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	Author	Mills, D.S.; Riezebos, M.
	Title	The role of the image of a conspecific in the regulation of stereotypic head movements in the horse			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	91	Issue	1-2	Pages	155-165
	Keywords	Behaviour; Horse; Nodding; Perception; Stereotypy; Vision; Weaving
	Abstract	Weaving is a common locomotor stereotypy in horses, which has been shown to be reduced by the presence of a mirror in the stable; however, it remains unclear whether this effect is due to the reflected image of a horse or some other property associated with the introduction of a mirror. To investigate this further, the response of six known weaving horses to each of three different types of poster was recorded. The poster images consisted of a life-size image of a horse's face (true image), a version of the same image cut into 54 squares, which were then randomly rearranged (pixilated image) and a blank (white image) display of the same size as the other posters. Four observation periods were included in each of the two days of observation per treatment condition: 08:00-08:30 h, 10:00-10:30 h, 12:00-12:30 h and 16:00-17:00 h. The first observation period was followed by the provision of concentrate feed and forage, the second observation period by exercise, the third by further forage and the fourth by concentrate feed and forage. The horses' activities and positions in the box were scanned at 1 min intervals and the number of observations registering each activity and position was calculated as a percentage of the total number of observations for each observation period. Weaving was significantly less when the horses were provided with the image of a horse's face (mean percentage of observations +/- S.E.M.; 5.56 +/- 1.57), compared to both the pixilated (14.85 +/- 3.06) and white (20.52 +/- 4.12) images. Nodding was significantly less when provided with the true image of a horse (0.45 +/- 0.15) compared to the pixilated image (2.15 +/- 0.67). The pattern of weaving throughout the day differed from that of nodding, and together these results suggest that the two behaviours are not analogous. The presence of the image of the horse's face was associated with a significant increase in alertness (F112,2 = 11.31, p < 0.001) and recorded time spent looking at the poster(F112,2 = 3.46, p < 0.05). The timing of stereotypic head movements in the horse in this study suggests that they are not associated with boredom, or lack of stimulation but rather acute frustration.
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	Corporate Author				Thesis
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	307
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	Author	Saslow, C.A.
	Title	Understanding the perceptual world of horses			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	78	Issue	2-4	Pages	209-224
	Keywords	Horse; Perception; Vision; Olfaction; Touch; Hearing; Pain; Training; Psychophysics; Umwelt
	Abstract	From the viewpoint of experimental psychology, there are two problems with our current knowledge of equine perception. The first is that the behavioral and neurophysiological research in this area has enormous gaps, reflecting that this animal is not a convenient laboratory subject. The second is that the horse, having been a close companion to humans for many millennia, entrenched anecdotal wisdom is often hard to separate from scientific fact. Therefore, any summary at present of equine perception has to be provisional. The horse appears to have developed a visual system particularly sensitive to dim light and movement, it may or may not have a weak form of color vision in part of the retina, it has little binocular overlap, and its best acuity is limited to a restricted horizontal band which is aimed primarily by head/neck movements. However, the total field of view is very large. Overall, as would be expected for a prey animal, horse vision appears to have evolved more for detection of predator approach from any angle than for accurate visual identification of stationary objects, especially those seen at a distance. It is likely that, as for most mammals except the primates, horses rely more heavily on their other senses for forming a view of their world. Equine high-frequency hearing extends far above that of humans, but horses may be less able to localize the point of origin of brief sounds. The horse's capacity for chemoreception and its reliance on chemical information for identification may more closely resemble that of the dog than of the human. Its tactile sensitivity is high, and the ability of its brain and body to regulate pain perception appears to be similar to that found in other mammals. There is room for a great deal of future research in both the area of equine perception and sensory-based cognition, but for the present time persons interacting with this animal should be made aware of the importance of the sounds they make, the movements of their bodies, the way they touch the animal, and the odors they emit or carry on their clothing.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	400
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	Author	Kendrick, K.M.
	Title	Intelligent perception			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	57	Issue	3-4	Pages	213-231
	Keywords	Intelligent perception; Environmental changes; Primates
	Abstract	For an animal from any species to exhibit intelligent perception it must be capable of being consciously aware of what it perceives and capable of learning from this experience. Although many organisms, and for that matter machines, are capable of rapid adaptive learning in response to perception of environmental changes, such adaptations can occur without them being consciously aware either of external stimuli or their response to them. While behavioural and neurophysiological evidence suggests that, apart from ourselves, other higher primates must also be capable of such awareness, an important central question is whether such awareness is a characteristic of primate evolution or if it also occurs in sub-primate mammals as well. In this review I will examine our behavioural and neurophysiological evidence from visual and olfactory recognition studies in the sheep to support the argument that they are likely to be aware of and learn about both social and non-social objects and that they are therefore capable of intelligent perception. However, the impact of motivational changes on these perceptual processes suggests that they may be limited in terms of both prospection and retrospection and dealing with symbolic associations.
