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Author Bergstrom, C.T.; Lachmann, M. openurl 
  Title Signaling among relatives. III. Talk is cheap Type Journal Article
  Year 1998 Publication Proceedings of the National Academy of Sciences of the United States of America Abbreviated Journal Proc. Natl. Acad. Sci. U.S.A.  
  Volume 95 Issue 9 Pages 5100-5105  
  Keywords Animal Communication; Animals; Costs and Cost Analysis; *Evolution; Interpersonal Relations; Models, Biological  
  Abstract The Sir Philip Sidney game has been used by numerous authors to show how signal cost can facilitate honest signaling among relatives. Here, we demonstrate that, in this game, honest cost-free signals are possible as well, under very general conditions. Moreover, these cost-free signals are better for all participants than the previously explored alternatives. Recent empirical evidence suggests that begging is energetically inexpensive for nestling birds; this finding led some researchers to question the applicability of the costly signaling framework to nestling begging. Our results show that cost-free or inexpensive signals, as observed empirically, fall within the framework of signaling theory.  
  Address Department of Biological Sciences, Stanford University, Stanford, CA 94305, USA. carl@charles.stanford.edu  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0027-8424 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:9560235 Approved no  
  Call Number refbase @ user @ Serial 561  
Permanent link to this record
 

 
Author Gary C. Jahn; Craig Packer,Robert Heinsohn openurl 
  Title Lioness leadership Type Journal Article
  Year 1996 Publication Science (New York, N.Y.) Abbreviated Journal Science  
  Volume 271 Issue 5253 Pages 1216-1219  
  Keywords Animals; *Behavior; Animal; Cooperative Behavior; Female; Lions/*psychology; Territoriality  
  Abstract  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0036-8075 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Jahn1996 Serial 2073  
Permanent link to this record
 

 
Author Broucek, J.; Ksac, P.; Uhrincat, M. openurl 
  Title The effect of sire line on learning and locomotor behaviour of heifers Type Journal Article
  Year 2003 Publication Czech Journal of Animal Science Abbreviated Journal Czech J. Anim. Sci  
  Volume 48 Issue Pages 387-394  
  Keywords heifers; sire; maze; open-field test; repeatability; learning; locomotor behaviour relationship  
  Abstract ABSTRACT: e aim of this study was to test the effect of sire line on maze learning ability and locomotor behaviour

in open-field tests of heifers, consistency over the time of grid crossing and relationship between the time of traversing the maze and grid crossings in open-field tests, respectively. We analysed the results of ethological tests for 54 Holstein heifers that descended from 7 sires. Maze behaviour was observed at the age of 15 weeks, an open-field test was applied at two age periods, 16 weeks and 18 months. We found out highly significant differences in the time of traversing the maze between heifers of different sire origin (P < 0.01). &#57426;e number of grid crossings over the five minutes of the open-field test did not differ between the daughters of the age of 16 weeks and 18 months. Repeatability between the number of grid crossings at the age of 16 weeks and 18 months was proved by significant correlation (r = 0.2713*). On the contrary, significant relationships between the times of traversing the maze and locomotor behaviour in the open-field test (r =-0.3739*) were found only when the sequence of observations followed

after a week pause (age of 15 and 16 weeks).
 
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4322  
Permanent link to this record
 

 
Author Connor, R.C.; Wells, R.S.; Mann, J.; Read,A.J. isbn  openurl
  Title The bottlenose dolphin: Social relationships in a fission-fusion society. Type Book Chapter
  Year 2000 Publication Cetacean Societies: Field Studies of Dolphins and Whales. Abbreviated Journal  
  Volume Issue Pages 91-126  
  Keywords cetacean social behavior, male alliance formation, most cetacean species, platanistid river dolphins, cetacean sociality, strategies and social bonds, female cetaceans, many cetologists, most mysticetes, sperm whale calves, passive fishing nets, variant whistles, historical whaling records, cetacean systematics, stable matrilineal groups, peak calving season, suction cup tags, mutualistic groups, cetacean vocalizations, focal animal studies, larger odontocetes, predictive signaling, individual cetaceans, sperm whale clicks, resident killer whales  
  Abstract Book Description

