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	Author	Saslow, C.A.
	Title	Understanding the perceptual world of horses			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	78	Issue	2-4	Pages	209-224
	Keywords	Horse; Perception; Vision; Olfaction; Touch; Hearing; Pain; Training; Psychophysics; Umwelt
	Abstract	From the viewpoint of experimental psychology, there are two problems with our current knowledge of equine perception. The first is that the behavioral and neurophysiological research in this area has enormous gaps, reflecting that this animal is not a convenient laboratory subject. The second is that the horse, having been a close companion to humans for many millennia, entrenched anecdotal wisdom is often hard to separate from scientific fact. Therefore, any summary at present of equine perception has to be provisional. The horse appears to have developed a visual system particularly sensitive to dim light and movement, it may or may not have a weak form of color vision in part of the retina, it has little binocular overlap, and its best acuity is limited to a restricted horizontal band which is aimed primarily by head/neck movements. However, the total field of view is very large. Overall, as would be expected for a prey animal, horse vision appears to have evolved more for detection of predator approach from any angle than for accurate visual identification of stationary objects, especially those seen at a distance. It is likely that, as for most mammals except the primates, horses rely more heavily on their other senses for forming a view of their world. Equine high-frequency hearing extends far above that of humans, but horses may be less able to localize the point of origin of brief sounds. The horse's capacity for chemoreception and its reliance on chemical information for identification may more closely resemble that of the dog than of the human. Its tactile sensitivity is high, and the ability of its brain and body to regulate pain perception appears to be similar to that found in other mammals. There is room for a great deal of future research in both the area of equine perception and sensory-based cognition, but for the present time persons interacting with this animal should be made aware of the importance of the sounds they make, the movements of their bodies, the way they touch the animal, and the odors they emit or carry on their clothing.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	400
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	Author	Blackmore, T.L.; Foster, T.M.; Sumpter, C.E.; Temple, W.
	Title	An investigation of colour discrimination with horses (Equus caballus)			Type	Journal Article
	Year	2008	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	78	Issue	3	Pages	387-396
	Keywords	Chromatic discrimination; Colour vision; Horse; Operant
	Abstract	The ability of four horses (Equus caballus) to discriminate coloured (three shades of blue, green, red, and yellow) from grey (neutral density) stimuli, produced by back projected lighting filters, was investigated in a two response forced-choice procedure. Pushes of the lever in front of a coloured screen were occasionally reinforced, pushes of the lever in front of a grey screen were never reinforced. Each colour shade was randomly paired with a grey that was brighter, one that was dimmer, and one that approximately matched the colour in terms of brightness. Each horse experienced the colours in a different order, a new colour was started after 85% correct responses over five consecutive sessions or if accuracy showed no trend over sessions. All horses reached the 85% correct with blue versus grey, three horses did so with both yellow and green versus grey. All were above chance with red versus grey but none reached criterion. Further analysis showed the wavelengths of the green stimuli used overlapped with the yellow. The results are consistent with histological and behavioural studies that suggest that horses are dichromatic. They differ from some earlier data in that they indicate horses can discriminate yellow and blue, but that they may have deficiencies in discriminating red and green.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5336
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	Author	Hirata, S.
	Title	A note on the responses of chimpanzees (Pan troglodytes) to live self-images on television monitors			Type	Journal Article
	Year	2007	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	75	Issue	1	Pages	85-90
	Keywords	Animals; Discrimination Learning; Female; Male; Pan troglodytes/psychology; *Self Concept; Self Psychology; Social Behavior; Television
	Abstract	The majority of studies on self-recognition in animals have been conducted using a mirror as the test device; little is known, however, about the responses of non-human primates toward their own images in media other than mirrors. This study provides preliminary data on the reactions of 10 chimpanzees to live self-images projected on two television monitors, each connected to a different video camera. Chimpanzees could see live images of their own faces, which were approximately life-sized, on one monitor. On the other monitor, they could see live images of their whole body, which were approximately one-fifth life-size, viewed diagonally from behind. In addition, several objects were introduced into the test situation. Out of 10 chimpanzees tested, 2 individuals performed self-exploratory behaviors while watching their own images on the monitors. One of these two chimpanzees successively picked up two of the provided objects in front of a monitor, and watched the images of these objects on the monitor. The results indicate that these chimpanzees were able to immediately recognize live images of themselves or objects on the monitors, even though several features of these images differed from those of their previous experience with mirrors.
	Address	Great Ape Research Institute, Hayashibara Biochemical Laboratories Inc., Okayama, Japan. hirata@gari.be.to
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	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17324534			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4145
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	Author	Kilian, A.; Fersen, L. von; Güntürkün, O.
