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	Author	Farmer, K.; Krüger, K.; Byrne, R.W.; Marr, I.
	Title	Sensory laterality in affiliative interactions in domestic horses and ponies (Equus caballus)			Type	Journal Article
	Year	2018	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	21	Issue	5	Pages	631-637
	Keywords
	Abstract	Many studies have been carried out into both motor and sensory laterality of horses in agonistic and stressful situations. Here we examine sensory laterality in affiliative interactions within four groups of domestic horses and ponies (N = 31), living in stable social groups, housed at a single complex close to Vienna, Austria, and demonstrate for the first time a significant population preference for the left side in affiliative approaches and interactions. No effects were observed for gender, rank, sociability, phenotype, group, or age. Our results suggest that right hemisphere specialization in horses is not limited to the processing of stressful or agonistic situations, but rather appears to be the norm for processing in all social interactions, as has been demonstrated in other species including chicks and a range of vertebrates. In domestic horses, hemispheric specialization for sensory input appears not to be based on a designation of positive versus negative, but more on the perceived need to respond quickly and appropriately in any given situation.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Farmer2018			Serial	6386
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	Author	Meriggi,A.; Lovari, S.
	Title	A Review of Wolf Predation in Southern Europe: Does the Wolf Prefer Wild Prey to Livestock?			Type	Journal Article
	Year	1996	Publication	Journal of Applled Ecology	Abbreviated Journal	J. Appl. Ecol
	Volume	33	Issue		Pages	1561-1571
	Keywords	Canis lupus, conservation, food habits, prey abundance, prey availability.
	Abstract	1. The recent recovery of the wolf in southern Europe has not yet removed the risk of local extinction. Wolf populations are fragmented and often comprise fewer than 500 individuals. In North America, northern and eastern Europe, wolves feed maiiily on wild herbivores. In southern Europe, this canid has apparently adapted to feed also on fruit, rubbish, livestock, small and medium-size mammals. 2. The main conservation problem lies with predation o n domestic ~ingulates,w liich leads to extensive killing of wolves. The reintroduction of wild large herbivores has been advocated as a means of reducing attacks on livestock, but predatiori on the latter may remain high if domestic ungulates are locally abundant. 3. Our synthesis of 15 studies, published in the last 15 years, on food habits of the wolf in southern Europe, has shown that ungulates have been the main diet component overall. A significant inverse correlation was found between the occurrence (%) of wild and domestic ungulates in the diet. The presence of relatively few wild ungulate species was necessary to reduce predation on livestock. 4. Selection of wild and domestic ungulate prey was influenced mainly by their local abundance, but also by their accessibility. Feeding dependence on rubbish was local and rare. In Italy, the consumption of riibbish/fruit and that of ungulates was significantly negatively correlated. Diet breadth increased as the presence of large prey in tlie diet decreased. 5. The simultaneous reintroduction of severa1 wild ungulate species is likely to reduce predation on livestock and may prove to be one of the most effective conservation measures.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6387
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	Author	Zlatanova, D.; Ahmed, A.; Valasseva, A.; Genov, P.
	Title	Adaptive Diet Strategy of the Wolf (Canis lupus L.) in Europe: a Review			Type	Journal Article
	Year	2014	Publication	ACTA ZOOLOGICA BULGARICA	Abbreviated Journal	Acta zool. bulg.
