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	Author	Reid, P.J.; Shettleworth, S.J.
	Title	Detection of cryptic prey: search image or search rate?			Type	Journal Article
	Year	1992	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	18	Issue	3	Pages	273-286
	Keywords	Animals; Appetitive Behavior; Attention; Color Perception; Columbidae; Discrimination Learning; Food Preferences/psychology; Imagination; Mental Recall; *Predatory Behavior
	Abstract	Animals' improvement in capturing cryptic prey with experience has long been attributed to a perceptual mechanism, the specific search image. Detection could also be improved by adjusting rate of search. In a series of studies using both naturalistic and operant search tasks, pigeons searched for wheat, dyed to produce 1 conspicuous and 2 equally cryptic prey types. Contrary to the predictions of the search-rate hypothesis, pigeons given a choice between the 2 cryptic types took the type experienced most recently. However, experience with 1 cryptic type improved accuracy on the other cryptic type, a result inconsistent with a search image specific to 1 prey type. Search image may better be thought of as priming of attention to those features of the prey type that best distinguish the prey from the background.
	Address	University of Toronto, Ontario, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:1619395			Approved	no
	Call Number	refbase @ user @			Serial	381
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	Author	Shettleworth, S.J.; Plowright, C.M.
	Title	How pigeons estimate rates of prey encounter			Type	Journal Article
	Year	1992	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	18	Issue	3	Pages	219-235
	Keywords	Analysis of Variance; Animals; Appetitive Behavior; Columbidae; Conditioning, Operant; Food Preferences/psychology; Motivation; Predatory Behavior; Probability Learning; *Reinforcement Schedule; Social Environment
	Abstract	Pigeons were trained on operant schedules simulating successive encounters with prey items. When items were encountered on variable-interval schedules, birds were more likely to accept a poor item (long delay to food) the longer they had just searched, as if they were averaging prey density over a short memory window (Experiment 1). Responding as if the immediate future would be like the immediate past was reversed when a short search predicted a long search next time (Experiment 2). Experience with different degrees of environmental predictability appeared to change the length of the memory window (Experiment 3). The results may reflect linear waiting (Higa, Wynne, & Staddon, 1991), but they differ in some respects. The findings have implications for possible mechanisms of adjusting behavior to current reinforcement conditions.
	Address	Department of Psychology, University of Toronto, Ontario, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:1619391			Approved	no
	Call Number	refbase @ user @			Serial	382
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	Author	Biro, D.; Sumpter, D.J.T.; Meade, J.; Guilford, T.
	Title	From Compromise to Leadership in Pigeon Homing			Type	Journal Article
	Year	2006	Publication	Current Biology	Abbreviated Journal	Curr Biol
	Volume	16	Issue	21	Pages	2123-2128
	Keywords	Animal Migration; Animals; Columbidae/physiology; Decision Making; Flight, Animal; Homing Behavior; Models, Biological; Orientation; Social Behavior; *Social Dominance
	Abstract	Summary A central problem faced by animals traveling in groups is how navigational decisions by group members are integrated, especially when members cannot assess which individuals are best informed or have conflicting information or interests , , , and . Pigeons are now known to recapitulate faithfully their individually distinct habitual routes home , and , and this provides a novel paradigm for investigating collective decisions during flight under varying levels of interindividual conflict. Using high-precision GPS tracking of pairs of pigeons, we found that if conflict between two birds' directional preferences was small, individuals averaged their routes, whereas if conflict rose over a critical threshold, either the pair split or one of the birds became the leader. Modeling such paired decision-making showed that both outcomes--compromise and leadership--could emerge from the same set of simple behavioral rules. Pairs also navigated more efficiently than did the individuals of which they were composed, even though leadership was not necessarily assumed by the more efficient bird. In the context of mass migration of birds and other animals, our results imply that simple self-organizing rules can produce behaviors that improve accuracy in decision-making and thus benefit individuals traveling in groups , and .
	Address	Department of Zoology, University of Oxford, Oxford OX1 3PS, United Kingdom. daro.biro@zoo.ox.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	Englisch	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0960-9822	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17084696			Approved	no
	Call Number				Serial	2026
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	Author	Nguyen, N.H.; Klein, E.D.; Zentall, T.R.