	Title	Left hemispheric advantage for numerical abilities in the bottlenose dolphin			Type	Journal Article
	Year	2005	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	68	Issue	2	Pages	179-184
	Keywords	Bottlenose dolphin; Hemispheric specialization; Monocular vision; Numerical ability
	Abstract	In a two-choice discrimination paradigm, a bottlenose dolphin discriminated relational dimensions between visual numerosity stimuli under monocular viewing conditions. After prior binocular acquisition of the task, two monocular test series with different number stimuli were conducted. In accordance with recent studies on visual lateralization in the bottlenose dolphin, our results revealed an overall advantage of the right visual field. Due to the complete decussation of the optic nerve fibers, this suggests a specialization of the left hemisphere for analysing relational features between stimuli as required in tests for numerical abilities. These processes are typically right hemisphere-based in other mammals (including humans) and birds. The present data provide further evidence for a general right visual field advantage in bottlenose dolphins for visual information processing. It is thus assumed that dolphins possess a unique functional architecture of their cerebral asymmetries.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5366
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	Author	Cheung, C.; Akiyama, T.E.; Ward, J.M.; Nicol, C.J.; Feigenbaum, L.; Vinson, C.; Gonzalez, F.J.
	Title	Diminished hepatocellular proliferation in mice humanized for the nuclear receptor peroxisome proliferator-activated receptor alpha			Type	Journal Article
	Year	2004	Publication	Cancer research	Abbreviated Journal	Cancer Res
	Volume	64	Issue	11	Pages	3849-3854
	Keywords	Animals; Anticholesteremic Agents/pharmacology; Carcinogens/pharmacology; Cell Division; DNA Replication/drug effects; Fatty Acids/metabolism; Hepatocytes/cytology/drug effects/metabolism/physiology; Humans; Mice; Mice, Transgenic; Oxidation-Reduction; Peroxisome Proliferators/pharmacology; Pyrimidines/pharmacology; Receptors, Cytoplasmic and Nuclear/genetics/physiology; Species Specificity; Transcription Factors/genetics/*physiology
	Abstract	Lipid-lowering fibrate drugs function as agonists for the nuclear receptor peroxisome proliferator-activated receptor alpha (PPARalpha). Sustained activation of PPARalpha leads to the development of liver tumors in rats and mice. However, humans appear to be resistant to the induction of peroxisome proliferation and the development of liver cancer by fibrate drugs. The molecular basis of this species difference is not known. To examine the mechanism determining species differences in peroxisome proliferator response between mice and humans, a PPARalpha-humanized mouse line was generated in which the human PPARalpha was expressed in liver under control of the tetracycline responsive regulatory system. The PPARalpha-humanized and wild-type mice responded to treatment with the potent PPARalpha ligand Wy-14643 as revealed by induction of genes encoding peroxisomal and mitochondrial fatty acid metabolizing enzymes and resultant decrease of serum triglycerides. However, surprisingly, only the wild-type mice and not the PPARalpha-humanized mice exhibited hepatocellular proliferation as revealed by elevation of cell cycle control genes, increased incorporation of 5-bromo-2'-deoxyuridine into hepatocyte nuclei, and hepatomegaly. These studies establish that following ligand activation, the PPARalpha-mediated pathways controlling lipid metabolism are independent from those controlling the cell proliferation pathways. These findings also suggest that structural differences between human and mouse PPARalpha are responsible for the differential susceptibility to the development of hepatocarcinomas observed after treatment with fibrates. The PPARalpha-humanized mice should serve as models for use in drug development and human risk assessment and to determine the mechanism of hepatocarcinogenesis of peroxisome proliferators.
	Address	Laboratory of Metabolism, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland 20892, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0008-5472	ISBN		Medium
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	Notes	PMID:15172993			Approved	no
	Call Number	refbase @ user @			Serial	74
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	Author	Smith, S.; Goldman, L.
	Title	Color discrimination in horses			Type	Journal Article
	Year	1999	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	62	Issue	1	Pages	13-25
	Keywords	Horses; Vision; Color; Discrimination; Behavior
	Abstract	Four Arabian horses and one Thoroughbred were presented with a series of two-choice color vs. gray discrimination problems. Testing was done in a stall containing a wall with two translucent panels that were illuminated from behind by light projected through color or gray filters to provide the discriminative stimuli. Horses first learned to push one of the panels in order to receive the food reward behind the positive stimulus in an achromatic light-dark discrimination task, and were then tested on their ability to discriminate between gray and four individual colors: red (617 nm), yellow (581 nm), green (538 nm), and blue (470 nm). The criterion for learning was set at 85% correct responses, and final testing for all color vs. gray discriminations involved grays of varying intensities, making brightness an irrelevant cue. Three subjects were tested with all four colors. Two of those subjects successfully reached the criterion for learning on all four color vs. gray discriminations, while the third reached criterion with red and blue, but performed at chance levels for yellow and green. A fourth horse was only tested with green and yellow, and a fifth only with blue, and both of those horses successfully reached criterion on the discriminations they attempted. With the exception of the one subject's poor performance with yellow and green, there was no significant difference between horses on any of the discrimination tasks, and no significant difference in their performance with different colors. The results suggest that horses have color vision that is at least dichromatic, although partial color-blindness may occur in some individuals.
