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Author Klingel, H. openurl 
  Title Social organization and reproduction in equids Type (up) Journal Article
  Year 1975 Publication Journal of Reproduction and Fertility. Supplement Abbreviated Journal J Reprod Fertil Suppl  
  Volume Issue 23 Pages 7-11  
  Keywords Animals; Behavior, Animal; Female; Male; Perissodactyla/*physiology; Reproduction; *Sexual Behavior, Animal; Social Behavior; Territoriality  
  Abstract There are two distinct types of social organization and, accordingly, two types of mating systems in equids. In the horse, Plains zebra and Mountain zebra, the adults live in non-territorial and cohesive one-male groups and in stallion groups. The family stallions have exclusive mating rights which are respected by all others. In Grevy's zebra and in the African and Asiatic wild asses, the stallions are permanently territorial and have exclusive mating rights within their territories. Ecological and evolutionary aspects are discussed.  
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  Series Volume Series Issue Edition  
  ISSN 0449-3087 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:1060868 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2303  
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Author Zehnder, A.M.; Ramer, J.C.; Proudfoot, J.S. openurl 
  Title The use of altrenogest to control aggression in a male Grant's Zebra (Equus burchelli boehmi) Type (up) Journal Article
  Year 2006 Publication Journal of zoo and wildlife medicine : official publication of the American Association of Zoo Veterinarians Abbreviated Journal J Zoo Wildl Med  
  Volume 37 Issue 1 Pages 61-63  
  Keywords Aggression/*drug effects; Animals; Animals, Zoo; Behavior, Animal/*drug effects; Dose-Response Relationship, Drug; Equidae/*physiology; Female; Horses; Male; Treatment Outcome; Trenbolone/*analogs & derivatives/therapeutic use  
  Abstract A male Grant's Zebra (Equus burchelli boehmi) housed with two mares at the Indianapolis Zoo had a 9-yr history of intermittent aggressive behavior toward mares and other animals. Periods of separation allowed the mares time to heal after sustaining superficial bite wounds. On 26 March 2003, the male (890293) was started on altrenogest at a dosage of 19.8 mg orally once daily to allow reintroduction. The dosage was doubled (40 mg once a day) because of a perceived lack of response. Reintroduction to the mares occurred on 17 May 2003 with no signs of aggression noted. Treatment was reduced to 19.8 mg orally once a day and then discontinued. Altrenogest was restarted at 39.5 mg orally once a day because of the planned introduction of a new mare. There have been no major aggressive displays at this dosage of altrenogest and the dosage has recently been reduced following successful introduction of a new mare.  
  Address University of Florida, 2015 SW 16th Street, Gainesville, Florida 32610, USA  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1042-7260 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17312816 Approved no  
  Call Number Serial 1772  
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Author Hall, C.A.; Cassaday, H.J.; Vincent, C.J.; Derrington, A.M. doi  openurl
  Title Cone excitation ratios correlate with color discrimination performance in the horse (Equus caballus) Type (up) Journal Article
  Year 2006 Publication Journal of comparative psychology (Washington, D.C. : 1983) Abbreviated Journal J Comp Psychol  
  Volume 120 Issue 4 Pages 438-448  
  Keywords Animals; Behavior, Animal; *Color Perception; *Discrimination (Psychology); Discrimination Learning; Horses; Photoreceptors, Vertebrate/*physiology  
  Abstract Six horses (Equus caballus) were trained to discriminate color from grays in a counterbalanced sequence in which lightness cues were irrelevant. Subsequently, the pretrained colors were presented in a different sequence. Two sets of novel colors paired with novel grays were also tested. Performance was just as good in these transfer tests. Once the horse had learned to select the chromatic from the achromatic stimulus, regardless of the specific color, they were immediately able to apply this rule to novel stimuli. In terms of the underlying visual mechanisms, the present study showed for the first time that the spectral sensitivity of horse cone photopigments, measured as cone excitation ratios, was correlated with color discrimination performance, measured as accuracy, repeated errors, and latency of approach.  