“Part review, part testament to extraordinary dedication, and part call to get involved, Cetacean Societies highlights the achievements of behavioral ecologists inspired by the challenges of cetaceans and committed to the exploration of a new world.”-from the preface by Richard Wrangham

Long-lived, slow to reproduce, and often hidden beneath the water's surface, whales and dolphins (cetaceans) have remained elusive subjects for scientific study even though they have fascinated humans for centuries. Until recently, much of what we knew about cetaceans came from commercial sources such as whalers and trainers for dolphin acts. Innovative research methods and persistent efforts, however, have begun to penetrate the depths to reveal tantalizing glimpses of the lives of these mammals in their natural habitats.

Cetacean Societies presents the first comprehensive synthesis and review of these new studies. Groups of chapters focus on the history of cetacean behavioral research and methodology; state-of-the-art reviews of information on four of the most-studied species: bottlenose dolphins, killer whales, sperm whales, and humpback whales; and summaries of major topics, including group living, male and female reproductive strategies, communication, and conservation drawn from comparative research on a wide range of species.

Written by some of the world's leading cetacean scientists, this landmark volume will benefit not just students of cetology but also researchers in other areas of behavioral and conservation ecology as well as anyone with a serious interest in the world of whales and dolphins.

Contributors are Robin Baird, Phillip Clapham, Jenny Christal, Richard Connor, Janet Mann, Andrew Read, Randall Reeves, Amy Samuels, Peter Tyack, Linda Weilgart, Hal Whitehead, Randall S. Wells, and Richard Wrangham.
 
  Address  
  Corporate Author Thesis  
  Publisher University of Chicago Press Place of Publication Chicago Editor Mann, J.;Connor, R.C.; Tyack, P.L.;Whitehead, H.  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN 978-0226503417 Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4427  
Permanent link to this record
 

 
Author Granquist,S.M.; Sigurjónsdóttir, H; Thórhallsdóttir, A.G. pdf  openurl
  Title Social structure and interactions within groups of horses containing a stallion Type Conference Article
  Year 2008 Publication IESM 2008 Abbreviated Journal  
  Volume Issue Pages  
  Keywords Social interactions,, Stallions, Hierarchy, Icelandic horses  
  Abstract Earlier research indicates that stallions might either prevent social interactions between mares in their

herds directly or indirectly by their presence (Feist and McCullough, 1976; Sigurjonsdottir et al.,

2003) The impact of stallions on the social interactions between harem members was studied in 6

groups of Icelandic horses in 2004, 2006 and 2007 for altogether 525 hours. Four of the groups were

permanently living together under semi-feral conditions, while two were temporary breeding groups. In

addition, temporary and permanent groups were compared and the effect of the stability of the group

on the social behaviour of horses was investigated .

The results show that stallions seldom intervene directly in social interactions between harem

members. However, the hierarchies were less rigid and fewer friendship bonds were found in the

groups compared to what has been found in groups without stallions in Iceland. These results give

some support to our prediction that the stallion does indirectly suppress social interactions of herd

members.

The stability of the group was found to affect the aggression rate, since a higher aggression rate was

found in the temporary groups compared to the permanent groups. The number of preferred

allogrooming partners of the horses was also affected to some extent, as a significantly lower number

of allogrooming partners was found in the most unstable group compared to all the other groups. The

results have significance for further research in the field of social structure of mammals, and may also

be applied in the management of horses and other domestic animals.
 
  Address  
  Corporate Author Sigurjónsdóttir, H Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference IESM 2008  
  Notes Talk 15 min IESM 2008 Fullpaper Approved yes  
  Call Number Equine Behaviour @ team @ Serial 4487  
Permanent link to this record
 

