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	Author	Dorrance, B.R.; Zentall, T.R.
	Title	Imitation of conditional discriminations in pigeons (Columba livia)			Type	Journal Article
	Year	2002	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	116	Issue	3	Pages	277-285
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Conditioning (Psychology)/physiology; Discrimination Learning/physiology; Habituation, Psychophysiologic; *Imitative Behavior; Light; Reinforcement (Psychology)
	Abstract	In the present experiments, the 2-action method was used to determine whether pigeons could learn to imitate a conditional discrimination. Demonstrator pigeons (Columba livia) stepped on a treadle in the presence of 1 light and pecked at the treadle in the presence of another light. Demonstration did not seem to affect acquisition of the conditional discrimination (Experiment 1) but did facilitate its reversal of the conditional discrimination (Experiments 2 and 3). The results suggest that pigeons are not only able to learn a specific behavior by observing another pigeon, but they can also learn under which circumstances to perform that behavior. The results have implications for proposed mechanisms of imitation in animals.
	Address	Department of Psychology, Augustana College, Rock Island, Illinois 61201, USA. psdorrance@augustana.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12234078			Approved	no
	Call Number	refbase @ user @			Serial	240
Permanent link to this record



	Author	Henning, J.M.; Zentall, T.R.
	Title	Imitation, social facilitation, and the effects of ACTH 4-10 on rats' bar-pressing behavior			Type	Journal Article
	Year	1981	Publication	The American journal of psychology	Abbreviated Journal	Am J Psychol
	Volume	94	Issue	1	Pages	125-134
	Keywords	Adrenocorticotropic Hormone/pharmacology; Animals; Conditioning, Operant/drug effects; Dose-Response Relationship, Drug; Extinction, Psychological/drug effects; Imitative Behavior/drug effects; Male; Peptide Fragments/pharmacology; Rats; *Social Facilitation
	Abstract	The effects of ACTH 4-10 on rats' imitation learning was examined during the acquisition and extinction of a bar-press response for water reinforcement. Rats were exposed to either a bar-pressing conspecific (OB), an experimentally naive conspecific (ON), or an empty box (OE) during bar-press acquisition. In a factorial design, each rat was then exposed to one of the same three conditions during extinction. An 80 mcg dose of ACTH 4-10 was administered to half of the rats in each group prior to observation. Performance differences during acquisition were generally small, but significant performance differences during extinction were found. Social facilitation was indicated by the finding that rats extinguished in the presence of a conspecific exhibited significantly greater resistance to extinction than rats extinguished in the presence of an empty box. An imitation effect was also found. Rats that observed a bar-pressing conspecific during both acquisition and extinction (group OB-OB) showed significantly greater resistance top extinction than did groups OB-ON, CB-OE, or OE-OE. There were no significant effects of the hormone, however, relative to saline controls.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0002-9556	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:6263117			Approved	no
	Call Number	refbase @ user @			Serial	267
Permanent link to this record



	Author	Zentall, T.R.
	Title	Imitation: definitions, evidence, and mechanisms			Type	Journal Article
	Year	2006	Publication	Animal cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	4	Pages	335-353
	Keywords	Adaptation, Psychological; Animals; Behavior, Animal; Imitative Behavior; Learning; Motivation; Social Environment; Transfer (Psychology)
	Abstract	Imitation can be defined as the copying of behavior. To a biologist, interest in imitation is focused on its adaptive value for the survival of the organism, but to a psychologist, the mechanisms responsible for imitation are the most interesting. For psychologists, the most important cases of imitation are those that involve demonstrated behavior that the imitator cannot see when it performs the behavior (e.g., scratching one's head). Such examples of imitation are sometimes referred to as opaque imitation because they are difficult to account for without positing cognitive mechanisms, such as perspective taking, that most animals have not been acknowledged to have. The present review first identifies various forms of social influence and social learning that do not qualify as opaque imitation, including species-typical mechanisms (e.g., mimicry and contagion), motivational mechanisms (e.g., social facilitation, incentive motivation, transfer of fear), attentional mechanisms (e.g., local enhancement, stimulus enhancement), imprinting, following, observational conditioning, and learning how the environment works (affordance learning). It then presents evidence for different forms of opaque imitation in animals, and identifies characteristics of human imitation that have been proposed to distinguish it from animal imitation. Finally, it examines the role played in opaque imitation by demonstrator reinforcement and observer motivation. Although accounts of imitation have been proposed that vary in their level of analysis from neural to cognitive, at present no theory of imitation appears to be adequate to account for the varied results that have been found.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA. Zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17024510			Approved	no
	Call Number	refbase @ user @			Serial	217
Permanent link to this record



