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	Author	Kacelnik, A.
	Title	The foraging efficiency of great tits (Parus major L.) in relation to light intensity			Type	Journal Article
	Year	1979	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	27	Issue	Part 1	Pages	237-241
	Keywords
	Abstract	I report an experiment aimed at testing whether foraging efficiency of great tits is limited by light intensity at the time of the dawn chorus. Captive great tits hunting for prey under different luminance conditions were less successful in finding prey when foraging, hunted for a lower proportion of their time, and handled individual prey items for longer when luminance was under approximately 7 cd/m2. This luminance is not reached in the field until after the time of the dawn chorus, suggesting that in the early morning foraging is limited by light intensity. I suggest that a satisfactory functional explanation of the dawn chorus must take into account the comparatively low foraging opportunity early in the morning, as well as the factors affecting the opportunity for singing and other territorial activities.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2125
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	Author	Clegg, H.A.; Buckley, P.; Friend, M.A.; McGreevy, P.D.
	Title	The ethological and physiological characteristics of cribbing and weaving horses			Type	Journal Article
	Year	2008	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	109	Issue	1	Pages	68-76
	Keywords	Horse; Stereotypy; Digestion; Gut transit; Stress
	Abstract	Data were gathered on the behavioural and physiological characteristics of five cribbers, six weavers and six non-stereotypic (control) mature Thoroughbred geldings for a period of 16 weeks. The horses were hired from their owners and stabled individually throughout the trial. Cribbers and weavers had been known to stereotype for at least 12 months prior to commencement of the study. Behavioural data were collected using video surveillance. Cribbers stereotyped most frequently (PÂ <Â 0.001) in the period 2-8Â h following delivery of concentrated food, reinforcing the suggestion that diet is implicated in cribbing behaviour. Weavers stereotyped most frequently (PÂ <Â 0.001) during periods of high environmental activity such as during routine pre-feeding activities and in the hour prior to daily turnout, presumably when anticipation and stimulation were at their highest levels. Cribbers and weavers took longer than control horses to fully consume their ration, suggesting possible differences in motivation to feed, distress levels, satiety mechanisms or abdominal discomfort. Physiological data were collected throughout the trial and there were no differences in oro-caecal transit time, digestibility, plasma cortisol concentration or heart rate among the three behavioural groups.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4768
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	Author	McKinley, S.; Young, R.J.
	Title	The efficacy of the model-rival method when compared with operant conditioning for training domestic dogs to perform a retrieval-selection task			Type	Journal Article
	Year	2003	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	81	Issue	4	Pages	357-365
	Keywords	Dogs; Learning; Operant conditioning; Model-rival; Cognition
	Abstract	Traditionally, dogs have been trained by operant conditioning techniques; that is, dogs make a desired behavioural response and this response is reinforced by a reward such as food. This type of training is very effective in training dogs to perform basic obedience behaviours (e.g. `stay'). However, dogs are social animals and should be predisposed to learn from social stimuli. In the present study, we used a modified version of the model-rival technique that has been extensively used in experiments investigating the cognitive ability of parrots. In this technique, social stimuli are used to create in the animal an interest in the object without the use of food or other rewards. Therefore, the animal learns the name of the object (intrinsic reward) and not that the object's name means food. In this experiment we compared the learning ability of nine pet dogs to solve the same retrieval-selection task having been previously trained using operant conditioning or model-rival techniques. The retrieval-selection task was the dogs had to correctly select the commanded object to bring to the experimenter from a group of three similar objects. The results show no difference in the speeds with which the dogs solved the test--demonstrating the efficacy of the model-rival method. This is the first time that the effectiveness of the model-rival technique has been experimentally demonstrated with dogs. Furthermore, we believe that the methodology reported in this paper has applications in dog training and in experiments into dog cognition.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2900
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	Author	Williams, J.L.; Friend, T.H.; Nevill, C.H.; Archer, G.