	Volume	66	Issue	4	Pages	439-452
	Keywords	Wolf, Canis lupus, prey, adaptive strategy
	Abstract	The diet strategy of the wolf in Europe is reviewed on the basis of 74 basic and 14 additional literature sources. The comparative analysis reveals clear dependence on the latitude (and, therefore, on the changing environmental conditions) correlated with the wild ungulate abundance and diversity. Following a geographic pattern, the wolf is specialised on different species of ungulates: moose and reindeer in Scandinavia, red deer in Central and Eastern Europe and wild boar in Southern Europe. Where this large prey is taken, the roe deer is hunted with almost the same frequency in every region. The wolf diet in Europe shows two ecological adaptations formed by a complex of variables: 1. Wolves living in natural habitats with abundance of wild ungulates feed mainly on wild prey. 2. In highly anthropogenic habitats, with low abundance of wild prey, wolves feed on livestock (where husbandry of domestic animals is available) and take also a lot of plant food, smaller prey (hares and rodents) and garbage food. The frequency of occurrence of wild ungulates in the diet of wolves in North Europe varies from 54.0% in Belarus to 132.7% in Poland, while that of livestock is in the range from 0.4% in Norway to 74.9% in Belarus. In South Europe, the frequency of occurrence of wild prey varies from 0% in Italy and Spain to 136.0% in Italy, while of domestic ungulates ranges between 0% and 100% in Spain. The low density or lack of wild prey triggers the switch of the wolf diet to livestock, plant food (32.2-85% in Italy) or even garbage (up to 41.5% in Italy).
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6388
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	Author	Nakamura, K.; Takimoto-Inose, A.; Hasegawa, T.
	Title	Cross-modal perception of human emotion in domestic horses (Equus caballus)			Type	Journal Article
	Year	2018	Publication	Scientific Reports	Abbreviated Journal
	Volume	8	Issue	1	Pages	8660
	Keywords
	Abstract	Humans have domesticated many kinds of animals in their history. Dogs and horses have particularly close relationships with humans as cooperative partners. However, fewer scientific studies have been conducted on cognition in horses compared to dogs. Studies have shown that horses cross-modally distinguish human facial expressions and recognize familiar people, which suggests that they also cross-modally distinguish human emotions. In the present study, we used the expectancy violation method to investigate whether horses cross-modally perceive human emotions. Horses were shown a picture of a human facial expression on a screen, and they then heard a human voice from the speaker before the screen. The emotional values of the visual and auditory stimuli were the same in the congruent condition and different in the incongruent condition. Horses looked at the speaker significantly longer in the incongruent condition than in the congruent condition when they heard their caretaker's voices but not when they heard the stranger voice. In addition, they responded significantly more quickly to the voice in the incongruent condition than in the congruent one. To the best of our knowledge, this is the first study to show that horses cross-modally recognized the emotional states of their caretakers and strangers.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	2045-2322	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Nakamura2018			Serial	6391
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	Author	Burla, J.-B.; Siegwart, J.; Nawroth, C.
	Title	Human Demonstration Does Not Facilitate the Performance of Horses (Equus caballus) in a Spatial Problem-Solving Task			Type	Journal Article
	Year	2018	Publication	Animal	Abbreviated Journal	Animal
	Volume	8	Issue	6	Pages	96
	Keywords	detour task; equids; social cognition; social learning; spatial cognition
	Abstract	Horses’ ability to adapt to new environments and to acquire new information plays an important role in handling and training. Social learning in particular would be very adaptive for horses as it enables them to flexibly adjust to new environments. In the context of horse handling, social learning from humans has been rarely investigated but could help to facilitate management practices. We assessed the impact of human demonstration on the spatial problem-solving abilities of horses during a detour task. In this task, a bucket with a food reward was placed behind a double-detour barrier and 16 horses were allocated to two test groups of 8 horses each. One group received a human demonstration of how to solve the spatial task while the other group received no demonstration. We found that horses did not solve the detour task more often or faster with human demonstration. However, both test groups improved rapidly over trials. Our results suggest that horses prefer to use individual rather than social information when solving a spatial problem-solving task
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6392
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	Author	Gille, C.; Hoischen-Taubner, S.; Spiller, A.