	Title	Imitation of a two-action sequence by pigeons			Type	Journal Article
	Year	2005	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	12	Issue	3	Pages	514-518
	Keywords	Animals; Behavior, Animal; Cognition; Columbidae; Imitative Behavior; *Learning
	Abstract	Developmental psychologists have described imitation as a process that suggests perspective-taking abilities. However, imitative behavior has been found in animals, which are generally not considered capable of taking the perspective of another. Previous studies with birds have demonstrated the imitation of a single response (sometimes referred to as action-level imitation). In the present experiment, we examined the extent to which pigeons would imitate an unfamiliar sequence of two behaviors (sometimes referred to as program-level imitation). Our results indicate that, although there are individual differences, pigeons show a significant tendency to match a demonstrated sequence of behavior involving, first, a response to a treadle (pecking at it or stepping on it) and, second, pushing aside a screen that blocks access to food (a left-vs.-right push).
	Address	University of Kentucky, Lexington, Kentucky 40506, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16235638			Approved	no
	Call Number	refbase @ user @			Serial	221
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	Author	Zentall, T.R.; Jackson-Smith, P.; Jagielo, J.A.; Nallan, G.B.
	Title	Categorical shape and color coding by pigeons			Type	Journal Article
	Year	1986	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	12	Issue	2	Pages	153-159
	Keywords	Animals; Color Perception; Columbidae; Discrimination Learning; Form Perception; Generalization, Stimulus; Psychophysics; Transfer (Psychology)
	Abstract	Categorical coding is the tendency to respond similarly to discriminated stimuli. Past research indicates that pigeons can categorize colors according to at least three spectral regions. Two present experiments assessed the categorical coding of shapes and the existence of a higher order color category (all colors). Pigeons were trained on two independent tasks (matching-to-sample, and oddity-from-sample). One task involved red and a plus sign, the other a circle and green. On test trials one of the two comparison stimuli from one task was replaced by one of the stimuli from the other task. Differential performance based on which of the two stimuli from the other task was introduced suggested categorical coding rules. In Experiment 1 evidence for the categorical coding of sample shapes was found. Categorical color coding was also found; however, it was the comparison stimuli rather than the samples that were categorically coded. Experiment 2 replicated the categorical shape sample effect and ruled out the possibility that the particular colors used were responsible for the categorical coding of comparison stimuli. Overall, the results indicate that pigeons can develop categorical rules involving shapes and colors and that the color categories can be hierarchical.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3701264			Approved	no
	Call Number	refbase @ user @			Serial	262
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	Author	Urcuioli, P.J.; Zentall, T.R.
	Title	Retrospective coding in pigeons' delayed matching-to-sample			Type	Journal Article
	Year	1986	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	12	Issue	1	Pages	69-77
	Keywords	Animals; Color Perception; Columbidae; Discrimination Learning; Form Perception; Memory; Mental Recall; Orientation; Pattern Recognition, Visual; Retention (Psychology)
	Abstract	In this study we examined how coding processes in pigeons' delayed matching-to-sample were affected by the stimuli to be remembered. In Experiment 1, two groups of pigeons initially learned 0-delay matching-to-sample with identical comparison stimuli (vertical and horizontal lines) but with different sample stimuli (red and green hues or vertical and horizontal lines). Longer delays were then introduced between sample offset and comparison onset to assess whether pigeons were prospectively coding the same events (viz., the correct line comparisons) or retrospectively coding different events (viz., their respective sample stimuli). The hue-sample group matched more accurately and showed a slower rate of forgetting than the line-sample group. In Experiment 2, pigeons were trained with either hues or lines as both sample and comparison stimuli, or with hue samples and line comparisons or vice versa. Subsequent delay tests revealed that the hue-sample groups remembered more accurately and generally showed slower rates of forgetting than the line-sample groups. Comparison dimension had little or no effect on performance. Together, these data suggest that pigeons retrospectively code the samples in delayed matching-to-sample.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3701260			Approved	no
	Call Number	refbase @ user @			Serial	263
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	Author	DiGian, K.A.; Friedrich, A.M.; Zentall, T.R.
	Title	Discriminative stimuli that follow a delay have added value for pigeons			Type	Journal Article
	Year	2004	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	11	Issue	5	Pages	889-895
	Keywords	Animals; Behavior, Animal; Choice Behavior; Columbidae; Cues; Discrimination (Psychology)
	Abstract	Clement, Feltus, Kaiser, and Zentall (2000) reported that pigeons prefer discriminative stimuli that require greater effort (more pecks) to obtain over those that require less effort. In the present experiment, we examined two variables associated with this phenomenon. First, we asked whether delay of reinforcement, presumably a relatively aversive event similar to effort, would produce similar effects. Second, we asked whether the stimulus preference produced by a prior relatively aversive event depends on its anticipation. Anticipation of delay was accomplished by signaling its occurrence. Results indicated that delays can produce preferences similar to those produced by increased effort, but only if the delays are signaled.