	Title	The efficacy of a secondary reinforcer (clicker) during acquisition and extinction of an operant task in horses			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	88	Issue	3-4	Pages	331-341
	Keywords	Equine; Horse; Clicker training; Operant conditioning; Secondary reinforcers
	Abstract	“Clicker training” is a popularly promoted training method based on operant conditioning with the use of a secondary reinforcer (the clicker). While this method draws from theories of learning and is used widely, there has been little scientific investigation of its efficacy. We used 60 horses, Equus callabus, and assigned each horse to one of six reinforcement protocols. The reinforcement protocols involved combinations of reinforcers administered (primary versus secondary plus primary), schedule of reinforcement (continuous versus variable ratio), and reinforcers applied during extinction (none or secondary). There were no differences (P>=0.11) between horses which received a secondary reinforcer (click) followed by the primary reinforcer (food) and those which received only the primary reinforcer (food) in the number of trials required to train the horses to touch their noses to a plastic cone (operant response). There also were no differences (P>=0.12) between horses which received the secondary reinforcer plus primary reinforcer and those which received only the primary reinforcer in regards to the number of trials to extinction. We conclude that there is no difference in the amount of training required to learn the operant task or in the task's resistance to extinction between horses that received a secondary reinforcer followed by a primary reinforcer versus horses which received only a primary reinforcer.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3581
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	Author	Waran, N.K.; Clarke, N.; Farnworth, M.
	Title	The effects of weaning on the domestic horse (Equus caballus)			Type	Journal Article
	Year	2008	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	110	Issue	1-2	Pages	42-57
	Keywords	Behaviour; Foal; Horse; Stress; Weaning; Welfare
	Abstract	For free-living or feral horses weaning takes place naturally at around 8-9 months [Gill, E.L., 1988. Factors affecting body condition of New Forest Ponies. Ph.D. Thesis. Department of Biology, University of Southampton]. Some mares will continue to suckle their foal until shortly before the arrival of their next foal, gestation being approximately 342 days depending upon the breed of the horse [Ropiha, R.T., Mathews, G., Butterfield, R.M., 1969. The duration of pregnancy in Thoroughbred mares. Vet. Rec. 84, 552-555]. Under domestic conditions, weaning tends to take place earlier, typically between 4 and 6 months of age. The weaning process has been identified as associated with potential psychological, physical and nutritional stressors that are of welfare concern. Following a review of the literature it is evident that there is a need for detailed research into what should constitute best practice with respect to foal and mare welfare. In addition, there is a need to understand the potential long-term impact of weaning on, for example, trainability and later maternal behaviour, and whether the stresses associated with early weaning have detrimental effects on the performance horse. There is also a lack of clear information concerning the most frequently observed weaning practices and the reasons why certain weaning methods are chosen. Some variables should be closely managed during weaning in order to minimise stress responses. These include: early creep feeding to familiarise the young animal with the food it will be exposed to during weaning, feeding a high fibre diet and keeping the animal in extensive conditions using a gradual approach to weaning. However, we conclude that there may not be one best method for weaning, since the chosen method must take into account a number of factors including: available resources, the housing environment, the individual foal's stage of development, the strength of the mare-foal attachment, the foal's ability to cope with changes in social conditions and the ability of the horse owner to implement the chosen method. We do however suggest that the fewest stress responses appear to occur where foals are weaned gradually and allowed to have social contact either with other foals or with older horses.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Admin @ knut @			Serial	4348
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	Author	Pollard, J.C.; Littlejohn, R.P.