  Address School of Animal, Rural and Environmental Sciences, Nottingham Trent University, Southwell, United Kingdom. carol.hall@ntu.ac.uk  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0735-7036 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17115866 Approved no  
  Call Number Serial 1780  
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Author Henry, S.; Richard-Yris, M.-A.; Hausberger, M. doi  openurl
  Title Influence of various early human-foal interferences on subsequent human-foal relationship Type (up) Journal Article
  Year 2006 Publication Developmental psychobiology Abbreviated Journal Dev Psychobiol  
  Volume 48 Issue 8 Pages 712-718  
  Keywords Animal Husbandry/methods; Animals; Animals, Newborn/*psychology; Animals, Suckling/*psychology; Behavior, Animal; Female; *Handling (Psychology); Horses/*psychology; Humans; Mothers/psychology; *Object Attachment; Species Specificity  
  Abstract Whereas the way animals perceive human contact has been particularly examined in pet animals, a small amount of investigations has been done in domestic ungulates. It was nevertheless assumed that, as pet animals, non-aggressive forms of tactile contact were as well rewarding or positive for these species, even though the features of intraspecific relationships in pet animals and domestic ungulates may be to some extent different.We test here the hypothesis that horses may not consider physical handling by humans as a positive event. When comparing different early human-foal interactions, we found that early exposure to a motionless human enhanced slightly foals reactions to humans whereas forced stroking or handling in early life did not improve later human-foal relation. Foals that were assisted during their first suckling (e.g., brought to the dam's teat) even tended to avoid human approach at 2 weeks, and physical contact at 1 month of age.We argue that interspecies differences may exist in how tactile stimulation is perceived. It may be important for the establishment of a bond that a young animal is active in the process and able, through its behavioral responses, to help define what is positive for it. This way of investigation may have important general implications in how we consider the development of social relations, both within and between species.  
  Address UMR CNRS 6552, Ethologie-Evolution-Ecologie, Universite de Rennes 1, Avenue du General Leclerc, Campus de Beaulieu, F-35042 Rennes Cedex, France. severine.henry@univ-rennes1.fr  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0012-1630 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17111402 Approved no  
  Call Number Serial 1781  
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Author Christensen, J.W.; Rundgren, M.; Olsson, K. openurl 
  Title Training methods for horses: habituation to a frightening stimulus Type (up) Journal Article
  Year 2006 Publication Equine veterinary journal Abbreviated Journal Equine Vet J  
  Volume 38 Issue 5 Pages 439-443  
  Keywords Animals; Behavior, Animal/*physiology; *Conditioning (Psychology); *Fear; Heart Rate/*physiology; Horses/physiology/*psychology; Learning; Male; Random Allocation; *Reinforcement (Psychology); Safety; Time Factors  
  Abstract REASONS FOR PERFORMING STUDY: Responses of horses in frightening situations are important for both equine and human safety. Considerable scientific interest has been shown in development of reactivity tests, but little effort has been dedicated to the development of appropriate training methods for reducing fearfulness. OBJECTIVES: To investigate which of 3 different training methods (habituation, desensitisation and counter-conditioning) was most effective in teaching horses to react calmly in a potentially frightening situation. HYPOTHESES: 1) Horses are able to generalise about the test stimulus such that, once familiar with the test stimulus in one situation, it appears less frightening and elicits a reduced response even when the stimulus intensity is increased or the stimulus is presented differently; and 2) alternative methods such as desensitisation and counter-conditioning would be more efficient than a classic habituation approach. METHODS: Twenty-seven naive 2-year-old Danish Warmblood stallions were trained according to 3 different methods, based on classical learning theory: 1) horses (n = 9) were exposed to the full stimulus (a moving, white nylon bag, 1.2 x 0.75 m) in 5 daily training sessions until they met a predefined habituation criterion (habituation); 2) horses (n = 9) were introduced gradually to the stimulus and habituated to each step before the full stimulus was applied (desensitisation); 3) horses (n = 9) were trained to associate the stimulus with a positive reward before being exposed to the full stimulus (counter-conditioning). Each horse received 5 training sessions of 3 min per day. Heart rate and behavioural responses were recorded. RESULTS: Horses trained with the desensitisation method showed fewer flight responses in total and needed fewer training sessions to learn to react calmly to test stimuli. Variations in heart rate persisted even when behavioural responses had ceased. In addition, all horses on the desensitisation method eventually habituated to the test stimulus whereas some horses on the other methods did not. CONCLUSIONS AND POTENTIAL RELEVANCE: Desensitisation appeared to be the most effective training method for horses in frightening situations. Further research is needed in order to investigate the role of positive reinforcement, such as offering food, in the training of horses.  
