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Author Alexander, R, MCN et al doi  openurl
  Title Fast locomotion of some african ungulates Type Journal Article
  Year (up) 1977 Publication Journal of Zoology Abbreviated Journal J Zool  
  Volume 183 Issue 3 Pages 291-300  
  Keywords  
  Abstract ABSTRACT

Ten species of ungulate were filmed, galloping in their natural habitat. They ranged in size from Thomson's gazelle (about 20 kg) to giraffe (about 1000 kg). They were pursued to make them run as fast as possible. The films have been analysed to determine speed, stride frequency, stride and step lengths, and duty factors. The dependence of these quantities on body size is discussed.



Summary:

Fast locomotion of zebra, giraffe, warthog and seven species of Bovidae has been studied. The animals were filmed from a pursuing vehicle while galloping in their natural habitat.

Stride frequency was more closely correlated with limb length (represented by hip height) than with body mass. Mean stride frequency was proportional to (hip height)-0·51 and maximum stride frequency to (hip height) -0·63.

Maximum speed was between 10 and 14 m s -1 for all species except buffalo (7 m s -1). It was not significantly correlated with body mass.

Since the small species ran at least as fast as the large ones they attained higher Froude numbers. Relative stride length was approximately 1·8 (Froude number)0·39 for all species, irrespective of size. Relative step length was approximately 0·65 (Froude number)0·2, both for the fore feet and for the hind ones. The vertical forces exerted by the feet are proportional to (body weight)×(Froude number)0·2 so the forces at maximum speed are larger multiples of body weight for small species than for large ones.
 
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  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes from Prof. Hans Klingels Equine Reference List Approved no  
  Call Number refbase @ user @ Serial 130  
Permanent link to this record
 

 
Author Clutton-Brock, T.H.; Harvey, P.H. doi  openurl
  Title Primates, brains and ecology Type Journal Article
  Year (up) 1980 Publication Journal of Zoology Abbreviated Journal J. Zool. Lond.  
  Volume 190 Issue 3 Pages 309-323  
  Keywords  
  Abstract The paper examines systematic relationships among primates between brain size (relative to body size) and differences in ecology and social system. Marked differences in relative brain size exist between families. These are correlated with inter-family differences in body size and home range size. Variation in comparative brain size within families is related to diet (folivores have comparatively smaller brains than frugivores), home range size and possibly also to breeding system. The adaptive significance of these relationships is discussed.  
  Address  
  Corporate Author Thesis  
  Publisher Blackwell Publishing Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1469-7998 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5451  
Permanent link to this record
 

 
Author Mace, G.M.; Harvey, P.H.; Clutton-Brock, T.H. doi  openurl
  Title Brain size and ecology in small mammals Type Journal Article
  Year (up) 1981 Publication Journal of Zoology Abbreviated Journal J Zool  
  Volume 193 Issue 3 Pages 333-354  
  Keywords  
  Abstract Relative brain size (measured as gross brain size after body size effects are removed) differs systematically between families of rodents, insectivores and lagomorphs. The Sciuridae have the largest relative brain size, the Soricidae and Bathyergidae the smallest. These results are discussed and compared with previous analyses of relative brain sizes among primates and bats. These differences complicate comparisons between relative brain size across phylogenetically diverse species and attempts to relate differences in relative brain size to ecological variables. To overcome these problems, best fit relationships were estimated for each family, and values for each genus were expressed as deviations from the lines of best fit. We refer to these values as Comparative Brain Size (CBS). Differences in CBS are related to differences in habitat type (forest-dwelling genera have larger CBS' than grassland forms), in diet (folivores have smaller CBS' than generalists or insectivores, frugivores and granivores), in zonation (arboreal genera have larger CBS' than terrestrial ones) and in activity timing (nocturnal genera have larger CBS' than dirurnal ones). However, these ecological categories are interrelated and, when the effects of other ecological differences are taken into account using analyses of variance, only the differences associated with diet, and possibly habitat remain.  
  Address  
  Corporate Author Thesis  
  Publisher Blackwell Publishing Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1469-7998 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5455  
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Author Smuts, M.M.S.; Penzhorn, B. L. openurl 
  Title Descriptions of antomical differences between skulls and mandibles of Equus zebra and E. burchelli from southern Africa Type Journal Article
  Year (up) 1988 Publication South African Journal of Zoology Abbreviated Journal South African Journal of Zoology  
  Volume 23 Issue (4)3 Pages 328-336  
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  Language Summary Language Original Title  
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  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes from Professor Hans Klingels Equine Reference List Approved yes  
  Call Number Serial 1617  
Permanent link to this record
 

