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	Author	Fischhoff, I.R.; Sundaresan, S.R.; Cordingley, J.; Larkin, H.M.; Sellier, M.-J.; Rubenstein, D.I.
	Title	Social relationships and reproductive state influence leadership roles in movements of plains zebra, Equus burchellii			Type	Journal Article
	Year	2007	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	73	Issue	5	Pages	825-831
	Keywords	collective movements; drinking; equid; Equus burchellii; foraging; group dynamics; motivation; plains zebra; reproductive status; social relationships
	Abstract	In animal groups, collective movements emerge from individual interactions. Biologists seek to identify how characteristics of actors in these groups, and their relationships, influence the decision-making process. We distinguished two basic factors determining leadership in group choices: identity and state. We hypothesized that identity is more important to leadership in groups with stable relationships, which permit the development of habitual roles. In groups with fluid membership, particular individuals or subgroups are less likely to emerge as consistent leaders. Instead, we predicted that movement initiation in unstable groups depends on individual state at the time of the decision. We characterized how identity and reproductive state influenced leadership patterns in the movements of plains zebra. As in many other mammals, lactation in this species significantly alters water and energy needs. We investigated leadership in tightly knit harems and loosely bonded herds of multiple harems. Harem females tended to have habitual roles in the initiation of harem movement. In herds, however, we found no consistent leaders among harems. At both levels of social organization, lactation was a key determinant of leadership. In harems, lactating females were more likely to initiate movement than nonlactating females. In turn, harems containing lactating females were more likely to lead herd movements. Thus, we conclude that social relationships and reproductive state together shape the interactions that produce group behaviours. One benefit to lactating females of leading herd movements is preferential access to scarce water. Thus, leadership roles in group decisions may have fitness consequences.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ room B 3.092			Serial	1825
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	Author	Lloyd, A.S.; Martin, J.E.; Bornett-Gauci, H.L.I.; Wilkinson, R.G.
	Title	Evaluation of a novel method of horse personality assessment: Rater-agreement and links to behaviour			Type	Journal Article
	Year	2007	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	105	Issue	1-3	Pages	205-222
	Keywords	Personality; Horse; Individual differences; Behaviour; Reliability
	Abstract	The efficacy of questionnaire-based personality assessment has been shown in a variety of animal and human personality studies. There has been a recent increase in questionnaire-based studies focussing on equine personality but with a lack of comparability to studies on other species. The aim of this study was to test the reliability of an assessment method originally developed for primates and demonstrate reliability using three criteria (1) assessments by independent observers must agree with one another, (2) these assessments must predict behaviours and real-world outcomes and (3) observer ratings must be shown to reflect genuine attributes of the individuals rated, not merely the observer's implicit personality theories about how traits co-vary. The personality of 61 horses (Equus caballus) was assessed using a questionnaire constructed of 30 behaviourally defined adjectives (BDAs). Horses were each assessed by three judges, in addition to a total of 2 h behaviour observations recorded per horse. Rater agreement was demonstrated for 72.1% of the horses and 25 of the BDAs. Principal component analysis was carried out on the rating data and revealed six underlying personality components that were labelled “dominance”, “anxiousness”, “excitability”, “protection”, “sociability” and “inquisitiveness”. Component scores for horses were correlated against behavioural observations for the same horses and revealed significant correlations with 20 of the recorded behaviour variables. Correlations between specific components and their associated behaviours were logical and consistent with the types of behaviours that would be expected to be linked with such personality types. The data were shown to meet all three criteria and provided strong evidence that the assessment method was reliably measuring horse personality.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	1981
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	Author	Visser, E.K.; van Reenen, C.G.; Schilder, M.B.H.; Barneveld, A.; Blokhuis, H.J.