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	Publisher		Place of Publication		Editor
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	796
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	Author	Pick, D.F.; Lovell, G.; Brown, S.; Dail, D.
	Title	Equine color perception revisited			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	42	Issue	1	Pages	61-65
	Keywords	Equine; Color perception; Dichromat
	Abstract	An attempt to replicate Grzimek (1952; Z. Tierpsychol., 27: 330-338) is reported where a Quarter-Horse mare chose between colored and gray stimuli for food reinforcement. Stimuli varied across a broad range of reflectance values. A double-blind procedure with additional controls for auditory, olfactory, tactile, and position cues was used. The subject could reliably discriminate blue (462 nm) vs. gray, and red (700 nm) vs. gray without regard to reflectance (P<0.001), but could not discriminate green (496 nm) vs. gray. It is suggested that horses are dichromats in a manner similar to swine and cattle.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4368
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	Author	Vallortigara, G.; Rogers, L.J.
	Title	Survival with an asymmetrical brain: advantages and disadvantages of cerebral lateralization			Type	Journal Article
	Year	2005	Publication	The Behavioral and Brain Sciences	Abbreviated Journal	Behav Brain Sci
	Volume	28	Issue	4	Pages	575-89; discussion 589-633
	Keywords	Animals; Attention/physiology; Behavior/physiology; Behavior, Animal/physiology; Dominance, Cerebral/physiology; *Evolution; Humans; Models, Biological; Visual Perception/physiology
	Abstract	Recent evidence in natural and semi-natural settings has revealed a variety of left-right perceptual asymmetries among vertebrates. These include preferential use of the left or right visual hemifield during activities such as searching for food, agonistic responses, or escape from predators in animals as different as fish, amphibians, reptiles, birds, and mammals. There are obvious disadvantages in showing such directional asymmetries because relevant stimuli may be located to the animal's left or right at random; there is no a priori association between the meaning of a stimulus (e.g., its being a predator or a food item) and its being located to the animal's left or right. Moreover, other organisms (e.g., predators) could exploit the predictability of behavior that arises from population-level lateral biases. It might be argued that lateralization of function enhances cognitive capacity and efficiency of the brain, thus counteracting the ecological disadvantages of lateral biases in behavior. However, such an increase in brain efficiency could be obtained by each individual being lateralized without any need to align the direction of the asymmetry in the majority of the individuals of the population. Here we argue that the alignment of the direction of behavioral asymmetries at the population level arises as an “evolutionarily stable strategy” under “social” pressures occurring when individually asymmetrical organisms must coordinate their behavior with the behavior of other asymmetrical organisms of the same or different species.
	Address	Department of Psychology and B.R.A.I.N. Centre for Neuroscience, University of Trieste, 34123 Trieste, Italy. vallorti@univ.trieste.it
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0140-525X	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16209828			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4622
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	Author	Hampton, R.R.; Shettleworth, S.J.
	Title	Hippocampal lesions impair memory for location but not color in passerine birds			Type	Journal Article
	Year	1996	Publication	Behavioral neuroscience	Abbreviated Journal	Behav Neurosci
	Volume	110	Issue	4	Pages	831-835
	Keywords	Animals; Appetitive Behavior/physiology; Birds/physiology; Brain Mapping; Color Perception/physiology; Discrimination Learning/physiology; Hippocampus/physiology; Long-Term Potentiation/physiology; Mental Recall/physiology; Orientation/*physiology; Species Specificity
	Abstract	The effects of hippocampal complex lesions on memory for location and color were assessed in black-capped chickadees (Parus atricapillus) and dark-eyed juncos (Junco hyemalis) in operant tests of matching to sample. Before surgery, most birds were more accurate on tests of memory for location than on tests of memory for color. Damage to the hippocampal complex caused a decline in memory for location, whereas memory for color was not affected in the same birds. This dissociation indicates that the avian hippocampus plays an important role in spatial cognition and suggests that this brain structure may play no role in working memory generally.
	Address	Department of Psychology, University of Toronto, Ontario, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7044	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:8864273			Approved	no
	Call Number	refbase @ user @			Serial	376
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	Author	Heffner, R.S.; Heffner, H.E.