  Address Danish Institute of Agricultural Sciences, Department of Animal Health, Welfare and Nutrition, Tjele, Denmark  
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  Series Volume Series Issue Edition  
  ISSN 0425-1644 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:16986605 Approved no  
  Call Number Serial 1783  
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Author McComb, K.; Clutton-Brock, T. doi  openurl
  Title Is mate choice copying or aggregation responsible for skewed distributions of females on leks? Type (up) Journal Article
  Year 1994 Publication Proceedings. Biological sciences / The Royal Society Abbreviated Journal Proc Biol Sci  
  Volume 255 Issue 1342 Pages 13-19  
  Keywords Animals; Deer/*physiology; Estrus/physiology; Female; Male; Phenotype; Sexual Behavior, Animal/*physiology; Territoriality  
  Abstract In several lek-breeding populations of birds and mammals, females arriving on leks tend to join males that already have females in their territories. This might occur either because females have an evolved preference for mating with males that are attractive to other females, or because they join groups of other females to obtain greater safety from predation or dangerous harassment by males. We have previously used controlled experiments to show that oestrous fallow deer females join males with established harems because they are attracted to female groups rather than to the males themselves. Here we demonstrate that the preference for males with females over males without females is specific to oestrous females and weak or absent in anoestrous ones, and that it is not associated with a preference for mating with males that have previously been seen to mate with other females. Furthermore, oestrous females given the choice between males that do not already have females with them show no significant preference for antlered over deantlered males or for older males over younger ones. We conclude that female attraction to other females on the lek is likely to be an adaptation to avoiding harassment in mixed-sex herds. In this situation, a male's ability to maintain the cohesion of his harem may be the principal cause of variation in mating success between males.  
  Address Department of Zoology, University of Cambridge, U.K  
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  ISSN 0962-8452 ISBN Medium  
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  Notes PMID:8153135 Approved no  
  Call Number Serial 1823  
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Author Uehara, T.; Yokomizo, H.; Iwasa, Y. doi  openurl
  Title Mate-choice copying as Bayesian decision making Type (up) Journal Article
  Year 2005 Publication The American naturalist Abbreviated Journal Am Nat  
  Volume 165 Issue 3 Pages 403-410  
  Keywords Animals; *Bayes Theorem; *Choice Behavior; Female; Male; *Models, Biological; *Sexual Behavior, Animal  
  Abstract Mate-choice copying by females has been reported in fishes (e.g., guppies) and lekking birds. Presumably, females assess males' quality using both information from direct observation of males and information acquired by observing other females' choices. Here, we study mathematically the conditions under which mate-choice copying is advantageous on the basis of Bayesian decision theory. A female may observe the mate choice of another female, called the model female, who has performed an optimal choice based on her own judgment. The conditions required for the focal female to choose the same mate as that chosen by the model female should depend on the male's appearance to her, the reliability of her own judgment of male quality, and the reliability of the model females. When three or more females are involved, the optimal mate choice critically depends on whether multiple model females make decisions independently or they themselves copy the choices of others. If two equally reliable females choose different males, the choice of the second female, made knowing the choice of the first, should have a stronger effect on the choice of the third (focal) female. This “last-choice precedence” should be tested experimentally.  
  Address Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka 812-8581, Japan. uehara@bio-math.biology.kyushu-u.ac.jp  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1537-5323 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:15729669 Approved no  
  Call Number Serial 1821  
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Author Dugatkin, L.A.; Godin, J.G. doi  openurl
  Title Reversal of female mate choice by copying in the guppy (Poecilia reticulata) Type (up) Journal Article
  Year 1992 Publication Proceedings. Biological sciences / The Royal Society Abbreviated Journal Proc Biol Sci  
  Volume 249 Issue 1325 Pages 179-184  
  Keywords Acclimatization; Animals; *Choice Behavior; Female; Male; Poecilia; *Sexual Behavior, Animal  
  Abstract Ever since Fisher (1958) formalized models of sexual selection, female mate choice has been assumed to be a genetically determined trait. Females, however, may also use social cues to select mates. One such cue might be the mate choice of conspecifics. Here we report the first direct evidence that a female's preference for a particular male can in fact be reversed by social cues. In our experiments using the Trinidadian guppy (Poecilia reticulata), this reversal was mediated by mate-copying opportunities, such that a female (the 'focal' female) is given the opportunity to choose between two males, followed by a period in which she observes a second female (the 'model' female) displaying a preference for the male she herself did not prefer initially. When allowed to choose between the same males a second time, compared with control tests, a significant proportion of focal females reversed their mate choice and copied the preference of the model female. These results provide strong evidence for the role of non-genetic factors in sexual selection and underlie the need for new models of sexual selection that explicitly incorporate both genetic and cultural aspects of mate choice.  