	Title	The effects of pen size on the behaviour of farmed red deer stags confined in yards			Type	Journal Article
	Year	1996	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	47	Issue	3-4	Pages	247-253
	Keywords	Red deer; Handling; Density; Aggression; Farming
	Abstract	To determine whether pen size affected the behaviour and welfare of farmed red deer confined temporarily in yards, four groups of ten 2-year-old stags were confined for 40 min or 2 days in each of spring and summer, in either large (5 m × 4 m ) or small (2.5 m × 4) pens. In the small pens, wall pacing and vertical/horizontal head movements at the walls were more frequently observed (P < 0.05) and were carried out by a greater percentage of the deer (P < 0.001), and distances between individuals were smaller (P < 0.01), than observations in the large pens. Aggressive activities varied seasonally, with head-butting and chasing being seen most frequently in the spring (P < 0.05) and biting and kicking being seen most frequently in the summer (P < 0.05), and the overall frequency of aggressive activities was low in summer. In spring, in small pens there were fewer threats to head-butt, head butts by moving animals, and less stepping activity than in large pens (P < 0.05). In summer, in small pens there were more threats to butt and more stepping activity than in the large pens (P < 0.05). In both seasons, aggressive activities were correlated with wall pacing (r = 0.58 and 0.55, respectively). It was concluded that the effect of pen size on the frequency and nature of aggressive and other activities varied seasonally. In order minimise aggression and stepping activity, small pens were favoured in spring and large pens were favoured in summer. However, in both seasons there were greater inter-individual distances and reduced pacing and head movements at the walls in large pens. This latter finding may indicate that the large pens were less aversive to the deer, regardless of season.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5472
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	Author	Heird, J.C.; Whitaker, D.D.; Bell, R.W.; Ramsey, C.B.; Lokey, C.E.
	Title	The effects of handling at different ages on the subsequent learning ability of 2-year-old horses			Type	Journal Article
	Year	1986	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	15	Issue	1	Pages	15-25
	Keywords
	Abstract	Forty weanling Quarter Horses, from the Pitchfork Ranch, Dickens, Texas, were randomly assigned to one of five groups based on the amount of handling they would receive. The five groups were: (1) limited handling; (2) 1 week of handling; (3) 2 weeks of handling; (4) 3 weeks of handling; (5) continuous handling for 18 months. As 2-year-olds, the horses were tested in a simple place-learning T-maze after being preconditioned. A maximum of 30 daily trials were conducted for 20 consecutive days, with feed location alternating between sides on successive days. If a horse met the criterion of 11 out of 12 correct responses with 8 correct responses being consecutive, it was retired without completing the 30 trials. Group 1 made fewer correct responses during the 20 days than any other group except Group 3. All groups achieved learning by Day 10, but the most-handled group reached a consistently higher percentage of correct responses earlier than the less-handled groups. Group 5, the group receiving the most handling, was the least emotional, as determined by the animals' reactivity to a novel stimulus, and received the highest scores for trainability after being ridden. Preconditioning trainability scores, or estimates of the horses' ability to learn prior to conditioning and testing, tended to predict maze-learning ability. These scores were also significantly correlated with post-maze training scores.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3590
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	Author	Williams, J.L.; Friend, T.H.; Toscano, M.J.; Collins, M.N.; Sisto-Burt, A.; Nevill, C.H.
	Title	The effects of early training sessions on the reactions of foals at 1, 2, and 3 months of age			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	77	Issue	2	Pages	105-114
	Keywords	Equine; Imprint training; Imprinting; Learning; Training; Handling; Foal
	Abstract	An early training procedure commonly termed “foal imprint training” is widely promoted in the horse industry. However, there have been no published scientific investigations of its efficacy. This study determined the effects of a training procedure on foals and their reaction to stimuli used in the early training procedure, and to a novel stimulus, at 1, 2 and 3 months of age. Twenty-five foals received a standard training procedure at 2, 12, 24, and 48 h after birth. After the training procedure, the foals received minimal additional handling that included veterinary treatments and occasional relocation. Twenty-two foals born over the same time period served as controls. All 47 (25 trained, 22 control) foals were tested at 1 month of age. Only 20 were available for testing at 2 months of age, and nine were available at 3 months. Percentage change from baseline heart rate, time required to complete exposure to each stimulus (foals that were more reactive took longer) and the behavior of each foal during the introduction of each stimulus were recorded. Overall, the control foals tended to receive lower (better) behavioral scores at 1 and 2 months of age. Foals that underwent the training procedure tended to require less time to complete exposure to the stimulus and had lower heart rates during exposure to the stimuli at 1 and 2 months of age. By 3 months of age, there were no significant differences between trained and control foals for any measures. Early training was not efficacious in this study.