	Title	Localization of tones by horses: use of binaural cues and the role of the superior olivary complex			Type	Journal Article
	Year	1986	Publication	Behavioral Neuroscience	Abbreviated Journal	Behav Neurosci
	Volume	100	Issue	1	Pages	93-103
	Keywords	Animals; Auditory Pathways/physiology; Auditory Perception/physiology; Avoidance Learning/physiology; Brain Mapping; Electroshock; Female; Horses/physiology; Male; Olivary Nucleus/anatomy & histology/physiology; Orientation/physiology; Pitch Perception/physiology; Sound Localization/physiology
	Abstract	The ability of horses to use binaural time and intensity difference cues to localize sound was assessed in free-field localization tests by using pure tones. The animals were required to discriminate the locus of a single tone pip ranging in frequency from 250 Hz to 25 kHz emitted by loudspeakers located 30 degrees to the left and right of the animals' midline (60 degrees total separation). Three animals were tested with a two-choice procedure; 2 additional animals were tested with a conditioned avoidance procedure. All 5 animals were able to localize 250 Hz, 500 Hz, and 1 kHz but were completely unable to localize 2 kHz and above. Because the frequency of ambiguity for the binaural phase cue delta phi for horses in this test was calculated to be 1.5 kHz, these results indicate that horses can use binaural time differences in the form of delta phi but are unable to use binaural intensity differences. This finding was supported by an unconditioned orientation test involving 4 additional horses, which showed that horses correctly orient to a 500-Hz tone pip but not to an 8-kHz tone pip. Analysis of the superior olivary complex, the brain stem nucleus at which binaural interactions first take place, reveals that the lateral superior olive (LSO) is relatively small in the horse and lacks the laminar arrangement of bipolar cells characteristic of the LSO of most mammals that can use binaural delta I.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7044	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3954885			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5634
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	Author	Zentall, T.R.
	Title	Mental time travel in animals: a challenging question			Type	Journal Article
	Year	2006	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	72	Issue	2	Pages	173-183
	Keywords	Animals; Behavior, Animal; Columbidae; Concept Formation; Conditioning, Operant; Imagination; Memory; Mental Recall; Planning Techniques; Rats; Time Perception; Transfer (Psychology)
	Abstract	Humans have the ability to mentally recreate past events (using episodic memory) and imagine future events (by planning). The best evidence for such mental time travel is personal and thus subjective. For this reason, it is particularly difficult to study such behavior in animals. There is some indirect evidence, however, that animals have both episodic memory and the ability to plan for the future. When unexpectedly asked to do so, animals can report about their recent past experiences (episodic memory) and they also appear to be able to use the anticipation of a future event as the basis for a present action (planning). Thus, the ability to imagine past and future events may not be uniquely human.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16466863			Approved	no
	Call Number	refbase @ user @			Serial	218
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	Author	Zentall, T.R.
	Title	Timing, memory for intervals, and memory for untimed stimuli: the role of instructional ambiguity			Type	Journal Article
	Year	2005	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	70	Issue	3	Pages	209-222
	Keywords	Animals; Attention; Columbidae; Discrimination Learning; Memory, Short-Term; Practice (Psychology); Reinforcement Schedule; Retention (Psychology); *Time Perception
	Abstract	Theories of animal timing have had to account for findings that the memory for the duration of a timed interval appears to be dramatically shorted within a short time of its termination. This finding has led to the subjective shortening hypothesis and it has been proposed to account for the poor memory that animals appear to have for the initial portion of a timed interval when a gap is inserted in the to-be-timed signal. It has also been proposed to account for the poor memory for a relatively long interval that has been discriminated from a shorter interval. I suggest here a simpler account in which ambiguity between the gap or retention interval and the intertrial interval results in resetting the clock, rather than forgetting the interval. The ambiguity hypothesis, together with a signal salience mechanism that determines how quickly the clock is reset at the start of the intertrial interval can account for the results of the reported timing experiments that have used the peak procedure. Furthermore, instructional ambiguity rather than memory loss may account for the results of many animal memory experiments that do not involve memory for time.
	Address	Department of Psychology, University of Kentucky, 202B Kastle Hall, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16095851			Approved	no
	Call Number	refbase @ user @			Serial	222
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	Author	Mrosovsky, N.; Shettleworth, S.J.
	Title	Further studies of the sea-finding mechanism in green turtle hatchlings			Type	Journal Article
	Year	1974	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	51	Issue	3-4	Pages	195-208
	Keywords	Animals; Animals, Newborn/physiology; Contact Lenses; Locomotion; Orientation; Retina/physiology; Turtles/physiology; Visual Fields; Visual Perception; Water
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0005-7959	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:4447586			Approved	no
	Call Number	refbase @ user @			Serial	389
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