 
Author Pierard, M. pdf  openurl
  Title Agonistic and affiliative interactions in group housed riding horses (Equus caballus) Type Conference Article
  Year 2012 Publication Proceedings of the 2. International Equine Science Meeting Abbreviated Journal Proc. 2. Int. Equine. Sci. Mtg  
  Volume in press Issue Pages  
  Keywords horse, group housing, social interactions  
  Abstract Group housed horses at a stud farm/riding stable in Belgium were observed on 17 days between 21 February and 25 April 2008, totalling 54hr25min of detailed data. The original group consisted of 8 Irish Cob mares, 1 Warmblood mare, 1 Arabian gelding and 2 Arabian mares. The group had been established in December 2007. During the course of the study 5 horses were removed from the group and 2 foals were born. 3 highly pregnant mares were housed adjacent to the group for part of the period. Horses were regularly used for lessons. Available surface area differed with the group on pasture at the end. Continuous all occurrence sampling of 10 agonistic and 2 affiliative behaviours was carried out for all group members present. Overall the group showed a frequency of 44.75 agonistic interactions per hour and 11.25 affiliative per hour. Of those agonistic interactions 46.3% were threats while 47% were less active interactions (displacement, being avoided), leaving only 6.7% more aggressive interactions ( mainly biting, some kicking and chasing). The effect on acting agonistically was not significant for age (p=0.1591) and borderline significant for density (p=0.0627). The analysis of the frequency of affiliative interactions showed there is no significant effect of age (p=0.1865) or density (p=0.7923). Agonistic and affiliative interactions were not significantly correlated (p=0.72). Affiliative behaviour a horse received showed a borderline effect (p=0.0787) on agonistic behaviour, as did the interaction between received agonistic and affiliative interactions (p=0.0725). Received agonistic interactions had a borderline negative effect (p=0.0656) on affiliative behaviour. A dominance hierarchy was calculated based on agonistic interactions using Empirical Bayes’ estimates based on Poisson regression with random effects. Agonistic behaviour expressed to other horses was significantly effected by relative rank (p=0.0243). Overall horses tended to be 3.7 times more aggressive towards lower ranking horses compared to higher ranking horses. Affiliative behaviour shown to other horses was not significantly influenced by the rank of the social partner (p=0.7915). Some individuals did show a significant effect whereby they showed more affiliative behaviour towards lower ranking individuals. This study was a small project to look at a practical situation of riding horses being kept in group housing. The frequent changes in group composition and available surface made it possible to look at agonistic and affiliative interactions in such circumstances. This is useful as instability in group composition is often used as main reason not to keep horses in social groups. The results from this study showed a borderline effect of density on agonistic behaviour. In reality it was also influenced by practical details, like a narrow paddock with only 2 hay crates on the smallest surface. Rank in a dominance hierarchy, based on agonistic behaviour, had a significant effect on the agonistic behaviour expressed towards higher or lower ranking horses. No injuries or escalating fights were observed. This study shows it is possible to keep a group of riding horses in a social context without excessive aggression.  
  Address  
  Corporate Author Pierard, M. Thesis  
  Publisher Xenophon Publishing Place of Publication Wald Editor Krueger, K.  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 978-3-9808134-26 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5515  
Permanent link to this record
 