	Title	Neue Reitsportmotive jenseits des klassischen Turniersports			Type	Journal Article
	Year	2011	Publication	Sportwissenschaft	Abbreviated Journal
	Volume	41	Issue	1	Pages	34-43
	Keywords
	Abstract	Während die traditionellen Pferdesportdisziplinen Dressur- und Springreiten Mitglieder verlieren, haben sich weitere Pferdesportarten in Deutschland etabliert und erfreuen sich wachsender Beliebtheit. Die vorliegende Arbeit beschäftigt sich mit den Hintergründen dieser Entwicklung. In einer empirischen Untersuchung wurden 1814 Reiter zu ihren Reitmotiven befragt. Mit Hilfe von Hauptkomponenten- und Clusteranalyse wurde eine Typologie gebildet, die ein differenziertes Bild der Motive verschiedener Reitergruppen ermöglicht. Während die leistungsorientierten Reiter eher in klassischen Reitsportdisziplinen vertreten sind, dominieren in moderneren Reitsportdisziplinen vor allem Genussmotive. Insgesamt entwickelt sich der Trend im Reitsport deutlich vom Drill in der Reitbahn hin zu mehr Entspannung, Erholung und Selbstverwirklichung. Der Wunsch, in der Freizeit Leistung zu bringen, sich mit anderen zu messen und Erfolg zu haben, ist nur noch für einen kleineren Teil der Pferdesportler bedeutsam. Aus der Verteilung der Motive ergeben sich neue Herausforderungen für den organisierten Reitsport, um den Spagat zwischen den Anforderungen der leistungsorientierten Sportreiter und den Erholungssuchenden zu meistern.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1868-1069	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Gille2011			Serial	6393
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	Author	Proops, L.; Grounds, K.; Smith, A.V.; McComb, K.
	Title	Animals Remember Previous Facial Expressions that Specific Humans Have Exhibited			Type	Journal Article
	Year	2018	Publication	Current Biology	Abbreviated Journal
	Volume	28	Issue	9	Pages	1428-1432.e4
	Keywords	affective processing; face processing; ; animal-human interaction; interspecific communication; animal memory
	Abstract	Summary For humans, facial expressions are important social signals, and how we perceive specific individuals may be influenced by subtle emotional cues that they have given us in past encounters. A wide range of animal species are also capable of discriminating the emotions of others through facial expressions [1, 2, 3, 4, 5], and it is clear that remembering emotional experiences with specific individuals could have clear benefits for social bonding and aggression avoidance when these individuals are encountered again. Although there is evidence that non-human animals are capable of remembering the identity of individuals who have directly harmed them [6, 7], it is not known whether animals can form lasting memories of specific individuals simply by observing subtle emotional expressions that they exhibit on their faces. Here we conducted controlled experiments in which domestic horses were presented with a photograph of an angry or happy human face and several hours later saw the person who had given the expression in a neutral state. Short-term exposure to the facial expression was enough to generate clear differences in subsequent responses to that individual (but not to a different mismatched person), consistent with the past angry expression having been perceived negatively and the happy expression positively. Both humans were blind to the photograph that the horses had seen. Our results provide clear evidence that some non-human animals can effectively eavesdrop on the emotional state cues that humans reveal on a moment-to-moment basis, using their memory of these to guide future interactions with particular individuals.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0960-9822	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6394
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	Author	McVey, A.; Wilkinson, A.; Mills, D.S.