	Address	University of Kentucky, Lexington, Kentucky 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15732699			Approved	no
	Call Number	refbase @ user @			Serial	226
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	Author	Zentall, T.R.; Weaver, J.E.; Clement, T.S.
	Title	Pigeons group time intervals according to their relative duration			Type	Journal Article
	Year	2004	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	11	Issue	1	Pages	113-117
	Keywords	Animals; Columbidae; *Discrimination (Psychology); Reinforcement (Psychology); Time Factors; Time Perception
	Abstract	In the present research, we asked whether pigeons tended to judge time intervals not only in terms of their absolute value but also relative to a duration from which they must be discriminated (i.e., longer or shorter). Pigeons were trained on two independent temporal discriminations. In one discrimination, sample durations of 2 and 8 sec were associated with, for example, red and green hue comparisons, respectively, and in the other discrimination, sample durations of 4 and 16 sec were associated with vertical and horizontal line comparisons, respectively. If pigeons are trained on a temporal discrimination and tested with intermediate durations, the subjective midpoint typically occurs close to the geometric mean of the two trained values. The 4- and 8-sec values were selected to be the geometric mean of the two values in the other discrimination. When a 4-sec test sample was presented with the comparisons from the 2- and 8-sec discrimination, the pigeons preferred the comparison associated with the shorter sample. Similarly, when an 8-sec test sample was presented with the comparisons from the 4- and 16-sec discrimination, the pigeons preferred the comparison associated with the longer sample. Thus, a relative grouping effect was found. That is, durations that should have produced indifferent choice were influenced by their relative durations (shorter than or longer than the alternative) during training.
	Address	Department of Psychology, University of Kentucky, Lexington, Kentucky 40506-0044, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15116995			Approved	no
	Call Number	refbase @ user @			Serial	231
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	Author	Clement, T.S.; Zentall, T.R.
	Title	Development of a single-code/default coding strategy in pigeons			Type	Journal Article
	Year	2000	Publication	Psychological science : a journal of the American Psychological Society / APS	Abbreviated Journal	Psychol Sci
	Volume	11	Issue	3	Pages	261-264
	Keywords	Animals; Attention; Columbidae; Discrimination Learning; Female; Male; Pattern Recognition, Visual; *Problem Solving; Retention (Psychology)
	Abstract	We tested the hypothesis that pigeons could use a cognitively efficient coding strategy by training them on a conditional discrimination (delayed symbolic matching) in which one alternative was correct following the presentation of one sample (one-to-one), whereas the other alternative was correct following the presentation of any one of four other samples (many-to-one). When retention intervals of different durations were inserted between the offset of the sample and the onset of the choice stimuli, divergent retention functions were found. With increasing retention interval, matching accuracy on trials involving any of the many-to-one samples was increasingly better than matching accuracy on trials involving the one-to-one sample. Furthermore, following this test, pigeons treated a novel sample as if it had been one of the many-to-one samples. The data suggest that rather than learning each of the five sample-comparison associations independently, the pigeons developed a cognitively efficient single-code/default coding strategy.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0956-7976	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11273414			Approved	no
	Call Number	refbase @ user @			Serial	246
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	Author	Clement, T.S.; Zentall, T.R.
	Title	Choice based on exclusion in pigeons			Type	Journal Article
	Year	2003	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	10	Issue	4	Pages	959-964
	Keywords	Animals; Appetitive Behavior; Association Learning; Choice Behavior; Color Perception; Columbidae; Discrimination Learning; Memory, Short-Term; *Problem Solving; Psychomotor Performance; Reaction Time; Transfer (Psychology)
	Abstract	When humans acquire a conditional discrimination and are given a novel-sample-comparison choice, they often reject a comparison known to be associated with a different sample and choose the alternative comparison by default (or by exclusion). In Experiment 1, we found that if, following matching training, we replaced both of the samples, acquisition took five times longer than if we replaced only one of the samples. Apparently, the opportunity to reject one of the comparisons facilitated the association of the other sample with the remaining comparison. In Experiment 2, we first trained pigeons to treat two samples differently (to associate Sample A with Comparison 1 and Sample B with Comparison 2) and then trained them to associate one of those samples with a new comparison (e.g., Sample A with Comparison 3) and to associate a novel sample (Sample C) with a different, new comparison (Comparison 4). When Sample B then replaced Sample C, the pigeons showed a significant tendency to choose Comparison 4 over Comparison 3. Thus, when given the opportunity, pigeons will choose by exclusion.
	Address	University of Kentucky, Lexington, Kentucky, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15000545			Approved	no
	Call Number	refbase @ user @			Serial	233
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