluded that IgA levels in saliva may be a useful marker of dog well-being and that stress results in decreased s-IgA levels.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5964
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	Author	Górecka-Bruzda, A.; Chruszczewski, M.H.; Jezierski, T.; Murphy, J.
	Title	Behaviour-related traits of Polish sport horse stallions participating in 100-day performance tests			Type	Journal Article
	Year		Publication	Livestock Science	Abbreviated Journal	Livest Sci
	Volume	In Press, Corrected Proof	Issue		Pages
	Keywords	Horse; 100-day performance tests; Behaviour; Temperament; Factor Analysis
	Abstract	Sport horse studbooks frequently use performance tests to identify, evaluate and approve stallions for future breeding programmes. The aim of this study was to analyse behaviour-related traits of Polish Warmblood stallions participating in studbook 100-day performance tests. Both relatedness and differences of traits and their contribution to overall variability were analysed. Three-year-old stallions (n = 374) participated in 100-day performance test programmes at two Polish training stations from 2004 to 2008. Traits including character, temperament and trainability (trainer rated), free-jumping, jumping under the rider, walk, trot and canter (independently rated by both trainer and the selection jury) and jumping ability, dressage-ability and rideability (rated by the independent test rider) were subjected to Factor Analysis (FA) with varimax rotation. The FA produced three factors [Cooperation (Cp), Jumping Potential (JP) and Dressage Potential (DP)] with eigenvalues > 1, which accounted for 64.6% of the total variance of the traits examined. After rotation, the factors represented Cp: 23.2.0%, JP: 21.5% and DP: 19.8% of variance respectively. The traits of character, temperament, trainability, jumping under the rider, rideability, dressage and jumping abilities had high loadings on the second factor (Cp) and thereafter, the assessment of all behaviour-related traits contributed to the general evaluation with approximately one-fifth of total variability. The findings may indicate that successful sport careers for any horse may be much more closely associated with willingness to cooperate with the rider and riding and handling behaviour, than to the horse's actual physical performance potential. It would appear that only so-called sport-suitable performance of the horse is being selected for in performance tests (during jumping and dressage suitability testing). This approach may be short-sighted and inadequate with serious implications for horse under-performance in leisure activities notwithstanding general horse welfare within the disciplines of equitation.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1871-1413	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5281
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	Author	Whistance, L.K.; Sinclair, L.A.; Arney, D.R.; Phillips, C.J.C.
	Title	Trainability of eliminative behaviour in dairy heifers using a secondary reinforcer			Type	Journal Article
	Year	2009	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	117	Issue	3-4	Pages	128-136
	Keywords	Cattle; Eliminative behaviour; Learning; Clicker training; Clean bedding
	Abstract	Soiled bedding influences cleanliness and disease levels in dairy cows and there is no evidence of an inherent latrine behaviour in cattle. If cows were trained to use a concrete area of the housing system as a latrine, a cleaner bed could be maintained. Thirteen group-housed, 14-16-month-old Holstein-Friesian heifers, were clicker trained with heifer-rearing concentrate pellets as a reward. Training was carried out in four phases. (Phase 1) Association of feed reward with clicker, criterion: 34/40 correct responses. (Phase 2) Simple task (nose-butting a disc) to reinforce phase 1 association, criterion: 17/20 correct responses. (Phase 3) Association of eliminative behaviour with reward where criterion was four sessions with only one incorrect response: criteria for each heifer in phases 1-3 were set using binomial tests. (Phase 4) Shaping eliminative behaviour to occur on concrete. Possible responses were, eliminating on concrete (C) or straw (S), or moving from one substrate to another immediately before eliminating: CÂ -->Â S, SÂ -->Â C. Heifers were rewarded for the desired behaviours C and SÂ -->Â C and ignored when S and CÂ -->Â S occurred. If learning was achieved, C should increase as CÂ -->Â S decreased and SÂ -->Â C should increase as S decreased: tested with Spearman rank correlations. All heifers achieved criterion by day 4 of phase 1 (PÂ =Â 0.001); day 1 of phase 2 (PÂ =Â 0.001) and day 10 of phase 3 (PÂ <Â 0.009). Responses changed throughout phase 3 beginning with (i) looking at the trainer whilst voiding then moving to trainer after the click, and later including (ii) moving to trainer immediately before- or (iii) during voiding. No relationship was found between S and SÂ -->Â C (rsÂ =Â -0.14; PÂ =Â 0.63) or C and CÂ -->Â S (rsÂ =Â -0.33; PÂ =Â 0.25). All group members eliminated more often on concrete (580) than on straw (141) but four heifers with consistently longer lying bouts also showed more CÂ -->Â S before lying down (Mann-Whitney, PÂ =Â 0.007). The present study is believed to be the first reported work to show that cattle can be trained to show an awareness of their own eliminative behaviour. This was not successfully shaped to latrine behaviour, however, and it is suggested that floor type may not have been a sufficiently salient cue. Voiding on straw occurred largely with response CÂ -->Â S (0.73) and general behaviour suggested that this was strongly linked to lying patterns of individual heifers.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4765
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	Author	Keil, N.M., Sambraus, H.H.