 
Author VanDierendonck, M.C.; de Vries, H.; Schilder, M.B.H. url  openurl
  Title An Analysis of Dominance, Its Behavioural Parameters and Possible Determinants in a Herd of Icelandic orses in Captivity Type Journal Article
  Year (up) 1995 Publication Netherlands Journal of Zoology Abbreviated Journal Netherl. J. Zool.  
  Volume 45 Issue 3-4 Pages 362-385  
  Keywords Dominance; rank order; horses; Icelandic horses.  
  Abstract Th e applicability of the concept of dominance was investigated in a captive herd of  Icelandic

horses and  ponies of diff erent breeds. Eight out of  behaviours possibly related

to dominance occurred frequently enough to be investigated in detail. For these eight agonistic

behaviours the coverage, the unidirectionality in the exchange, and the degree of

transitivity (Landau`s linearity index) were calculated. Four off ensive behaviours, together

with avoidance, were suitable for further analysis with regard to dominance. Th e patterns

of asymmetries with which these behaviours were exchanged were suffi ciently similar as to

justify the application of the dominance concept and to construct a (nearly) linear dominance

hierarchy. Th e rank order of the castrated stallions was completely linear, the hierarchy

of the mares was almost completely linear. Th e results suggest that off ensive and defensive

aggressive behaviours should be treated separately and that the concept of dominance

is applicable. However, ritualized formal dominance signals between adult horses appear to

be (almost) absent. Th e rank positions of the individuals were correlated with age and residency

in the herd but not with height. Middle ranking horses tended to be more frequently

in the close vicinity of another horse than high ranking or low ranking horses. Over and

above this correlation at the individual level, it was found that pairs of horses close in rank

to each other were more often also spatially close to each other. Being in oestrus did not infl

uence the dominance relationships between mares. For castrated stallions the rank positions

were correlated with the age at which they were castrated. Th is suggests that in male

horses experience prior to neutering infl uences the behaviour afterwards.
 
  Address  
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  Publisher Place of Publication Editor  
  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number refbase @ user @ Serial 440  
Permanent link to this record
 