	Title	Learning performances in young horses using two different learning tests			Type	Journal Article
	Year	2003	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	80	Issue		Pages	311-326
	Keywords	Horse; Personality; Learning performance; Consistency; Emotionality
	Abstract	To achieve optimal performance in equine sports as well as in leisure not only the physical abilities of the horse should be considered, but also the horse's personality. Besides temperamental aspects, like emotionality, or the horse's reactivity towards humans in handling situations, the learning ability of the horse is another relevant personality trait. To study whether differences in learning performance are consistent over time and whether individual learning performance differs between learning tests or is affected by emotionality, 39 young horses (Dutch Warmblood) were tested repeatedly in two learning tests. An aversive stimulus (AS) was used in one learning test (the avoidance learning test) and a reward was used in the other learning test (the reward learning test). During both learning tests behaviour as well as heart rate were measured. Each test was executed four times, twice when horses were 1 year of age, and twice when they were 2 years of age. Half of the horses received additional physical training from 6 months onwards. In both tests horses could be classified as either performers, i.e. completing the daily session, or as non-performers, i.e. returning to the home environment without having completed the daily session. There were some indications that emotionality might have caused non-performing behaviour, but these indications are not convincing enough to exclude other causes. Furthermore, there seem to be no simple relationships between measures of heart rate, behavioural responses putatively related to emotionality and learning performance. Horses revealed consistent individual learning performances within years in both tests, and in the avoidance learning test also between years. There was no significant correlation between learning performances in the avoidance learning test and the learning performances in the reward learning test. It is concluded that individual learning abilities are consistent over a short time interval for an avoidance learning test and a reward learning test and over a longer time for the avoidance learning test. Furthermore, results indicate that some horses perform better when they have to learn to avoid an aversive stimulus while others perform better when they are rewarded after a correct response. It is suggested that these differences may be relevant to design optimal individual training programmes and methods.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	2009
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	Author	Dumont, B.; Boissy, A.; Achard, C.; Sibbald, A.M.; Erhard, H.W.
	Title	Consistency of animal order in spontaneous group movements allows the measurement of leadership in a group of grazing heifers			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	95	Issue	1-2	Pages	55-66
	Keywords	Cattle; Grazing; Leadership; Movement order; Walking
	Abstract	The term `leadership' has been used in several different senses, resulting in very different ways of identifying leaders and apparently inconsistent conclusions on how leadership is determined in herbivores. We therefore propose the following definitions: (i) a leader is the individual that is consistently the one who initiates long-distance, spontaneous group movements toward a new feeding site and (ii) long-distance spontaneous group movements are movements which happen when an animal changes activity and location and is immediately followed by a similar change in activity and location by other members of the group. Using these definitions, we tested for consistency of movement order across time and situation within a group of fifteen 2-year-old heifers. We found that the same individual was recorded as the very first animal in 48% of movements toward a new feeding site and could therefore be identified as the `leader'. We also showed that movement order when the animals entered an experimental plot, or progressed slowly through the field during a grazing bout, did not produce the same result. This method, which enables us to identify leaders in groups of animals at pasture, should improve our knowledge of how leadership is determined in grazing herbivores.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ room B 3.029			Serial	2027
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	Author	Lynch, J.J.; Hinch, G.N.; Bouissou, M.F.; Elwin, R.L.; Green, G.C.; Davies, H.I.
	Title	Social organization in young Merino and Merino x Border Leicester ewes			Type	Journal Article
	Year	1989	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	22	Issue	1	Pages	49-63
	Keywords
	Abstract	The social behaviour of two groups of Merino ewes and one group of Merino x Border Leicester ewes was studied. Each group comprised eight sheep, 15 months of age and, within each group, the animals were of similar liveweight. Dominance rankings were established at each test, but there was little consistency in ewe rank over time. Similarly, little consistency was found in ewe ranking for movement order between pens, and for exploratory and fear test rankings. However, with tests on movement orders, some consistency in the sheep which ranked first was shown. In the field, no aggression was seen while sheep were grazing and there were no occasions when ranking related to movement could be observed. There were short-term associations between pairs of sheep, but these occurred in less than half the individuals. Although the spatial distribution was not studied, the lack of long-term association between pairs would suggest that strong spatial preference does not occur. It is concluded that the social organization of single-age Merino and Merino x Border Leicester ewes is not based on dominance or leadership ranking nor on long-term associations between individuals.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ room B 3.029			Serial	2036
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	Author	Southwick, C.H.; Siddiqi, M.R.
	Title	The role of social tradition in the maintenance of dominance in a wild rhesus group			Type	Journal Article
	Year	1967	Publication	Primates	Abbreviated Journal	Primates
	Volume	8	Issue	4	Pages	341-353
	Keywords
	Abstract	Following the injury and disability of the dominant male, the home range of a group of rhesus in a rural habitat in Aligarh district was significantly reduced from 40 acres to less than 10 acres. Throughout this injury and prior to his death, the male maintained his dominance in reference to a peripheral male who frequently attempted to enter the group. Upon the death of the dominant male, group leadership and dominance was assumed by a young subdominant male within the group and the peripheral male still remained outside the group. These observations indicate a strong social tradition in the maintenance of dominance within this wild rhesus group, and they emphasize the role of the dominant male in maintaining home range.
	Address
	Corporate Author				Thesis
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	Notes				Approved	no
	Call Number				Serial	2064
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	Author	Meunier, H.; Leca, J.B.; Deneubourg, J.L.; Petit, O.
