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	Author	Leblanc, M.-A.; Duncan, P.
	Title	Can studies of cognitive abilities and of life in the wild really help us to understand equine learning?			Type	Journal Article
	Year	2007	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	76	Issue		Pages	49-52
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	Publisher		Place of Publication		Editor
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	621
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	Author	Duncan, P.; Vigne, N.
	Title	The effect of group size in horses on the rate of attacks by blood-sucking flies			Type	Journal Article
	Year	1979	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	27	Issue	Part 2	Pages	623-625
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	763
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	Author	Duncan, P.
	Title	Determinants of the use of habitat by horses in a mediterranean wetland			Type	Journal Article
	Year	1983	Publication		Abbreviated Journal	J. Anim. Ecol.
	Volume	52	Issue		Pages	93-109
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
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	Area		Expedition		Conference
	Notes	from Professor Hans Klingels Equine Reference List			Approved	no
	Call Number				Serial	1031
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	Author	Boy, V.; Duncan, P.
	Title	Time-budgets of Camargue horses. I. Developmental changes in the time-budgets of foals.			Type	Journal Article
	Year	1979	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	71	Issue		Pages	187-201
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	Corporate Author	20			Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	1803
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	Author	Duncan, P.
	Title	Time-budgets of Camargue horses III. Environmental influences			Type	Journal Article
	Year	1985	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	92	Issue		Pages	188-208
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	Publisher		Place of Publication		Editor
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2283
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	Author	Grange, S.; Duncan, P.
	Title	Bottom-up and top-down processes in African ungulate communities: resources and predation acting on the relative abundance of zebra and grazing bovids			Type	Journal Article
	Year	2006	Publication	Ecography	Abbreviated Journal
	Volume	29	Issue	6	Pages	899-907
	Keywords
	Abstract	African ungulate populations appear to be limited principally by their food resources. Within ungulate communities, plains zebras coexist with grazing bovids of similar body size, but rarely are the dominant species. Given the highly effective nutritional strategy of the equids and the resistance of zebras to drought, this is unexpected and suggests that zebra populations may commonly be limited by other mechanisms. Long-term research in the Serengeti ecosystem and in the Kruger National Park suggests that zebra could be less sensitive to food shortage, and more sensitive to predation, than grazing bovids: if this is a general principle, then, at a larger scale, resource availability should have a weaker effect on the abundance of zebra than on grazing ruminants of similar body size (wildebeest and buffalo), and zebras should be relatively more abundant in ecosystems where predators are rare or absent. We test these expectations using data on 23 near-natural ecosystems in east and southern Africa. The abundance of wildebeest is more closely related to resources than is that of zebra; buffalo are intermediate. We show that hyena densities are closely correlated with those of lions, and use the abundance of lions as an index of predation by large predators. The numerical response of lions to increases in the abundance of their prey was linear for mesoherbivores, and apparently so for the three species alone. Finally, the abundance of zebra relative to grazing bovids is lower in ecosystems with high biomasses of lions. These results indicate that zebras may commonly be more sensitive to top-down processes than grazing bovids: the mechanism(s) have not been demonstrated, but predation could play a role. If it is true, then when numbers of the large mammalian predators decline, zebra populations should increase faster than buffalo and wildebeest.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Grange2006			Serial	2313
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	Author	Mayes, E.; Duncan, P.
	Title	Temporal patterns of feeding behaviour in free-ranging horses			Type	Journal Article
	Year	1986	Publication	Behavior	Abbreviated Journal	Behav.
	Volume	96	Issue		Pages	105-129
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	Corporate Author				Thesis
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2351
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	Author	Monard, A.-M.; Duncan, P.
	Title	Consequences of natal dispersal in female horses			Type	Journal Article
	Year	1996	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	52	Issue	3	Pages	565-579
	Keywords
	Abstract	Social, genetic and reproductive consequences of natal dispersal were investigated in female horses,Equus caballus, living in a herd with a natural social structure. Dispersal did not as a rule reduce the level of competition the young mares faced: they did not selectively join groups with fewer resident females than the groups they left, and they did not attain higher ranks; there was also no tendency for females to disperse to groups with the fewest resident females, and they suffered more aggression from the mares in their new groups than in their natal groups. These results therefore do not support the hypothesis that a function of natal dispersal is to reduce intra-sexual competition. The young mares nevertheless dispersed non-randomly, generally joining harems with one stallion and at least two subadult females; and they preferred to move to groups with familiar females but no familiar males. As a result, most were closely related to some females of their new groups, but distantly related or unrelated to the male(s). Since after dispersal the young mares bred only with a male of their new groups, inbreeding coefficients of most (85%) of their offspring were lower than from matings between half siblings. These results are consistent with the hypothesis that a function of natal dispersal is to avoid close inbreeding. Dispersal did not appear to involve reproductive costs: the young mares suffered no delay in age at first reproduction, and the survival rates of their first foals tended to be higher if the females had emigrated, although not significantly so.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2386
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	Author	Monard, A.-M.; Duncan, P.; Fritz, H.; Feh, C.