	Title	“Intervenors” in agonistic interactions amongst domesticated goats			Type	Journal Article
	Year	1998	Publication	Zeitschrift fur Säugetierkunde	Abbreviated Journal	Z. Säugetierk.
	Volume	63	Issue	5	Pages	266-272
	Keywords	Behaviour; Domestication; Goat; Intervention; Rank order
	Abstract	Social behaviour was observed in individually marked goats in two herds. The goats from one herd (n = 98) were horned, those of the other herd (n = 83) were polled. By recording agonistic interactions within the herds, a dominance index was determined for each animal. In both herds, intervention took place. Intervention is defined as one animal pushing in between two fighters, and thus ending the fight. More cases of intervention took place per individual animal amongst the horned goats than amongst the polled ones. Goats which intervened in fights on several occasions usually had a high dominance index. Members of the herd which were observed intervening only once had an average dominance index in both herds of almost 0.5. In some cases, goats very low in the rank order intervened a fight. Only rarely did the intervenors have a lower dominance index than the two fighters. In 103 cases, the direct dominance relationship between a fighting animal and the intervenor was known. In 95 cases (92.2%), the intervenor was dominant to the herd member in this fight and in just eight cases (7.8%), it was subordinate. It could not be determined what advantage the intervenor gained from its activity. It is possible that, at least in certain cases, a particularly relationship existed between the intervenor and one of the fighters.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5236
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	Author	Wey, T.; Blumstein, D.T.; Shen, W.; Jordán, F.
	Title	Social network analysis of animal behaviour: a promising tool for the study of sociality			Type	Journal Article
	Year	2008	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	75	Issue	2	Pages	333-344
	Keywords	animal behaviour; quantifying sociality; social network analysis; social structure
	Abstract	Social animals live and interact together, forming complex relationships and social structure. These relationships can have important fitness consequences, but most studies do not explicitly measure those relationships. An approach that explicitly measures relationships will further our understanding of social complexity and the consequences of both direct and indirect interactions. Social network analysis is the study of social groups as networks of nodes connected by social ties. This approach examines individuals and groups in the context of relationships between group members. Application of social network analysis to animal behaviour can advance the field by identifying and quantifying specific attributes of social relationships, many of which are not captured by more common measures of sociality, such as group size. Sophisticated methods for network construction and analysis exist in other fields, but until recently, have seen relatively little application to animal systems. We present a brief history of social network analysis, a description of basic concepts and previous applications to animal behaviour. We then highlight relevance and constraints of some network measures, including results from an original study of the effect of sampling on network parameter estimates, and we end with promising directions for research. By doing so, we provide a prospective overview of social network analysis' general utility for the study of animal social behaviour.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4691
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	Author	Weissing, F.J.
