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	Author	Kelly, D.M.; Spetch, M.L.
	Title	Pigeons encode relative geometry			Type	Journal Article
	Year	2001	Publication	Journal of Experimental Psychology. Animal Behavior Processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	27	Issue	4	Pages	417-422
	Keywords	Animals; Behavior, Animal/physiology; Cognition/physiology; Columbidae; Discrimination Learning/physiology; Form Perception/physiology; Space Perception/*physiology
	Abstract	Pigeons were trained to search for hidden food in a rectangular environment designed to eliminate any external cues. Following training, the authors administered unreinforced test trials in which the geometric properties of the apparatus were manipulated. During tests that preserved the relative geometry but altered the absolute geometry of the environment, the pigeons continued to choose the geometrically correct corners, indicating that they encoded the relative geometry of the enclosure. When tested in a square enclosure, which distorted both the absolute and relative geometry, the pigeons randomly chose among the 4 corners, indicating that their choices were not based on cues external to the apparatus. This study provides new insight into how metric properties of an environment are encoded by pigeons.
	Address	Department of Psychology, University of Alberta, Edmonton, Alberta, Canada, T6G 2E9. kelly@bio.psy.ruhr-uni-bochum.de
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11676090			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2770
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	Author	Merchant, H.; Fortes, A.F.; Georgopoulos, A.P.
	Title	Short-term memory effects on the representation of two-dimensional space in the rhesus monkey			Type	Journal Article
	Year	2004	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	7	Issue	3	Pages	133-143
	Keywords	Analysis of Variance; Animals; Discrimination Learning/physiology; Macaca mulatta; Male; Memory, Short-Term/physiology; Mental Processes/physiology; Pattern Recognition, Visual/physiology; Space Perception/*physiology
	Abstract	Human subjects represent the location of a point in 2D space using two independent dimensions (x-y in Euclidean or radius-angle in polar space), and encode location in memory along these dimensions using two levels of representation: a fine-grain value and a category. Here we determined whether monkeys possessed the ability to represent location with these two levels of coding. A rhesus monkey was trained to reproduce the location of a dot in a circle by pointing, after a delay period, on the location where a dot was presented. Five different delay periods (0.5-5 s) were used. The results showed that the monkey used a polar coordinate system to represent the fine-grain spatial coding, where the radius and angle of the dots were encoded independently. The variability of the spatial response and reaction time increased with longer delays. Furthermore, the animal was able to form a categorical representation of space that was delay-dependent. The responses avoided the circumference and the center of the circle, defining a categorical radial prototype around one third of the total radial length. This radial category was observed only at delay durations of 3-5 s. Finally, the monkey also formed angular categories with prototypes at the obliques of the quadrants of the circle, avoiding the horizontal and vertical axes. However, these prototypes were only observed at the 5-s delay and on dots lying on the circumference. These results indicate that monkeys may possess spatial cognitive abilities similar to humans.
	Address	Brain Sciences Center (11B), Veterans Affairs Medical Center, One Veterans Drive, MN 55417, Minneapolis, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14669074			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2548
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	Author	Petruso, E.J.; Fuchs, T.; Bingman, V.P.
	Title	Time-space learning in homing pigeons (Columba livia): orientation to an artificial light source			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	2	Pages	181-188
	Keywords	Animals; Circadian Rhythm; Columbidae/physiology; Homing Behavior/physiology; Learning/physiology; Light; Orientation/physiology; Space Perception/physiology; Time Perception/physiology
	Abstract	Time-space learning reflects an ability to represent in memory event-stimulus properties together with the place and time of the event; a capacity well developed in birds. Homing pigeons were trained in an indoor octagonal arena to locate one food goal in the morning and a different food goal in the late afternoon. The goals differed with respect to their angular/directional relationship to an artificial light source located outside the arena. Further, the angular difference in reward position approximated the displacement of the sun's azimuth that would occur during the same time period. The experimental birds quickly learned the task, demonstrating the apparent ease with which birds can adopt an artificial light source to discriminate among alternative spatial responses at different times of the day. However, a novel midday probe session following successful learning revealed that the light source was interpreted as a stable landmark and not as a surrogate sun that would support compass orientation. Probe sessions following a phase shift of the light-dark cycle revealed that the mechanism employed to make the temporal discrimination was prevailingly based on an endogenous circadian rhythm and not an interval timing mechanism.
	Address	Department of Psychology and J.P. Scott Center for Neuroscience, Mind and Behavior Bowling Green State University, Bowling Green, OH 43403, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17160343			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2432
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	Author	Shettleworth, S.J.; Krebs, J.R.