 
Author De Giorgio, F.; Schoorl, J.M. pdf  openurl
  Title Why isolate during training? Social learning and social cognition applied as training approach for young horses (Equus caballus) Type Conference Article
  Year 2012 Publication Proceedings of the 2. International Equine Science Meeting Abbreviated Journal Proc. 2. Int. Equine. Sci. Mtg  
  Volume in press Issue Pages  
  Keywords Cognitive approach; Horse training; Horse-Human relationship; Social environment; Social learning  
  Abstract In the last decade an increasing number of studies has been oriented towards equine social learning and their social behavior within the herd (Kruger‚ 2006-2008). In social species, social learning is important to learn and gain useful skills to move and live in their own social and environmental context. Group housing has been recognized as an important element to fulfill the physical and behavioral needs of horses, especially their need for social contact (Søndergaard‚ 2011). Still‚ when it comes to horse training, the social aspect and‚ in general‚ cognitive abilities of the horse are rarely taken into account. Although it is widely accepted that social isolation is stressful for horse (Mal et al, 1991a and 1991b) still isolating a young horse is the first step when it comes to training methods. Due to tradition and culture and our performance-oriented society it is both difficult to accept and apply a different social/cognitive training approach. Training sessions are focused on immediate results whereas in cognitive learning part of the process is latent and will not be visible immediately‚ but taking the cognitive skills into account plays an important role in avoiding tension both in the horse as in the human-horse interaction (Baragli and De Giorgio, 2011). In this study we tested the possibility to apply social learning by creating a social environment‚ favoring a cognitive learning approach‚ for the training of six young horses. The group existed in three males and three females, between two and three years old. All six showed initial difficulties and defense to human interaction. They were housed in two groups in adjacent spacious paddocks where they had ample opportunity to move and express their individual and social behavioral repertoire. Each horse had one training session per week without isolating it from the others. The training sessions were held following a cognitive-relational model defined as the equine-zooanthropologic approach (De Giorgio, 2010 – Marchesini, 2011). The learning objectives were to be able to handle each horse‚ conduct it‚ saddle and ride it within a maximum time-frame of two years. Every time a defensive or alert behavior would occur the training activity was re-arranged to not over-pressure the horse. Therefore the persons working with the horses carried out the activities without tight expectations focusing on the horses’ positive attention. After eighteen months all six horses were used to the saddle and to riding. None of the horses ever fled or showed defense behavior and in the case of unexpected events they showed no emotional reactivity/reactive behavior. Today the horses show the same calm behavior whenever worked individually. This preliminary study highlights how social learning applied to equestrian activity can be fundamental for safety and welfare and the establishment of a more problem-free relationship between horse and human. Safety as the defensive behavior seems to have been reduced and welfare as the horses have been trained in a social context without being isolated and thus without being stressed during the training experience.  
  Address  
  Corporate Author De Giorgio, F. Thesis  
  Publisher Xenophon Publishing Place of Publication Wald Editor Krueger, K.  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 978-3-9808134-26 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5528  
Permanent link to this record
 

 
Author Zeitler-Feicht, M. H.; Streit, S.; Dempfle, L. openurl 
  Title Automatic feeding systems for horses in group housing systems with regard to animal welfare. Part 1: Feeding stalls versus automatic feeding systems Type Journal Article
  Year 2010 Publication Tierärztliche Praxis Großtiere Abbreviated Journal Tierärztl Prax  
  Volume 38 Issue G Pages 363-370  
  Keywords Pferd, Fütterungseinrichtungen, soziale Interaktionen, Herzfrequenz, Verletzungsrisiko, Betriebseinfluss [Horse, feeding stations, social interactions, heart rate, risk of injury, individual farm management]  
  Abstract Gegenstand und Ziel: Überprüfung der Tiergerechtheit von Abrufstationen

für Kraftfutter und Heu in der Gruppenhaltung von Pferden anhand

ethologischer und physiologischer Parameter im Vergleich zu konventionellen

Fressständen. Material und Methoden: Die Verhaltensbeobachtungen

fanden in 11 bzw. 10 Offenlaufställen mit Abrufstationen

bzw. Fressständen statt. 270 Pferde wurden individuell unterschieden.

Untersuchungsparameter waren: Drohverhalten mit und ohne Verletzungsrisiko,

Meideverhalten, Verdrängung aus dem Fressstand sowie

Herzfrequenz und Integumentverletzungen. In jedem Betrieb erfolgten

kontinuierliche Direktbeobachtungen für je einen 24-Stunden-Tag (Tortenstückverfahren).

Ergebnisse: Im Wartebereich der Abrufautomaten

traten signifikant mehr Drohgesten ohne bzw. mit Verletzungsgefahr je

Pferd und Tag (8,6 bzw. 3,0) auf als im Wartebereich der Fressstände

(3,4 bzw. 0,9). Demgegenüber konnten die Pferde in Abrufstationen

(1,4 Drohgesten je Pferd und Tag) ungestörter fressen als in Fressständen

(3,2 Drohgesten je Pferd und Tag). Insgesamt betrachtet ergab sich

jedoch bei beiden Fütterungssystemen eine geringe Anzahl an sozionegativen

Interaktionen im Bereich der Futtereinrichtungen. Die Herzfrequenz

lag im Warteareal im Durchschnitt im physiologischen Bereich

(44,59 ± 11,73 Schläge/min). Integumentverletzungen im Zusammenhang

mit dem Fütterungssystem traten nicht auf. Der wichtigste Einflussfaktor

auf die Untersuchungskriterien war der Betrieb (Fläche, Konzeption,

Management). Schlussfolgerung: Unter dem Aspekt der Tiergerechtheit

hinsichtlich des Stress- und Verletzungsrisikos durch soziale

Interaktionen eignen sich bei ordnungsgemäßer Gruppenhaltung mit

fachgerechtem Management sowohl Fressstände als auch Abrufstationen

für Pferde im Offenlaufstall.