	Title	Animal behaviour: Born leaders			Type	Journal Article
	Year	2011	Publication		Abbreviated Journal	Nature
	Volume	474	Issue	7351	Pages	288-289
	Keywords	* Animal behaviour * Evolution * Psychology
	Abstract	Social animals face a dilemma. To reap the benefits of group living, they have to stay together. However, individuals differ in their preferences as to where to go and what to do next. If all individuals follow their own preferences, group coherence is undermined, resulting in an outcome that is unfavourable for everyone. Neglecting one's own preferences and following a leader is one way to resolve this coordination problem. But what attributes make an individual a 'leader'? A modelling study by Johnstone and Manica1 illuminates this question.
	Address
	Corporate Author				Thesis
	Publisher	Nature Publishing Group, a division of Macmillan Publishers Limited. All Rights Reserved.	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0028-0836	ISBN		Medium
	Area		Expedition		Conference
	Notes	10.1038/474288a			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5396
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	Author	Murphy, J.; Arkins, S.
	Title	Equine learning behaviour			Type	Journal Article
	Year	2007	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	76	Issue	1	Pages	1-13
	Keywords	Horse; Behaviour; Learning; Processes; Memory
	Abstract	Scientists and equestrians continually seek to achieve a clearer understanding of equine learning behaviour and its implications for training. Behavioural and learning processes in the horse are likely to influence not only equine athletic success but also the usefulness of the horse as a domesticated species. However given the status and commercial importance of the animal, equine learning behaviour has received only limited investigation. Indeed most experimental studies on equine cognitive function to date have addressed behaviour, learning and conceptualisation processes at a moderately basic cognitive level compared to studies in other species. It is however, likely that the horses with the greatest ability to learn and form/understand concepts are those, which are better equipped to succeed in terms of the human-horse relationship and the contemporary training environment. Within equitation generally, interpretation of the behavioural processes and training of the desired responses in the horse are normally attempted using negative reinforcement strategies. On the other hand, experimental designs to actually induce and/or measure equine learning rely almost exclusively on primary positive reinforcement regimes. Employing two such different approaches may complicate interpretation and lead to difficulties in identifying problematic or undesirable behaviours in the horse. The visual system provides the horse with direct access to immediate environmental stimuli that affect behaviour but vision in the horse is of yet not fully investigated or understood. Further investigations of the equine visual system will benefit our understanding of equine perception, cognitive function and the subsequent link with learning and training. More detailed comparative investigations of feral or free-ranging and domestic horses may provide useful evidence of attention, stress and motivational issues affecting behavioural and learning processes in the horse. The challenge for scientists is, as always, to design and commission experiments that will investigate and provide insight into these processes in a manner that withstands scientific scrutiny.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	629
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	Author	Christensen, J.W.; Keeling, L.J.; Nielsen, B.L.
	Title	Responses of horses to novel visual, olfactory and auditory stimuli			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	93	Issue	1-2	Pages	53-65
	Keywords	Horses; Novelty responses; Fear; Behaviour; Heart rate
	Abstract	Responses of horses towards fear-eliciting stimuli can have important consequences for both human and horse safety. This experiment was designed to investigate behavioural and heart rate (HR) responses of horses to novel visual, auditory and olfactory stimuli. Twenty-four 2-year-old, previously unhandled, stallions were habituated to receive a food reward from a container in a test arena. Each horse was exposed to three 2 min tests in a balanced design where in addition to the feed container, either a traffic cone (visual test), white noise (auditory test) or eucalyptus oil applied to the inside of the container (olfactory test) were used as the novel stimuli. Compared to the control, less time was spent eating during all tests. There was no difference in locomotion activity in the different test situations, but presentation of the novel visual and auditory stimuli elicited significantly increased HR responses in the horses, compared to their response to the arena without novel stimuli (control), whereas there was no increase in HR response to the olfactory stimulus. However, during the olfactory test, the horses had an increased number of eating bouts and became more vigilant towards their surroundings, whereas during the visual and auditory tests, more time was spent alert towards the stimulus. The horses also took significantly more steps backwards in response to the auditory test. The heart rate responses correlated between tests and reflect a non-differentiated activation of the sympathetic nervous system, while the behavioural responses were linked to the type of stimulus.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2256
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