	Title	How marsh tits find their hoards: the roles of site preference and spatial memory			Type	Journal Article
	Year	1982	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	8	Issue	4	Pages	354-375
	Keywords	Animals; Appetitive Behavior; Birds; Cues; Discrimination Learning; Memory; Mental Recall; Orientation; *Space Perception
	Abstract	Marsh tits (Parus palustris) store single food items in scattered locations and recover them hours or days later. Some properties of the spatial memory involved were analyzed in two laboratory experiments. In the first, marsh tits were offered 97 sites for storing 12 seeds. They recovered a median of 65% of them 2-3 hr later, making only two errors per seed while doing so. Over trials, they used some sites more often than others, but during recovery they were more likely to visit a site of any preference value if they had stored a seed there that day than if they had not. Recovery performance was much worse if the experimenters moved the seeds between storage and recovery. A fixed search strategy that had some of the same average properties as the tits' search behavior also did worse than the real birds. In Experiment 2, any tendency to visit the same sites on successive daily tests in the aviary was placed in opposition to memory for storage sites by allowing the tits to store more seeds 2 hr after storing a first batch. They tended to avoid individual storage sites holding seeds from the first batch. When the tits searched for all the seeds 2 hr later, they tended to recover more seeds from the second batch than from the first, i.e., there was a recency effect.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7175447			Approved	no
	Call Number	refbase @ user @			Serial	385
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	Author	Shettleworth, S.J.; Sutton, J.E.
	Title	Multiple systems for spatial learning: dead reckoning and beacon homing in rats			Type	Journal Article
	Year	2005	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	31	Issue	2	Pages	125-141
	Keywords	Animals; Behavior, Animal; Cues; Feeding Behavior; Habituation, Psychophysiologic; Homing Behavior; Learning; Male; Rats; Rats, Long-Evans; Space Perception
	Abstract	Rats homed with food in a large lighted arena. Without visual cues, they used dead reckoning. When a beacon indicated the home, rats could also use the beacon. Homing did not differ in 2 groups of rats, 1 provided with the beacon and 1 without it; tests without the beacon gave no evidence that beacon learning overshadowed dead reckoning (Experiment 1). When the beacon was at the home for 1 group and in random locations for another, there was again no evidence of cue competition (Experiment 2). Dead reckoning experience did not block acquisition of beacon homing (Experiment 3). Beacon learning and dead reckoning do not compete for predictive value but acquire information in parallel and are used hierarchically.
	Address	Department of Psychology, University of Toronto, Toronto, ON M5S 3G3, Canada. shettle@psych.utoronto.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15839771			Approved	no
	Call Number	refbase @ user @			Serial	364
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	Author	Shettleworth, S.J.; Westwood, R.P.
	Title	Divided attention, memory, and spatial discrimination in food-storing and nonstoring birds, black-capped chickadees (Poecile atricapilla) and dark-eyed juncos (Junco hyemalis)			Type	Journal Article
	Year	2002	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	28	Issue	3	Pages	227-241
	Keywords	Animals; Attention/physiology; Behavior, Animal/physiology; Birds; Discrimination (Psychology); Food Habits; Memory/physiology; Space Perception/physiology; Spatial Behavior/physiology
	Abstract	Food-storing birds, black-capped chickadees (Poecile atricapilla), and nonstoring birds, dark-eyed juncos (Junco hyemalis), matched color or location on a touch screen. Both species showed a divided attention effect for color but not for location (Experiment 1). Chickadees performed better on location than on color with retention intervals up to 40 s, but juncos did not (Experiment 2). Increasing sample-distractor distance improved performance similarly in both species. Multidimensional scaling revealed that both use a Euclidean metric of spatial similarity (Experiment 3). When choosing between the location and color of a remembered item, food storers choose location more than do nonstorers. These results explain this effect by differences in memory for location relative to color, not division of attention or spatial discrimination ability.
	Address	Department of Psychology, University of Toronto, 100 Saint George Street, Toronto, Ontario M5S 3G3, Canada. shettle@psych.utoronto.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12136700			Approved	no
	Call Number	refbase @ user @			Serial	370
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	Author	Skov-Rackette, S.I.; Miller, N.Y.; Shettleworth, S.J.
	Title	What-where-when memory in pigeons			Type	Journal Article
	Year	2006	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	32	Issue	4	Pages	345-358
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Conditioning, Operant/physiology; Memory/physiology; Reinforcement (Psychology); Space Perception/physiology; Spatial Behavior/physiology; Teaching; Visual Perception/physiology
	Abstract	The authors report a novel approach to testing episodic-like memory for single events. Pigeons were trained in separate sessions to match the identity of a sample on a touch screen, to match its location, and to report on the length of the retention interval. When these 3 tasks were mixed randomly within sessions, birds were more than 80% correct on each task. However, performance on 2 different tests in succession after each sample was not consistent with an integrated memory for sample location, time, and identity. Experiment 2 tested binding of location and identity memories in 2 different ways. The results were again consistent with independent feature memories. Implications for tests of episodic-like memory are discussed.
	Address	Department of Psychology, University of Toronto, Toronto, ON, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17044738			Approved	no
	Call Number	refbase @ user @			Serial	357
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	Author	Sovrano, V.A.; Bisazza, A.; Vallortigara, G.
	Title	How fish do geometry in large and in small spaces			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	1	Pages	47-54
	Keywords	Animals; Association Learning; Color Perception; Cues; Discrimination Learning; Distance Perception; Fishes; Male; Pattern Recognition, Visual; Social Environment; *Space Perception; Visual Perception
	Abstract	It has been shown that children and non-human animals seem to integrate geometric and featural information to different extents in order to reorient themselves in environments of different spatial scales. We trained fish (redtail splitfins, Xenotoca eiseni) to reorient to find a corner in a rectangular tank with a distinctive featural cue (a blue wall). Then we tested fish after displacement of the feature on another adjacent wall. In the large enclosure, fish chose the two corners with the feature, and also tended to choose among them the one that maintained the correct arrangement of the featural cue with respect to geometric sense (i.e. left-right position). In contrast, in the small enclosure, fish chose both the two corners with the features and the corner, without any feature, that maintained the correct metric arrangement of the walls with respect to geometric sense. Possible reasons for species differences in the use of geometric and non-geometric information are discussed.
	Address	Department of General Psychology, University of Padua, Via Venezia, 8, 35131, Padova, Italy. valeriaanna.sovrano@unipd.it
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16794851			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2462
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	Author	Sturz, B.R.; Bodily, K.D.; Katz, J.S.
	Title	Evidence against integration of spatial maps in humans			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	3	Pages	207-217
	Keywords	Adult; Association Learning; Computer Graphics; Humans; Male; Orientation; Problem Solving; Reference Values; Space Perception; *Spatial Behavior; User-Computer Interface
	Abstract	A dynamic 3-D virtual environment was constructed for humans as an open-field analogue of Blaisdell and Cook's (2005) pigeon foraging task to determine if humans, like pigeons, were capable of integrating separate spatial maps. Participants used keyboard keys and a mouse to search for a hidden goal in a 4x4 grid of raised cups. During Phase 1 training, a goal was consistently located between two landmarks (Map 1: blue T and red L). During Phase 2 training, a goal was consistently located down and left of a single landmark (Map 2: blue T). Transfer trials were then conducted in which participants were required to make choices in the presence of the red L alone. Cup choices during transfer assessed participants' strategies: association (from Map 1), generalization (from Map 2), or integration (combining Map 1 and 2). During transfer, cup choices increased to a location which suggested an integration strategy and was consistent with results obtained with pigeons. However, additional analyses of the human data suggested participants initially used a generalization strategy followed by a progressive shift in search behavior away from the red L. This shift in search behavior during transfer was responsible for the changes in cup choices across transfer trials and was confirmed by a control condition. These new analyses offer an alternative explanation to the spatial integration account proposed for pigeons.
	Address	Department of Psychology, Auburn University, Auburn, AL 36849, USA. sturzbr@auburn.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16767470			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2464
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	Author	Sutton, J.E.; Shettleworth, S.J.
	Title	Internal sense of direction and landmark use in pigeons (Columba livia)			Type	Journal Article
	Year	2005	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	119	Issue	3	Pages	273-284
	Keywords	Animals; Columbidae; Conflict (Psychology); Cues; Discrimination Learning; Homing Behavior; Intuition; Orientation; Space Perception; Transfer (Psychology); Visual Perception
	Abstract	The relative importance of an internal sense of direction based on inertial cues and landmark piloting for small-scale navigation by White King pigeons (Columba livia) was investigated in an arena search task. Two groups of pigeons differed in whether they had access to visual cues outside the arena. In Experiment 1, pigeons were given experience with 2 different entrances and all pigeons transferred accurate searching to novel entrances. Explicit disorientation before entering did not affect accuracy. In Experiments 2-4, landmarks and inertial cues were put in conflict or tested 1 at a time. Pigeons tended to follow the landmarks in a conflict situation but could use an internal sense of direction to search when landmarks were unavailable.
	Address	Department of Psychology, University of Toronto, ON, Canada. jsutton7@uwo.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16131256			Approved	no
	Call Number	refbase @ user @			Serial	360
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