[Objective: A comparison with regard to animal welfare of feeding

stalls and automatic feeding systems for hay and concentrates in group

housing systems for horses using parameters of ethology and physiology.

Material and method: The observations of animal behaviour took

place in 10 stables with feeding stalls and in 11 stables with automatic

feeding systems. The field around the feeding systems was divided into

three areas with comparable dimensions (waiting area, exit area and

inside of the feeding system). 270 horses were individually observed.

Parameters of research comprised: threatening behaviour with and

without risk of injury, avoiding behaviour, ”chasing away from the feeding

place”, as well as heart rate and injuries of the integument. Every

group of horses was continuously observed for 24 hours. This observation

took place on four different days and comprised six sessions, each

of 4 hours. Results: In the waiting area of the automatic systems there

were significantly more threatening gestures with and without risk of

injuries for each horse and day (8.6 and 3.0, respectively) than in the

waiting area of the feeding stalls. In contrast, the horses could eat more

relaxed in the automatic systems (1.4 threatening gestures for each

horse and day) than in the feeding stalls (3.2 threatening gestures for

each horse and day). All together the number of negative interactions in

the feeding area of both feeding systems was relatively low. The heart

rate was within the physiological range (44,59 ± 11,73 beats/min) in

the waiting area. There were no injuries of the integument in correlation

with the feeding systems. The most important factor of the observation

criteria was the individual group housing system with its different dimension,

conception and management. Conclusion: Under the aspect

of animal welfare both feeding systems are suitable for horses with respect

to the risk of stress and injuries by social interactions and under

the condition of proper group housing with professional management.]
 
  Address  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5765  
Permanent link to this record
 

 
Author Zeitler-Feicht, M. H.; Streit, S.; Dempfle, L. openurl 
  Title Automatic feeding systems for horses in group housing systems with regard to animal welfare. Part 2: Comparison of different automatic feeding systems Type Journal Article
  Year 2011 Publication Tierärztliche Praxis Großtiere Abbreviated Journal Tierärztl Prax  
  Volume 39 Issue G Pages 33-40  
  Keywords Pferd, Fütterungssystem, Bauvarianten, soziale Interaktionen, Besuchshäufigkeit, Aufenthaltsdauer, Herzfrequenz, Verletzungsrisiko [Horse, feeding system, different types of construction, social interactions, visiting frequency, duration of stay, heart rate, risk of injury]  
  Abstract Ziel: Überprüfung der Tiergerechtheit von in der Bauweise unterschiedlichen

Futterabrufstationen für Pferde in Gruppenhaltung anhand ethologischer

und physiologischer Parameter. Material und Methoden: In 32

Offenlaufställen (452 Pferde) mit computergesteuerten Abrufstationen

für Kraftfutter und Heu erfolgten kontinuierliche Direktbeobachtungen

für je einen 24-Stunden-Tag (Tortenstückverfahren). Erfasst wurden: Aufenthaltsdauer,

Besuchshäufigkeit, Droh- und Meideverhalten im Fütterungsbereich,

Blockaden sowie Herzfrequenz und Integumentverletzungen.

Ergebnisse: Eine Durchlaufstation reduzierte die Zahl der Auseinandersetzungen

im Fütterungsbereich signifikant. Ansonsten erhöhten die

für das fressende Pferd tiergerechten Varianten (Fressstand mit Eingangssperre,

ohne Austreibehilfe) Besuchshäufigkeit und Aufenthaltsdauer und

steigerten somit auch die Anzahl an Drohgesten je Tier und Tag. Insgesamt

betrachtet kann jedoch die Anzahl an sozionegativen Interaktionen

im Fütterungsbereich der Abrufstationen als relativ gering eingestuft

werden. Die Herzfrequenz lag im Warteareal bei den meisten Pferden im

physiologischen Bereich (45,1 ± 12,42 Schläge/min), erhöhte sich jedoch

in der Abrufstation um ca. 20 Schläge/min. Einige Tiere zeigten möglicherweise

stressbedingt kurzfristig auffallend hohe Werte (&#8805; 100 Schläge/

min). Integumentverletzungen im Zusammenhang mit dem Fütterungssystem

traten nicht auf. Der wichtigste Einflussfaktor auf die Untersuchungskriterien

war der Betrieb (Fläche, Konzeption, Management).

Schlussfolgerung: Durchlaufstationen sind pferdegerechter als Rücklaufstationen.

Nicht tiergerecht sind Stationen mit stromführender Austreibehilfe.

Weitere bauliche Unterschiede der derzeitigen Futterabrufstationen

dürften eher von untergeordneter Bedeutung sein, vor allem da

Flächengebot und Konzeption der Offenstallanlage sowie das Management

die überprüften Kriterien zur Tiergerechtheit maßgeblich beeinflussen.

Untersuchungen zur Abklärung der Ursache für die vereinzelt aufgetretenen

sehr hohen Herzfrequenzwerte sollten durchgeführt werden.

Objective: Comparison with regard to animal welfare of different automatic

feeding systems for hay and concentrate in group housing systems

for horses using parameters of ethology and physiology. Material and

methods: Parameters of research comprised: duration of stay, frequency

of visit, threatening behaviour with and without risk of injury, and avoiding

behaviour as well as heart rate and injuries of the integument. 452

horses were observed at the feeding area of 32 run-out-sheds. Every group

of horses was continuously observed following the pie chart system for

24 hours. Results: The “walk-through” station significantly reduced the

number of conflicts in the feeding area, whereas those systems which are

appropriate for the feeding horses (feeding station with access barrier and

without stimulation device by electric shock) led to a higher frequency of

visits and a longer duration of stay resulting in more threatening gestures.

However, the number of negative interactions in the feeding area of the

feeding systems can all together be classified as relatively insignificant.

The heart rate was within the physiological range (45.1 ± 12.42 beats/min)

in the waiting area, but increased by approximately 20 beats/min on average

within the feeding station. Some horses showed a very high heart rate

(&#8805; 100 beats/min) while entering the feeding station, possibly stress-related.

There were no injuries of the integument associated with the feeding

systems. The most important factor of the observation criteria was the

individual group housing system with its different dimensions, conception

and management. Conclusion: “Walk-through” stations are better than

”walk-back” stations with regard to animal welfare. Likewise, automatic

feeding stations with a current-carrying stimulation device are not supportive

of good welfare. The other differences between the constructions

of feeding stations of present systems are probably of less importance, particularly

as it was shown that the stable (management, stable area, conception)

had a significant influence on the surveyed parameters. Investigations

to obtain information on the cause for the sporadic occurrence of

very high heart rate values should be undertaken.]
 
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5766  
Permanent link to this record
 

 
Author Obergfell, J. openurl 
  Title Einf lus s v on St ruktur e l ement en auf da s Li eg ev e rha l t en v on Pf e rden in Gruppenha l tung unt e r Be rücks i cht igung de s Ag g r e s s i ons v e rha l t ens Type Manuscript
  Year 2012 Publication Abbreviated Journal  
  Volume Issue Pages  
  Keywords Pferde, Ruheverhalten, Liegeverhalten, Aggressionsverhalten, Strukturelemente, Rangordnung [horses, rest behavior, lying behavior, aggression behavior, structural elements, rank order]  
  Abstract Durch die vorliegende Studie wurde der Einfluss von Strukturelementen auf das Liege- und Aggressionsverhalten von Pferden in Gruppenhaltung untersucht. Die Strukturelemente sollten Rückzugsmöglichkeiten bieten, sowie in ihrem Bereich zur Aufhebung der Individualdistanz führen und dadurch die Fläche relativ vergrößern. Für die Versuche stellte das Haupt- und Landesgestüt Marbach drei unabhängige Pferdegruppen mit verschiedener Herdengröße zur Verfügung, die jeweils in Einraum- Innenlaufställen gehalten wurden. Die Datenerfassung fand zwischen 23 und sieben Uhr statt. Insgesamt wurden 366 Stunden Videomaterial ausgewertet. Beim Ruheverhalten wurden mit Hilfe des event-sampling- Verfahrens die Parameter Gesamtliegedauer, Dauer in Seitenlage, Dauer der Einzelphasen in

Seitenlage und Abliegehäufigkeit pro Nacht erfasst. Es wurden Versuchsphasen ohne

Strukturelemente und mit hängenden Planen als Strukturierung durchgeführt. Im ersten Stall

kamen außerdem über einander gestapelte Strohballen zum Einsatz. Diese Art der

Strukturierung stellte sich jedoch als nicht praktikabel heraus und führte im Vergleich zu den Planen zu einer signifikanten Verschlechterung der Gesamtliegedauer. In Stall 1 konnte man eine tendenzielle Verbesserung der Parameter Gesamtliegedauer und Gesamtdauer in Seitenlage durch das Anbringen von Planen sehen. In Stall 3 dagegen verschlechterte sich das Ruheverhalten in den Versuchsphasen mit Planen gegenüber den Versuchsphasen ohne Struktur. Die Werte der Gesamtdauer in Seitenlage nahmen signifikant ab. In Stall 2 zeigten sich keine Unterschiede in den verschiedenen Versuchsphasen. Möglicherweise ist die Wirkung der Strukturelemente auf das Liegeverhalten abhängig von der Flächengröße. Stall 1 hatte bezogen auf die Leitlinien des BMELV die größte und Stall 3 die kleinste Fläche. Bei den anderen Parametern des Liegeverhaltens gab es keine signifikanten Unterschiede in den verschiedenen Versuchsphasen. Mit Hilfe des time-sampling-Verfahrens wurde die Anzahl gleichzeitig liegender Pferde und gleichzeitig liegender Pferde in Seitenlage bestimmt. Auch hier zeigten sich keine Unterschiede in den verschiedenen Versuchsphasen. Bei der Gegenüberstellung der Werte der Gesamtliegedauer und der Gesamtdauer in Seitenlage mit dem Alter der Pferde (Stall 1 und Stall 3) und mit dem Integrationszeitpunkt (Stall 1) konnte kein Zusammenhang festgestellt werden. Beim Aggressionsverhalten wurden mit Hilfe des

event-sampling-Verfahrens in den Ställen 2 und 3 verschiedene Arten von Aggressionen

erfasst, die dann in die drei Intensitätsgrade Low-Level-, Mid-Level- und High-Level-

Aggressionen unterteilt wurden. Neben der Anzahl wurde die Dauer der verschiedenen

Aggressionen bewertet. Insgesamt konnte eine positive Wirkung der Planen auf das

Aggressionsverhalten beobachtet werden. Die Gesamtanzahl an Aggressionen nahm in beiden

Ställen tendenziell in den Versuchsphasen mit Planen ab. In Stall 3 konnte, wenn man die

Aggressionen stundenweise betrachtet, ein signifikanter Unterschied festgestellt werden.

Auch der Hinterhandschlag und die Aggressionen, welche das Ruheverhalten stören,

verringerten sich tendenziell nach dem Anbringen von Strukturelementen. In beiden Ställen

nahm die relative Häufigkeit von Mid-Level-Aggressionen nach dem Anbringen von Planen

zu. Dagegen konnte bei den High-Level-Aggressionen und in Stall 3 bei den Low-Level-

Aggressionen eine relative Abnahme beobachtet werden. Sowohl die Anzahl als auch die

Dauer der Mid-Level-Aggressionen verringerten sich in Stall 3 stundenweise betrachtet

signifikant in den Versuchsphasen mit Planen. In Stall 2 war bei den High-Level-

Aggressionen sowohl bei der Dauer als auch bei der Anzahl eine signifikante Abnahme zu

sehen. Wenn man das Aggressionsverhalten in Bezug zu der Fläche in den zwei Ställen

betrachtet, schien diese vor allem einen Einfluss auf die High-Level-Aggressionen zu

nehmen. Mit Hilfe des Rangindex der Pferde im Stall 3 wurde eine Rangordnung aufgestellt.

Zwischen dem Platz der Pferde in der Rangordnung und den Parametern des Ruheverhaltens

(Gesamtliegedauer, Gesamtdauer in Seitenlage) sowie dem Alter der Pferde konnte kein

Zusammenhang festgestellt werden. Zwischen dem Rangindex und der Gesamtanzahl an

Aggressionen bestand dagegen ein hoch signifikanter Zusammenhang. Im Rahmen dieser

Studie ist das Anbringen von Strukturelementen in Bezug auf das Aggressionsverhalten in

Einraum-Innenlaufställen von Pferden zu empfehlen. Die Wirkung auf das Liegeverhalten der

Pferde sollte in weiteren Studien untersucht werden.

[In this study the influence of structural elements on lying and aggression behavior in horses

kept in groups was exanimated. Structural elements should offer the possibility of retreat.

Furthermore, within the respective area, the individual distance should be limited, thus

increasing the overall space relatively. For the studies the “Haupt- und Landesgestüt

Marbach” provided three non-related groups of different sized horses, each kept in separate

“Einraum-Innenlaufställen”. Data collection took place between 11 pm and 7 am. Altogether,

366 hours of video material were analyzed. Using the event-sampling-method, the following

parameters within the horses´ resting behavior were determined: The total lying period, the

total lying period in lateral position, the periods of the single lying in lateral position and the

frequency of lying down. Test phases were carried out without structural elements and with

hanging canvases as structural elements. In the first stable there were also three bales of straw

piled up on top of each other. This kind of structuring emerged as impractical and in

comparison to the canvas, led to a significant deterioration to the total period of lying. After

the canvases were fixed in the first stable, a slight improvement with the parameters of total

lying period and total lying period in the lateral position could be seen. In contrast, the resting

behavior in stable 3 worsened in the test phases with the canvases in comparison to the test phases without structural elements. The values for the total lying period in the lateral position

decreased significantly. In stable 2, there were no differences in the different test phases.

Possibly the effect of the structural elements is dependent of the space. Stable 1 had the

largest space and stable 3 the least space referred to the guideline of the BMELV. The other

parameters of the lying behavior showed no significant differences in the different test phases.

The number of horses lying at the same time, and the number of horses lying at the same time

in the lateral position were detected with the aid of the time-sampling-method. There were

also no differences between the different test phases. If the values of the total lying period and

the total lying period in lateral position were compared to the age of the horses (stable 1 and

stable 3) and to the point of integration (stable 1), there was no correlation.

Different kinds of aggression were detected by the means of the event-sampling-method in

stable 2 and 3 and were subdivided into the three levels of intensity: Low-level-, mid-level

and high-level-aggressions. Both the number and the duration of the different aggressions

were evaluated. All in all a positive influence of the canvas on the aggression behavior could

be observed. In both stables, the total number of aggressions decreased in the test phases with

canvases. In stable 3 a significant difference could be detected, if the aggressions were

observed per hour. Also hind limb kicks and aggressions disturbing the resting behavior,

decreased by trend, after fixing the canvases in the stable. In both stables the relative

frequency of mid-level-aggressions increased after the canvases were mounted. In contrast

there was a relative decrease at the high-level-aggressions and in stable 3 at the low-levelaggressions.

The number and the duration of the mid-level-aggressions in stable 3 decreased,

when considered by the hour in the test phases with canvases. In stable 2 a significant

decrease in duration and number of high-level-aggressions was observed. When examining

aggression behavior in reference to space in the two stables, high-level-aggressions seemed to

be mostly influenced. In stable 3 a rank order was established with the aid of the rank index of

the horses. There was no correlation between the horses´ rank and the parameters of the

resting behavior (the total lying period, the total lying period in lateral position) and between

horses´ rank and age. In contrast, there was a high significant correlation between the rank

index and the total number of aggressions. In context to this study, structural elements in

“Einraum-Innenlaufställen” (BMELV 2009) seem to have a positive influence on the

aggression behavior and thus seem to be recommendable. The influence on the lying behavior

should be investigated in further studies.]
 
  Address  
  Corporate Author Thesis Ph.D. thesis  
  Publisher Place of Publication Karlsruhe Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5772  
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