	Title	Group movement decisions in capuchin monkeys: the utility of an experimental study and a mathematical model to explore the relationship between individual and collective behaviours			Type	Journal Article
	Year	2006	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	143	Issue		Pages	1511-1527
	Keywords	animal society – collective decision-making – primates – group movement – mathematical modeling
	Abstract	In primate groups, collective movements are typically described as processes dependent on leadership mechanisms. However, in some species, decision-making includes negotiations and distributed leadership. These facts suggest that simple underlying processes may explain certain decision mechanisms during collective movements. To study such processes, we have designed experiments on white-faced capuchin monkeys (Cebus capucinus) during which we provoked collective movements involving a binary choice. These experiments enabled us to analyse the spatial decisions of individuals in the group. We found that the underlying process includes anonymous mimetism, which means that each individual may influence all members of the group. To support this result, we created a mathematical model issued from our experimental data. A totally anonymous model does not fit perfectly with our experimental distribution. A more individualised model, which takes into account the specific behaviour of social peripheral individuals, revealed the validity of the mimetism hypothesis. Even though white-faced capuchins have complex cognitive abilities, a coexistence of anonymous and social mechanisms appears to influence their choice of direction during collective movements. The present approach may offer vital insights into the relationships between individual behaviours and their emergent collective acts.
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	Notes				Approved	no
	Call Number				Serial	2066
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	Author	di Bitetti, M.S.; Janson, C.H.
	Title	Social foraging and the finder's share in capuchin monkeys, Cebus apella			Type	Journal Article
	Year	2001	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	62	Issue	1	Pages	47-56
	Keywords
	Abstract	Group living can confer advantages to individuals, but it can also impose severe costs through resource competition. Kleptoparasitism is one example in which some individuals (joiners) can exploit the food discovered by other animals (finders). This type of social foraging has been modelled either as an information-sharing model or as a producer-scrounger game. An important variable in these models is the finder's advantage: the number of items obtained by the finder before the arrival of other individuals. In this study we describe how the spatial position and rank of individuals in a group of wild tufted capuchin monkeys affect their ability to discover and exploit new food sources. We also analyse the factors that affect the finder's share and the total amount of food obtained by the finder from a newly discovered resource. By placing platforms filled with bananas at novel locations in their home range, we show that animals in the leading edge of a foraging group have a higher probability of discovering new food sources than animals occupying other spatial positions. The alpha male and the alpha female, which tended to occupy central-forward positions, were able to monopolize newly discovered food sources and thus obtain a major share of them. The finder's share at the feeding platforms was smaller when there was more food on a platform, but increased with longer delays before the arrival of other individuals. The total amount of food obtained by the finder from the feeding platforms was larger when there was more food on the platform, when the finder was of higher social status, and when it took longer for other individuals to arrive. Animals can increase their finder's share and total amount consumed from a newly discovered resource by keeping large interindividual distances and by avoiding giving cues about the presence of food (such as food-associated vocalizations) to other animals.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2078
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	Author	Cassini, M.H.; Kacelnik, A.; Segura, E.T.
	Title	The tale of the screaming hairy armadillo, the guinea pig and the marginal value theorem			Type	Journal Article
	Year	1990	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	39	Issue	6	Pages	1030-1050
	Keywords
	Abstract	Foraging by screaming hairy armadillos, Chaetophractus vellerosus, and guinea pigs, Cavia porcellus, was studied in the laboratory. The main question was whether patch exploitation varies with overall capture rate as predicted by the marginal value theorem (MVT). Armadillos in experiment I and guinea pigs in experiment II experienced a single travel time between depleting patches of two kinds: good and poor. There were two treatments, which differed in the quality of poor patches. MVT predicts that within a treatment, more prey should be taken from good than from poor patches and between treatments, good patches should be exploited in inverse relation to the quality of poor patches and poor patches should be exploited in direct relation to their own quality. In experiment III, guinea pigs experienced three treatments which differed in the travel requirement, while the two patch types remained the same. MVT predicts that within a treatment more prey should be taken from good than from poor patches and that between treatments more prey should be taken from both patch types as travel requirement increases. The qualitative predictions were supported in the three experiments. The quantitative fit was good but there was a bias towards more severe patch exploitation than predicted. The results indicate that in these species patch exploitation depends on overall food availability as predicted by the MVT when overall food availability differs either because of patch type composition or because of differences in travel requirement between patches.
	Address
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	Notes				Approved	no
	Call Number				Serial	2120
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	Author	Klingel, H.
	Title	*Zur Sozialstruktur des Steppenzebras, Equus quagga boehmi Matschie.*			Type	Journal Article
	Year	1964	Publication	Die Naturwissenschaften	Abbreviated Journal
	Volume	51	Issue	14	Pages	347
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	Abstract
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
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	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2161
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