	Title	Social learning in horses: the effect of using a group leader demonstrator on the performance of familiar conspecifics in a detour task			Type	Journal Article
	Year	2018	Publication	Applied Animal Behaviour Science	Abbreviated Journal
	Volume		Issue		Pages
	Keywords	Equine; Imitation; Leader; Social facilitation; Social learning
	Abstract	Learning through the observation of others allows the transfer of information without the costs incurred during individual trial and error learning. Horses (Equus caballus) are a highly social species, which might be expected to be capable of learning from others, but experimental findings are inconsistent, and potentially confounded by social facilitation effects not related directly to the learning of the task. We refined the methods used in previous equine social learning studies, to examine and distinguish specific social influences on learning of a task: we used predefined group leaders rather than agonistically dominant individuals to demonstrate a detour task to familiar conspecific observers; in addition we had two control groups: a non-observer (true control) and a group with the demonstrator simply present at the goal (social facilitation control). 44 socially kept horses were allocated to one of the three test conditions and took part in five trials each. Success rate, latency and detour direction were recorded. There was no significant difference between the three groups in the likelihood of them succeeding in the task nor latency to succeed; however there was a significant difference in the route chosen by the groups, with the true control choosing the side with the entrance gate significantly more than either the observer group or social facilitation group. Both of the latter two groups chose to go in the same direction relative to themselves, regardless of which side the gate was. Seven out of nine horses in the observer group chose the same direction as their demonstrator every time. Our results show a significant role of social facilitation on detour behaviour and highlight the importance of including adequate controls for simpler cognitive influences on behaviour before claims can be made about the specific learning of motor actions or goal directed behaviour. Social cues may be important to horses if the task is sufficiently challenging and motivationally important, so future work should consider more demanding, but ecologically relevant situations, in order to maximise the potential revelation of social learning effects which do not depend on simple local or stimulus enhancement effects.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6395
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	Author	Ronnenberg, K.; Habbe, B.; Gräber, R.; Strauß, E.; Siebert, U.
	Title	Coexistence of wolves and humans in a densely populated region (Lower Saxony, Germany)			Type	Journal Article
	Year	2017	Publication	Basic and Applied Ecology	Abbreviated Journal	Basic. Appl. Ecol.
	Volume	25	Issue		Pages	1-14
	Keywords	Grey wolf; ; Habitat use; Species distribution models; Predator-prey interactions
	Abstract	Since the first sporadic occurrences of grey wolves (Canis lupus) west of the Polish border in 1996, wolves have shown a rapid population recovery in Germany. Wolves are known to avoid people and wolf attacks on humans are very rare worldwide. However, the subjectively perceived threat is considerable, especially as food-conditioned habituation to humans occurs sporadically. Lower Saxony (Germany) has an exceedingly higher human population density than most other regions with territorial wolves; thus, the potential for human-wolf conflicts is higher. Using hunters' wildlife survey data from 455 municipalities and two years (2014-2015) and data from the official wolf monitoring (557 confirmed wolf presences and 500 background points) collected between 2012-2015, grey wolf habitat selection was modelled using generalized additive models with respect to human population density, road density, forest cover and roe deer density. Moreover, we tested whether habitat use changed in response to human population and road density between 2012/2013 and 2014/2015. Wolves showed a preference for areas of low road density. Human population density was less important as a covariate in the model of the survey data. Areas with higher prey abundance (5-10 roe deer/km2) and areas with >20% forest cover were preferred wolf habitats. Wolves were mostly restricted to areas with the lowest road and human population densities. However, between the two time periods, avoidance of human density decreased significantly. Recolonization of Germany is still in its early stages and it is unclear where this process will halt. To-date authorities mainly concentrate on monitoring measures. However, to avoid conflict, recolonization will require more stringent management of wolf populations and an improved information strategy for rural populations.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-1791	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6397
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	Author	Galef, B.G.; Laland, K.N.
	Title	Social Learning in Animals: Empirical Studies and Theoretical Models			Type	Journal Article
	Year	2005	Publication	BioScience	Abbreviated Journal
	Volume	55	Issue	6	Pages	489-499
	Keywords
	Abstract	AbstractThe last two decades have seen a virtual explosion in empirical research on the role of social interactions in the development of animals' behavioral repertoires, and a similar increase in attention to formal models of social learning. Here we first review recent empirical evidence of social influences on food choice, tool use, patterns of movement, predator avoidance, mate choice, and courtship, and then consider formal models of when animals choose to copy behavior, and which other animals' behavior they copy, together with empirical tests of predictions from those models.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-3568	ISBN		Medium
	Area		Expedition		Conference
	Notes	10.1641/0006-3568(2005)055[0489:Sliaes]2.0.Co;2			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6398
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