 
Author VanDierendonck, M.C., de Vries, H., Schilder, M.B.H. url  doi
openurl 
  Title An Analysis of Dominance, Its Behavioural Parameters and Possible Determinants in a Herd of Icelandic horses in captivity Type Journal Article
  Year (up) 1995 Publication Netherlands Journal of Zoology Abbreviated Journal Netherl. J. Zool.  
  Volume 45 Issue 3-4 Pages 362-385  
  Keywords Dominance; rank order; horses; Icelandic horses.  
  Abstract Feral horses are social animals, which have to rely on survival strategies centered on the formation of cohesive social bonds within their bands. Many problems in the husbandry of social animals such as horses, are due to the fact that the limits of their adaptive abilities are exceeded. Evidence suggests that the fundamental social characteristics of domestic horses have remained relatively unchanged. The social structure, social strategies and social interactions were investigated (3 non-consecutive years, 24 hr per day for several weeks) in long term established groups of domestic horses (mares and geldings of all ages) and a few small introduced groups, kept in (semi)natural environments. The general aim was to investigate the social needs of domestic horses. The social life of domestic horses was characterised by long lasting bonds with preferred partners which were established and maintained by allogrooming, play, proximity and dominance behaviours. Bonding partners were mainly found within the same sex-age group, but adult geldings also bonded with sub-adult mares and geldings. Adult mares were clustered in a group, while the other animals formed a second group. Among the adult mares, subgroups according to reproductive state were formed. Individuals regulated their social network by interfering with interactions between other members of the herd, which in itself is complex. An intervention is a behavioural action of one animal that actively interferes with an ongoing interaction between a dyad with the apparent aim of altering that interaction. This was verified by post-hoc analyses of disturbed and undisturbed interactions. Interventions in allogrooming or play were performed significantly more often when at least one member of the initial dyad was a preferred partner of, or familiar to (within the small introduced bands) the intervener. The stronger the preferred association in allogrooming between the intervener and member(s) of the initial dyad, the higher the probability the intervener would displace one initial member and continue allogrooming with the other. Just five behaviours were extracted which reliably reflected the dominance relations among horses. Aggression with the hind quarters was used both offensively and defensively and therefore not suitable as a reliable parameter. Individual dominance relationships were related to social experience. The implications of these findings for horse husbandry were assessed. It is argued that the execution of affiliative behaviours may be rewarding in itself, and therefore always will be a highly motivated behaviour. It is shown that social positive physical interactions (allogrooming, play) with other horses is an ethological need and therefore indispensable in modern husbandry systems. Ethological needs are so important for the animal that husbandry systems that lack the possibilities to execute such behaviours will cause chronic stress. It is concluded that all horses need physical social contact, and that horses, which lack appropriate social learning experiences during ontogeny, may be hampered in their social functioning later in life. Solutions for problems, including dominance problems, in individual social housing and group housing are presented.  
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  Series Editor Series Title Abbreviated Series Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 2368  
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Author Weckerly, Floyd W. url  doi
openurl 
  Title Social bonding and aggression in female Roosevelt elk Type Journal Article
  Year (up) 1999 Publication Canadian Journal of Zoology Abbreviated Journal Can J Zool  
  Volume 77 Issue 9 Pages 1379-1384  
  Keywords  
  Abstract Abstract: The relationship between degree of social bonding (extent of association among individuals) and level of aggression in ruminants is unclear. I examined social bonding and aggression in three groups of female Roosevelt elk (Cervus elaphus roosevelti) over 2 years. I hypothesized that when animals are socially bonded, bouts of aggression will be won by the individual initiating the aggression, occur quickly, and involve little physical contact, and the level of aggression does not correlate with group size. The degree of social bonding was high among individuals in all groups. Dyads of known individuals were together >80% of the time. A permutation analysis indicated that groups with the observed sizes had <0.001 chance of random association, except on one occasion when the probability was 0.72 for one group. Using focal-animal sampling, aggressive interactions were won 72% of the time by the initiator, occurred quickly (<5 s), and involved little physical contact, and the level of aggression was not correlated with group size. The level of aggression was, however, significantly lower in one of the groups. This group may have had access to more abundant food resources than the other groups. Socially bonded elk conducted aggressive interactions in a fashion that did not disrupt social stability. Résumé : La relation entre le degré de liaison sociale (importance des associations entre individus) et l`agressivité n`est pas claire chez les ruminants. J`ai étudié les liaisons sociales et l`agressivité chez trois groupes de femelles du Cerf de Roosevelt (Cervus elaphus roosevelti) pendant 2 ans. J`ai posé en hypothèse que, chez les animaux liés socialement, la victoire devrait être emportée par l`individu qui entreprend l`agression, l`agression devrait être de courte durée, se faire avec peu de contacts physiques et la fréquence des agressions ne devrait pas être liée à la taille du groupe. Des paires d`individus passaient plus de 80% de leur temps ensemble. Une analyse des permutations a démontré que, chez les groupes des tailles observées, la probabilité d`une association aléatoire était de moins de 0,001, sauf en un cas où cette probabilité a été évaluée à 0,72 chez un groupe. Par échantillonnage directionnel, j`ai observé que les interactions agressives étaient gagnées par l`individu attaquant 72% du temps, étaient de courte durée (<5 s), se faisaient avec peu de contacts physiques et leur fréquence n`était pas reliée à la taille du groupe. Il y avait cependant moins d`agressivité chez l`un des groupes. Il se peut que ce groupe ait eu accès à plus de ressources alimentaires que les autres. Chez les cerfs liés par des liens sociaux, l`agressivité ne se manifeste pas de façon à déséquilibrer la stabilité sociale.  
  Address  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4268  
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Author Peterson R.O.; Jacobs A.K.; Drummer T.D.; Mech L.D.; Smith D.W. url  openurl
  Title Leadership behavior in relation to dominance and reproductive status in gray wolves, Canis lupus Type Journal Article
  Year (up) 2002 Publication Canadian Journal of Zoology Abbreviated Journal  
  Volume 80 Issue Pages 1405-1412  
  Keywords  
  Abstract We analyzed the leadership behavior of breeding and nonbreeding gray wolves (Canis lupus) in three packs during winter in 1997-1999. Scent-marking, frontal leadership (time and frequency in the lead while traveling), initiation of activity, and nonfrontal leadership were recorded during 499 h of ground-based observations in Yellowstone National Park. All observed scent-marking (N = 158) was done by breeding wolves, primarily dominant individuals. Dominant breeding pairs provided most leadership, consistent with a trend in social mammals for leadership to correlate with dominance. Dominant breeding wolves led traveling packs during 64% of recorded behavior bouts (N = 591) and 71% of observed travel time (N = 64 h). During travel, breeding males and females led packs approximately equally, which probably reflects high parental investment by both breeding male and female wolves. Newly initiated behaviors (N = 104) were prompted almost 3 times more often by dominant breeders (70%) than by nonbreeders (25%). Dominant breeding females initiated pack activities almost 4 times more often than subordinate breeding females (30 vs. 8 times). Although one subordinate breeding female led more often than individual nonbreeders in one pack in one season, more commonly this was not the case. In 12 cases breeding wolves exhibited nonfrontal leadership. Among subordinate wolves, leadership behavior was observed in subordinate breeding females and other individuals just prior to their dispersal from natal packs. Subordinate wolves were more often found leading packs that were large and contained many subordinate adults.

