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Smith, J.E.; Kolowski, J.M.; Graham, K.E.; Dawes, S.E.; Holekamp, K.E. Social and ecological determinants of fission-fusion dynamics in the spotted hyaena 2008 Animal Behaviour. 4676 76 619-636 details   doi
Smith, J.E.; Van Horn, R.C.; Powning, K.S.; Cole, A.R.; Graham, K.E.; Memenis, S.K.; Holekamp, K.E. Evolutionary forces favoring intragroup coalitions among spotted hyenas and other animals 2010 Behavioral Ecology 5285 21 284-303 details   doi
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