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	850
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	Author	Lomas, C.A.; Piggins, D.; Phillips, C.J.C.
	Title	Visual awareness			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	57	Issue	3-4	Pages	247-257
	Keywords	Visual awareness; Colour vision; Rhythm
	Abstract	Awareness varies between different species and humans can never truly appreciate what it is like to be another individual, either of the same species or another. Visual perceptual faculties provide some evidence of the extent to which domesticated animals derive information from objects in their environment, whilst changes in behaviour resulting from different visual stimuli can also provide valuable information on the state of visual awareness. Extensive processing of potentially visual information must occur in all domesticated species, but is much less well understood than purely sensory based information. For example, sensory aspects of colour vision are reasonably well understood, but the role of wavelength variables in an animal's cognition and its colour experience is not clear. Considerable use is made of diurnal changes in photoperiod to synchronise endogenous rhythms to particular times of the day and the year. Variation in light intensity in natural images is also important for social reasons for animals to be able to discriminate between, e.g., different faces, but little is known about intensity preferences or the effects of intensity on behaviour. It appears likely that in many cases visual stimuli represent some of the most important influences on an animal's awareness, either alone or in combination with, e.g., olfactory cues. However, a much greater understanding of their processing is required before we can make useful deductions about visual awareness in domesticated animals.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4309
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	Author	van Niekerk, H.P.
	Title	Ethological studies within the man-horse relationship			Type	Journal Article
	Year	1980	Publication	Journal of the South African Veterinary Association	Abbreviated Journal	J S Afr Vet Assoc
	Volume	51	Issue	4	Pages	237-238
	Keywords	Animals; Behavior, Animal; Hearing; Horses/*physiology; Humans; Smell; Taste; Touch; Vision
	Abstract	Certain aspects of ethology and the horse's senses are discussed to bring about a better understanding between man and horse. Furthermore the behaviour of horses with respect to housing, feeding, breeding, veterinary treatment and work are considered.
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	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN	1019-9128	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7241494			Approved	no
	Call Number	refbase @ user @			Serial	1960
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	Author	Macuda, T.; Timney, B.
	Title	Luminance and chromatic discrimination in the horse (Equus caballus)			Type	Journal Article
	Year	1999	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	44	Issue	3	Pages	301-307
	Keywords	Colour vision; Chromatic discrimination; Luminance discrimination; Horse
	Abstract	Equine colour vision was measured under conditions that minimised the possibility of animals using brightness cues to make chromatic discriminations. In a two-stage study, we first obtained luminance discrimination functions for achromatic targets then tested for chromatic discrimination over a range of target luminances. Horses were trained on a two-choice discrimination task. The positive stimulus was varied in luminance and/or colour using neutral density and broad band colour filters. The negative stimulus appeared as a uniform grey. In the brightness discrimination task, the horses performed well at large luminance differences but their percentage of correct responses declined to near chance levels at differences of less than 0.2 log units. In addition, a decrement in performance was noted at luminance differences of less than 0.2 log units for green and yellow chromatic discrimination functions, suggesting that horses cannot easily discriminate yellow and green from grey. However, the chromatic discrimination functions for red and blue showed that animals performed very well across the full range of target luminances. These results suggest that horses are at least dichromats.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	refbase @ user @			Serial	844
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	Author	Bartoš, L.; Bartošová, J.; Starostová, L.
	Title	Position of the head is not associated with changes in horse vision			Type	Journal Article
	Year	2008	Publication	Equine Veterinary Journal	Abbreviated Journal
	Volume	40	Issue	6	Pages	599-601
	Keywords	horse; horse vision; head position; pupil rotation
	Abstract	It has become accepted that the horse cannot see directly in front when the nose is lowered and must therefore rely on the rider. We tested the hypothesis that this conclusion would be correct only if the horse did not adjust the eyeball horizontal axis to changes of the head position. The results of the present study suggest that it is unlikely that horses have limited vision in relation to their head position when driven by the rider, and that the horse maintains the optimal horizontal eyeball position regardless of head position relative to the ground.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	2042-3306	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5679
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