  Address Department of Biology, Mount Allison University, Sackville, New Brunswick, Canada  
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  Series Volume Series Issue Edition  
  ISSN 0962-8452 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:1360679 Approved no  
  Call Number Serial 1824  
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Author Elhay, M.; Newbold, A.; Britton, A.; Turley, P.; Dowsett, K.; Walker, J. doi  openurl
  Title Suppression of behavioural and physiological oestrus in the mare by vaccination against GnRH Type (up) Journal Article
  Year 2007 Publication Australian Veterinary Journal Abbreviated Journal Aust Vet J  
  Volume 85 Issue 1-2 Pages 39-45  
  Keywords Animals; Antibodies/blood; Estradiol/blood; *Estrus/drug effects/physiology; Female; Gonadotropin-Releasing Hormone/*immunology/*pharmacology; Horses/*physiology; Luteinizing Hormone/blood; Ovulation/*drug effects/physiology; Progesterone/blood; Safety; Sexual Behavior, Animal/drug effects/physiology; Time Factors; Vaccination/veterinary  
  Abstract OBJECTIVE: To examine the immunogenicity of an equine immunocontraceptive vaccine and its efficacy in controlling hormone-related behaviour. DESIGN: A total of 24 mares at two sites in Australia were vaccinated with an immunocontraceptive vaccine comprising gonadotrophin releasing hormone (GnRH) conjugated to a carrier protein in immunostimulating complex as an adjuvant. Twelve animals at each site received a placebo of adjuvant alone and served as controls for seasonal oestrus, hormonal and behaviour patterns. Animals were observed for injection site reactions, ovarian and follicular activity, and serum levels of antibody, 17beta-oestradiol and progesterone in the weeks following vaccination. Mares were also examined for oestrous behaviour by teasing with a stallion. RESULTS: All mares responded to vaccination. Two weeks following the second vaccination there was a peak in antibody response to GnRH that declined gradually over the following weeks. Commensurate with the elevated anti-GnRH antibody there was a marked effect on ovarian activity with a reduction in 17beta-oestradiol and progesterone levels in the 24 vaccinated mares. There was also a reduction of oestrus-related behaviour as determined by a teaser stallion. This effect lasted a minimum of 3 months and correlated with the initial level of antibody response. CONCLUSION: Following a conventional two-dose immunisation regime this commercially available equine immunocontraceptive vaccine was effective at inhibiting oestrous behaviour for at least 3 months. This vaccine has a high level of safety since there were no significant local reactions nor were there any adverse systemic responses to vaccination.  
  Address Veterinary Medicines Research and Development, Pfizer Animal Health, Parkville, VIC 3052. Martin.Elhay@pfizer.com  
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  Series Volume Series Issue Edition  
  ISSN 0005-0423 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17300452 Approved no  
  Call Number Serial 1831  
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Author Momozawa, Y.; Takeuchi, Y.; Tozaki, T.; Kikusui, T.; Hasegawa, T.; Raudsepp, T.; Chowdhary, B.P.; Kusunose, R.; Mori, Y. doi  openurl
  Title SNP detection and radiation hybrid mapping in horses of nine candidate genes for temperament Type (up) Journal Article
  Year 2007 Publication Animal Genetics Abbreviated Journal Anim Genet  
  Volume 38 Issue 1 Pages 81-83  
  Keywords Animals; *Behavior, Animal; Breeding; Horses/*genetics/physiology; *Polymorphism, Single Nucleotide; Radiation Hybrid Mapping; *Temperament  
  Abstract  
  Address Laboratory of Veterinary Ethology, The University of Tokyo, Tokyo 113-8657, Japan  
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  Series Volume Series Issue Edition  
  ISSN 0268-9146 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17257195 Approved no  
  Call Number Serial 1834  
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