	Author	Dorrance, B.R.; Zentall, T.R.
	Title	Imitative learning in Japanese quail (Coturnix japonica) depends on the motivational state of the observer quail at the time of observation			Type	Journal Article
	Year	2001	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	115	Issue	1	Pages	62-67
	Keywords	Animals; Behavior, Animal; Coturnix; Female; Imitative Behavior; Learning; Male; *Motivation; Reinforcement (Psychology); Time Factors
	Abstract	The 2-action method was used to examine whether imitative learning in Japanese quail (Coturnix japonica) depends on the motivational state of the observer quail at the time of observation of the demonstrated behavior. Two groups of observers were fed before observation (satiated groups), whereas 2 other groups of observers were deprived of food before observation (hungry groups). Quail were tested either immediately following observation or after a 30-min delay. Results indicated that quail in the hungry groups imitated, whereas those in the satiated groups did not, regardless of whether their test was immediate or delayed. The results suggest that observer quail may not learn (through observation) behavior that leads to a reinforcer for which they are unmotivated at the time of test. In addition, the results show that quail are able to delay the performance of a response acquired through observation (i.e., they show deferred imitation).
	Address	Department of Psychology, University of Kentucky, Lexington 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11334220			Approved	no
	Call Number	refbase @ user @			Serial	245
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	Author	Akins, C.K.; Klein, E.D.; Zentall, T.R.
	Title	Imitative learning in Japanese quail (Coturnix japonica) using the bidirectional control procedure			Type	Journal Article
	Year	2002	Publication	Animal learning & behavior	Abbreviated Journal	Anim Learn Behav
	Volume	30	Issue	3	Pages	275-281
	Keywords	Animals; Attention; Behavior, Animal; Coturnix; Discrimination Learning; Imitative Behavior; Male; Smell
	Abstract	In the bidirectional control procedure, observers are exposed to a conspecific demonstrator responding to a manipulandum in one of two directions (e.g., left vs. right). This procedure controls for socially mediated effects (the mere presence of a conspecific) and stimulus enhancement (attention drawn to a manipulandum by its movement), and it has the added advantage of being symmetrical (the two different responses are similar in topography). Imitative learning is demonstrated when the observers make the response in the direction that they observed it being made. Recently, however, it has been suggested that when such evidence is found with a predominantly olfactory animal, such as the rat, it may result artifactually from odor cues left on one side of the manipulandum by the demonstrator. In the present experiment, we found that Japanese quail, for which odor cues are not likely to play a role, also showed significant correspondence between the direction in which the demonstrator and the observer push a screen to gain access to reward. Furthermore, control quail that observed the screen move, when the movement of the screen was not produced by the demonstrator, did not show similar correspondence between the direction of screen movement observed and that performed by the observer. Thus, with the appropriate control, the bidirectional procedure appears to be useful for studying imitation in avian species.
	Address	University of Kentucky, Lexington, Kentucky 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0090-4996	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12391793			Approved	no
	Call Number	refbase @ user @			Serial	239
Permanent link to this record



	Author	Akins, C.K.; Zentall, T.R.
	Title	Imitative learning in male Japanese quail (Coturnix japonica) using the two-action method			Type	Journal Article
	Year	1996	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	110	Issue	3	Pages	316-320
	Keywords	Animals; Appetitive Behavior; Attention; Coturnix; Imitative Behavior; Male; Motivation; Transfer (Psychology)
	Abstract	The study of imitative learning in animals has suffered from the presence of a number of confounding motivational and attentional factors (e.g., social facilitation and stimulus enhancement). The two-action method avoids these problems by exposing observers to demonstrators performing a response (e.g., operating a treadle) using 1 of 2 distinctive topographies (e.g., by pecking or by stepping). Japanese quail (Coturnix japonica) observers exposed to conspecific demonstrators showed a high correlation between the topography of the response they observed and the response they performed. These data provide strong evidence for the existence of true imitative learning in an active, precocious bird under conditions that control for alternative accounts.
	Address	Department of Psychology, University of Kentucky, Lexington 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:8858851			Approved	no
	Call Number	refbase @ user @			Serial	254
Permanent link to this record



	Author	Zentall, T.R.; Kaiser, D.H.
	Title	Interval timing with gaps: gap ambiguity as an alternative to temporal decay			Type	Journal Article
	Year	2005	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	31	Issue	4	Pages	484-486
	Keywords	Animals; Behavior, Animal; Discrimination (Psychology)/physiology; Memory/physiology; Rats; Time Perception/*physiology
	Abstract	C. V. Buhusi, D. Perera, and W. H. