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	Author	Schooening, B.
	Title	Ethology of the horse			Type	Journal Article
	Year	1998	Publication	Praktische Tierarzt	Abbreviated Journal	Prakt. Tierarzt
	Volume	79	Issue	6 Suppl.	Pages	25-28
	Keywords	Dominance; Ethology; Horse; Propedeutics
	Abstract	The paper starts with a short introduction/definition about ethology and the used methods in this scientific field, giving special examples for horses and about how their “normal behaviour” is measured. The behaviour repertoire of horses is described in a brief outline with special emphasis on their social systems and hierarchies and the problem of dominance, especially in interaction with humans. Schlütersche GmbH & Co. KG, Verlag und Druckerei.
	Address	Saselbergweg 32, 22395 Hamburg, Germany
	Corporate Author				Thesis
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	ISSN	0032681x (Issn)	ISBN		Medium
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	Notes	Cited By (since 1996): 1; Export Date: 21 April 2007; Source: Scopus; Language of Original Document: German; Correspondence Address: Schoßning, B.Saselbergweg 32 22395 Hamburg, Germany; email: Dr.B.Schoening@t-online.de			Approved	no
	Call Number	refbase @ user @			Serial	789
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	Author	Araba, B.D.; Crowell-Davis, S.L.
	Title	Dominance relationships and aggression of foals (Equus caballus)			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	41	Issue	1-2	Pages	1-25
	Keywords	aggression; dominance; horse; Equus caballus
	Abstract	Studied a herd of 15 Belgian brood-mares and 10 foals. Specific aspects of social structure studied were dominance-subordinance relationships, preferred associates, social spacing, aggression rates, the frequency of aggressions administered down the dominance hierarchy, and interactive play bouts. The rank order of the foals, both before and after weaning, was positively correlated with the rank order of their dams. There was also a significant relationship between a foal's rank and its total aggression or aggression rate per subordinate post-weaning. Higher ranking foals had higher rates of aggression. Over 80% of threats were directed down the dominance hierachy. The play-rank order of the foals, scored by the number of times foal left a play bout, was not significantly correlated with the rank order as scored by agonistic interactions. -from Authors
	Address	Dept Anatomy and Radiology, Univ of Georgia, Athens, GA, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	01681591 (Issn)	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	790
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	Author	Lehmann, K.; Kallweit, E.; Ellendorff, F.
	Title	Social hierarchy in exercised and untrained group-housed horses--A brief report			Type	Journal Article
	Year	2006	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	96	Issue	3-4	Pages	343-347
	Keywords	Horses; Social hierarchy; Exercise; Group-housing
	Abstract	Changes in social hierarchy were evaluated in a herd of 3-year-old Hanoverian geldings. One group (n = 5) was exposed to a training programme, the other (n = 5) remained untrained. After 6 months, the groups were reversed. Hierarchical positions were evaluated by field observations and/or paired-feeding tests at the beginning, the middle, the end of the first and at the end of the second training period. Both methods yielded identical results. Almost all horses changed position in only one direction: either up or down. Neither increase in aggression nor mutual injuries were recorded during the whole experiment. No statistically verified differences in dominance ranking occurred between trained and untrained groups, but apparent differences were consistent. Thus, if horses are kept in the same group for a longer period of time, exercise induced changes in hierarchy are probably of minor importance and are unlikely to increase the incidence of injuries. This may have implications for the promotion of group-housing for sport horses.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	800
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	Author	Sappington, B.K.F.; McCall, C.A.; Coleman, D.A.; Kuhlers, D.L.; Lishak, R.S.
	Title	A preliminary study of the relationship between discrimination reversal learning and performance tasks in yearling and 2-year-old horses			Type	Journal Article
	Year	1997	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	53	Issue	3	Pages	157-166
	Keywords	Cognition; Learning; Horse; Training
	Abstract	A study was conducted to determine the relationship between discrimination reversal learning and performance tasks in horses. Ten yearling and seven 2-year-old mares and geldings of Arabian (n = 4), Quarter Horse (n = 9), and Thoroughbred (n = 4) breeding were given a two-choice discrimination task in which either a black or a white bucket contained a food reward for ten trials per day during 19 test days. The spatial position of the buckets was varied on a random schedule. The rewarded bucket color was reversed each time a subject met criterion of eight correct choices per day for 2 consecutive days. Discrimination reversal testing was followed by 6 days of performance tasks: three crossing a wooden bridge and three jumping an obstacle to reach food and conspecifics, within a maximum allotted time of 15 min day-1. Total reversals attained by the horses were low (x = 1.5 +/- 0.9). All subjects did attain at least one reversal, and six had two or more reversals. No differences (P > .05) were detected between ages or sexes, nor among breeds in discrimination reversal learning or performance test measurements. However, there was a trend towards a breed difference (P <= 0.09) in the mean number of correct responses to the first reversal criterion. Correlations between reversal learning results and performance task results were extremely low, indicating that the discrimination reversal learning test was not useful for predicting success at these performance tasks. Results from the two performance tasks also showed little correlation (r = 0.04, P < 0.91), indicating that horses might not use the same approach when solving the problem of crossing these two obstacles. The overall poor performance of the horses on the discrimination reversal task suggests horses may have difficulty reversing previously learned tasks.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	826
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	Author	Hall, C.A.; Cassaday, H.J.
	Title	An investigation into the effect of floor colour on the behaviour of the horse			Type	Journal Article
	Year	2006	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	99	Issue	3-4	Pages	301-314
	Keywords	Horse; Colour aversion; Flooring; Vision
	Abstract	Adverse reactions of the domestic horse to environmental stimuli can be problematic in training and management. Hesitation and alarm reactions to visual features of the ground can occur in both ridden work and when handling horses. To assess the effect of one visual feature (colour) on the behaviour of the domestic horse, the reactions of 16 riding horses to 8 different coloured mats were recorded. The effect of stimulus position on these reactions was assessed by presenting them in two different positions, either on the ground (where the horses had to walk over them) or against a wall (where the horses walked past them). Each colour/position combination was presented twice in order to assess the effect of previous experience. An alleyway was constructed to allow the horses to be tested unconstrained and freely walking throughout. The time taken to traverse the alleyway and the observed reaction to the colour was recorded. Significant differences in both measures were found in relation to the position of the colour and whether the subject had previous experience of that colour/position combination. The initial presentation of the colours on the ground produced the highest percentage of adverse reactions. Certain colours encountered for the first time on the ground (yellow, white, black and blue) were found to cause a greater number of adverse reactions than others (green, red, brown and grey) and an increase in time taken to traverse the alleyway. Although a significant difference in relation to colour was found in the behaviour observed during the second presentation on the ground, no difference was found in relation to the time taken to traverse the alleyway. No significant effect of colour was found when the coloured stimuli were presented against the wall. These findings have important implications for situations where the colour of flooring could be controlled in order to minimise adverse behavioural reactions, in particular during initial training.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	834
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	Author	Hall, C.A.; Cassaday, H.J.; Derrington, A.M.
	Title	The effect of stimulus height on visual discrimination in horses			Type	Journal Article
	Year	2003	Publication	Journal of Animal Science	Abbreviated Journal	J. Anim Sci.
	Volume	81	Issue	7	Pages	1715-1720
	Keywords	Animals; Discrimination Learning/physiology; Female; Horses/physiology/psychology; Male; Orientation; Photic Stimulation; Vision/physiology
	Abstract	This study investigated the effect of stimulus height on the ability of horses to learn a simple visual discrimination task. Eight horses were trained to perform a two-choice, black/white discrimination with stimuli presented at one of two heights: ground level or at a height of 70 cm from the ground. The height at which the stimuli were presented was alternated from one session to the next. All trials within a single session were presented at the same height. The criterion for learning was four consecutive sessions of 70% correct responses. Performance was found to be better when stimuli were presented at ground level with respect to the number of trials taken to reach the criterion (P < 0.05), percentage of correct first choices (P < 0.01), and repeated errors made (P < 0.01). Thus, training horses to carry out tasks of visual discrimination could be enhanced by placing the stimuli on the ground. In addition, the results of the present study suggest that the visual appearance of ground surfaces is an important factor in both horse management and training.
	Address	School of Land-based Studies, Nottingham Trent University, Brackenhurst College Campus, Southwell, Nottinghamshire, England NG25 0QF. carol.hall@ntu.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0021-8812	ISBN		Medium
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	Notes	PMID:12854807			Approved	no
	Call Number	refbase @ user @			Serial	835
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	Author	Harman, A.M.; Moore, S.; Hoskins, R.; Keller, P.
	Title	Horse vision and an explanation for the visual behaviour originally explained by the 'ramp retina'			Type	Journal Article
	Year	1999	Publication	Equine veterinary journal	Abbreviated Journal	Equine Vet J
	Volume	31	Issue	5	Pages	384-390
	Keywords	Animals; Behavior, Animal; Cell Count; Eye/anatomy & histology; Ganglia, Sensory/cytology; Horses/physiology; Refractive Errors/veterinary; Retina/cytology/physiology; Vision/physiology; Visual Acuity; Visual Fields
	Abstract	Here we provide confirmation that the 'ramp retina' of the horse, once thought to result in head rotating visual behaviour, does not exist. We found a 9% variation in axial length of the eye between the streak region and the dorsal periphery. However, the difference was in the opposite direction to that proposed for the 'ramp retina'. Furthermore, acuity in the narrow, intense visual streak in the inferior retina is 16.5 cycles per degree compared with 2.7 cycles per degree in the periphery. Therefore, it is improbable that the horse rotates its head to focus onto the peripheral retina. Rather, the horse rotates the nose up high to observe distant objects because binocular overlap is oriented down the nose, with a blind area directly in front of the forehead.
	Address	Department of Psychology, University of Western Australia, Nedlands, Australia
	Corporate Author				Thesis
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	Language	English	Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN	0425-1644	ISBN		Medium
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	Notes	PMID:10505953			Approved	no
	Call Number	refbase @ user @			Serial	836
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	Author	Lansade, L.; Bertrand, M.; Bouissou, M.-F.
	Title	Effects of neonatal handling on subsequent manageability, reactivity and learning ability of foals			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	92	Issue	1-2	Pages	143-158
	Keywords	Horse; Neonatal handling; Manageability; Emotional reactivity; Learning-ability; Human-animal relationship
	Abstract	Behaviour is an important factor to be taken into account in the various uses of horses. Today horses are mainly used for sport and leisure activities. They should therefore be easy to manage, calm and not fearful. Early handling is known to improve manageability and learning ability and to reduce fearfulness in various species. It has become fashionable in the horse industry to use an early training procedure, referred to as “imprint training”, which is said to produce durable if not permanent effects. However, no studies concerning the long-term effects of such neonatal handling have been carried out in horses. The present study examines the short- and long-term effects of neonatal handling on manageability, general reactivity and learning ability of foals. Twenty-six Welsh foals were studied: 13 were handled daily for 14 days from birth and 13 were non-handled controls. The handling procedure consisted of fitting a halter, gently patting all parts of each foal's body, picking up feet and leading over 40 m. Two days, 3 months, 6 months and 1 year after the end of the handling period, foals underwent behavioural tests to measure their manageability and various aspects of their reactivity. The results showed that neonatal handling has only short-term effects on manageability: 2 days after the handling period, handled animals were significantly easier to handle than controls for the four parameters measured during this test (time to fit a halter, time to pick up feet, walk ratio that is time during which foal walks under constraint/total time measured during leading and number of defensive reactions). Two parameters (time to fit a halter and walk ratio) were still lower in handled foals than in non-handled foals 3 months later and only one 6 months later (walk ratio). One year later there was no difference between groups. In addition, there was no effect of handling on reactivity at any time of testing or in any of the tests (reaction to isolation from conspecifics, presence of a human, presence of a novel object and to a surprise effect). Finally, neonatal handling did not improve the spatial or discriminative learning abilities measured at 14 months of age. To conclude, the effects of neonatal handling are only temporary.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	841
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	Author	Lansade, L.; Bertrand, M.; Boivin, X.; Bouissou, M.-F.
	Title	Effects of handling at weaning on manageability and reactivity of foals			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	87	Issue	1-2	Pages	131-149
	Keywords	Horse; Handling; Weaning; Manageability; Emotional reactivity; Fearfulness; Human-animal relationship
	Abstract	The horse's temperament, including its manageability and reactivity and/or fearfulness, is of importance as it can result in problems and can render horses unsuitable for inexperienced riders. Early experience, including handling during infancy, may influence the horse's adult behaviour and reduce its fear of humans and other potentially frigthening situations. In the various species studied, handling has generally been undertaken during the neonatal period. The aim of the present study was to test the effects of handling young horses around the time of weaning, a period which has been demonstrated to be effective in increasing ease of handling in cattle and goats. Sixteen Anglo-Arab foals were handled for 12 days either immediately following weaning (early handled: EH) or 21 days later (late handled: LH); eight additional non-handled foals served as controls (C). Handling consisted of haltering, gently petting all parts of the body, picking up feet and leading the foal over 120 m. During handling sessions, EH were easier to handle than LH: time taken to fit them with a halter, to pick up feet, and “walk-ratio” (time walking under constraint/total time walking) were significantly lower for EH. During subsequent tests conducted over 2 days, 4, and 7 months, as well as 10 months and to some extent 18 months after the end of handling period, EH and LH were easier to handle and less reactive than controls, although differences diminished with time. The period following weaning can therefore be qualified as an “optimal period” for handling. Some of the effects persist for at least 18 months.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	842
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	Author	Macuda, T.; Timney, B.
	Title	Luminance and chromatic discrimination in the horse (Equus caballus)			Type	Journal Article
	Year	1999	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	44	Issue	3	Pages	301-307
	Keywords	Colour vision; Chromatic discrimination; Luminance discrimination; Horse
	Abstract	Equine colour vision was measured under conditions that minimised the possibility of animals using brightness cues to make chromatic discriminations. In a two-stage study, we first obtained luminance discrimination functions for achromatic targets then tested for chromatic discrimination over a range of target luminances. Horses were trained on a two-choice discrimination task. The positive stimulus was varied in luminance and/or colour using neutral density and broad band colour filters. The negative stimulus appeared as a uniform grey. In the brightness discrimination task, the horses performed well at large luminance differences but their percentage of correct responses declined to near chance levels at differences of less than 0.2 log units. In addition, a decrement in performance was noted at luminance differences of less than 0.2 log units for green and yellow chromatic discrimination functions, suggesting that horses cannot easily discriminate yellow and green from grey. However, the chromatic discrimination functions for red and blue showed that animals performed very well across the full range of target luminances. These results suggest that horses are at least dichromats.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	844
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