	Title	Variations in the birth sex ratio and neonatal mortality in a natural herd of horses			Type	Journal Article
	Year	1997	Publication	Behavioral Ecology and Sociobiology	Abbreviated Journal	Behav. Ecol. Sociobiol.
	Volume	41	Issue	4	Pages	243-249
	Keywords
	Abstract	Variations in birth sex ratios and sex differences in juvenile mortality occur in a number of mammalian species, and in many cases have been linked to resource availability. Most of these biases in offspring sex ratios concern polygynous species with pronounced sexual dimorphism, and where females only are philopatric. Data on species with unusual life-history strategies, such as slight sexual dimorphism or dispersal by both sexes, are of particular interest. In this study of a natural herd of horses (Equus caballus) which experienced an eruptive cycle, and therefore a period of nutritional stress, male offspring had higher neonatal mortality rates in nutritionally poor years than in good ones, whereas “year quality” had no effect on the mortality of female offspring; year quality could therefore be used by mares as predictor of sex-specific offspring survival. We show that the environmental conditions that predicted lower survival of males were negatively related to their production: the birth sex ratio the following year was female-biased; and mares were less likely to produce a son when they had produced a son the preceding year. There was no significant effect of mother's parity, age or rank, or the timing of conception or birth on offspring sex ratios. The mechanism leading to biases in the birth sex ratio could have been the loss of male embryos by mares that did not foal. As there was no evidence for selective abortion of male foetuses in females that did foal the next year, it is not necessary to invoke maternal adjustment, though this remains a possibility. Finally, there was a suggestion that male offspring were more costly to raise than females, since mothers that reared a son in poor years tended to experience an increase in the interbirth interval between their two subsequent offspring.
	Address
	Corporate Author				Thesis
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2388
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	Author	Fleurance, G.; Duncan, P.; Fritz, H.; Cabaret, J.; Cortet, J.; Gordon, I.J.
	Title	Selection of feeding sites by horses at pasture: Testing the anti-parasite theory			Type	Journal Article
	Year	2007	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	108	Issue	3-4	Pages	228-301
	Keywords	Foraging strategies; Horses; Parasite risk; Patch choice
	Abstract	Management of grazed grasslands for production and/or conservation objectives requires a thorough understanding of the choices of feeding sites by herbivores, and of the biological processes involved. Most models of the feeding strategies of herbivores are based on the principle that optimising the intake of energy (or some nutrient) is the primary goal of foragers but other selective forces, such as parasitism, could be important. Gastrointestinal parasites (including cyathostome nematodes) have powerful effects on the fitness of herbivores and may act as a major selection pressure favouring host behaviour that reduces the risk of encountering parasites. Among large herbivores, horses have perhaps the most marked tendency to select particular feeding sites within grasslands. We test here: (1) whether horses select feeding patches with relatively low parasite densities and (2) if their choice is affected by their parasite load. We used 10 two-year old saddle-horses and three periods. In the first period, the horses were under natural parasitism which varied strongly among individuals; in the second period they were all dewormed, and in the third, a sub-set of the horses was experimentally infected with cyathostome larvae. Ninety-eight percent of the infective larvae in the pasture were found <1 m from faeces. The main determinant of the choice of feeding patch by horses was the availability of patches of different parasite risk and grass height. Controlling for availability, the horses used tall grasses (>16 cm) less than expected, whether the grass was contaminated or not, and they selected for short patches >1 m from faeces, where the risk of encountering parasites was low. These results suggest that selection of feeding sites by horses is driven by an interaction between their nutritional and anti-parasite strategies: the horses avoid the patches of tall grass which are generally of low quality and areas contaminated by parasite larvae which leads them to prefer the patches of short grass far from faeces. The parasite status of the horses at the time of the experiment had no effect on their feeding choices. However, before concluding that the challenge by cyathostomes has no effect on the selection of feeding sites in horses, it will be necessary to test whether the history of parasitism of the individuals, rather than the current status, is important.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4228
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