Nous avons analysé le comportement de commandement chez des loups gris (Canis lupus) reproducteurs et non reproducteurs appartenant à  trois meutes durant les hivers de 1997-1999. Le marquage d'odeurs, la position en tête de meute (la durée et la fréquence au cours des déplacements), l'initiation des activités et la prise de décisions ailleurs qu'en tête du groupe ont été notés pendant 499 h d'observations au sol dans le Parc national de Yellowstone. Tous les marquages (N = 158) ont été faits par des loups reproducteurs, surtout des individus dominants. Ce sont surtout les couples dominants qui assurent le commandement, en accord avec une tendance chez les mammifères sociaux chez lesquels la fonction de chef est en corrélation avec la dominance. Les loups reproducteurs dominants ont conduit les meutes en déplacement pendant 64 % (N = 591) des épisodes de comportement et pendant 71 % des épisodes de déplacement (N = 64 h). Les mâles et les femelles reproducteurs ont dirigé les meutes en déplacement à peu près également, ce qui reflète probablement l'investissement parental important aussi bien de la part des reproducteurs mâles que des femelles. Les comportements nouveaux (N = 104) ont été adoptés presque trois fois plus souvent par des reproducteurs dominants (70 %) que par des individus non reproducteurs (25 %). Des femelles reproductrices dominantes ont été instigatrices des activités de leur meute environ quatre fois plus souvent que les femelles reproductrices subordonnées (30 vs. 8 fois). Bien qu'une femelle reproductrice subordonnée ait pris la direction de sa meute plus souvent que les individus non reproducteurs au cours d'une saison, cela n'est pas habituel. Dans 12 cas, des loups reproducteurs ont pris la direction de leur meute sans être en tête. Chez les individus subordonnés, le comportement de commandement a été observé chez des femelles reproductrices et chez d'autres individus juste avant qu'ils ne quittent leur meute d'origine au moment de la dispersion. Les loups subordonnés mènent surtout de grands troupeaux qui comptent beaucoup d'individus subordonnés.[Traduit par la Rédaction]
 