	Address
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	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4331
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	Author	Nicol, C.J.; Badnell-Waters, A.J.; Bice, R.; Kelland, A.; Wilson, A.D.; Harris, P.A.
	Title	The effects of diet and weaning method on the behaviour of young horses			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	95	Issue	3-4	Pages	205-221
	Keywords	Horse; Diet; Weaning; Temperament test
	Abstract	The effects of diet on horse behaviour have not previously been quantified in detail. In this study, we examined the behaviour of 17 foals from the age of 2 to 40 weeks. Each foal received either a starch and sugar (SS) diet or a fat and fibre (FF) diet. The two diets contained similar digestible energy, crude protein and micronutrients, but differed in the fat and non-structural carbohydrate balance. The baseline behaviour of the foals was observed every 2 weeks by focal animal sampling. Additional behavioural observations were conducted when the foals were weaned by one of two methods. Approximately 2 months after weaning, the temperament and tractability of the young horses was assessed using standardised tests. Responses to a novel object, to a novel person, and during a handling test were observed and quantified. Horses grew well on both diets with no apparent effects of diet on growth rate or baseline behaviour. Immediately after weaning, horses receiving the FF diet cantered less frequently (F = 5.10; p < 0.05), for a shorter duration (F = 7.23; p < 0.05) and appeared to be more settled. Foals that were barn-weaned appeared more stressed than foals that were paddock-weaned. In the temperament tests, horses receiving the FF diet spent significantly more time investigating (F = 6.78; p < 0.05), and less time looking at (F = 7.93; p < 0.05), the novel object than horses receiving the SS diet. They also spent less time walking away from the novel person (F = 5.16; p < 0.05) and their time taken to complete the handling test was significantly lower (F = 8.72; p = 0.01). Overall, the horses that received the FF diet appeared less distressed immediately after weaning and seemed calmer and more inquisitive during a range of temperament tests.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3642
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	Author	Nicol, C.J.; Pope, S.J.
	Title	The effects of demonstrator social status and prior foraging success on social learning in laying hens			Type	Journal Article
	Year	1999	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	57	Issue	1	Pages	163-171
	Keywords
	Abstract	Opportunities for social learning within a group of animals are likely to be influenced by the social dynamics of that group. Some individuals may be more influential demonstrators than others even when there are no differences in their skill level or performance. In this study of domestic hens,Gallus gallus domesticus, differences in demonstrator salience were examined. From 24 separate flocks we selected as demonstrators a dominant cockerel, a dominant hen, a mid-ranking hen or a subordinate hen. Demonstrators were pretrained to perform an operant discrimination task to obtain food. Six observers from each flock individually watched the demonstrator perform the task for four 5-min sessions held on consecutive days. On the fifth day observers were tested individually in the operant chamber. We analysed data from 19 flocks, where there were no quantitative differences in demonstrator performance. Observer hens of relatively high social status performed more correct operant pecks than observer hens of relatively low social status. Demonstrator category also had a significant effect on subsequent observer behaviour. Hens that had observed cockerels performed very few general pecks or operant pecks. Hens that had observed dominant hens performed more operant pecks, but hens that had observed sub-ordinate hens performed more general pecks in the chamber. The results suggested either that there was an interaction between dominance and gender in demonstrator salience or that dominant hens might have been influential because of some factor imperfectly associated with their dominance status. A possible candidate was the foraging ability of the dominant hens. In a second experiment using the same protocol, we manipulated the prior foraging success of dominant hens from four additional flocks but this had no significant effect on their subsequent influence as demonstrators.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	715
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