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  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4700  
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Author Reader, S.M. doi  openurl
  Title Innovation and social learning: individual variation and brain evolution Type Journal Article
  Year (up) 2003 Publication Animal Biology (formerly Netherlands Journal of Zoology) Abbreviated Journal Anim. Biol. Leiden.  
  Volume 53 Issue 2 Pages 147-158  
  Keywords  
  Abstract This paper reviews behavioural, neurological and cognitive correlates of innovation at the individual, population and species level, focusing on birds and primates. Innovation, new or modified learned behaviour not previously found in the population, is the first stage in many instances of cultural transmission and may play an important role in the lives of animals with generalist or opportunistic lifestyles. Within-species, innovation is associated with low neophobia, high neophilia, and with high social learning propensities. Indices of innovatory propensities can be calculated for taxonomic groups by counting the frequency of reports of innovation in published literature. These innovation rate data provide a useful comparative measure for studies of behavioural flexibility and cognition. Innovation rate is positively correlated with the relative size of association areas in the brain, namely the hyperstriatum ventrale and neostriatum in birds, and the neocortex and striatum in primates. Innovation rate is also positively correlated with the reported variety of tool use, as well as interspecific differences in learning. Current evidence thus suggests similar patterns of cognitive evolution in primates and birds.  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 3395  
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Author de Villiers, M.S.; Richardson, P.R.K.; van Jaarsveld, A.S. url  doi
openurl 
  Title Patterns of coalition formation and spatial association in a social carnivore, the African wild dog (Lycaon pictus) Type Journal Article
  Year (up) 2003 Publication Journal of Zoology Abbreviated Journal J Zool  
  Volume 260 Issue 4 Pages 377-389  
  Keywords coalitions; social systems; Carnivora; Lycaon pictus  
  Abstract In many social species, relationships within groups seem to be non-random but related to variables such as rank, kinship or sexual attractiveness. The endangered African wild dog Lycaon pictus is a social carnivore that lives in large, stable packs, and intra-pack associations might be expected to display similar patterns. We investigated patterns of coalition formation (support during dominance interactions, and partnership interactions) and resting associations between members of a captive pack of 19 wild dogs. The social organization of the captive pack was similar to that of free-ranging packs in many respects. Polyadic (group) incidents of coalition support were also observed in a free-ranging pack. Patterns of coalition formation in the captive pack were related to rank. Most aggressive interactions involved high-ranking individuals (particularly the alpha, beta and third-ranking males) and coalitionary support tended to reinforce the existing hierarchy. However, there was at least one example of support influencing a successful rank challenge. Support was affected by potential risks and benefits, the latter including dominance through association and revolutionary alliances. An even stronger pattern overlaid associations between pack members: coalitions and resting associations were strongest between members of the same age–sex cohort, and may have enabled the eventual dominance of younger pack members over adults. Among adults, coalitionary associations were sometimes overridden by intersexual relationships. The results from this captive pack suggest that wild dogs are sensitive to differences in competitive ability. This information, in conjunction with strong affiliative bonds between littermates, is used to manoeuvre for position in the social hierarchy. It may also be important during dispersal, in encounters with other dispersing groups of the same sex. Although most features of the social structure of the captive pack were comparable to those of free-ranging packs, aspects such as the influence of relatedness on coalition formation still need to be explored.  
  Address  
  Corporate Author Thesis  
  Publisher Blackwell Publishing Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1469-7998 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5249  
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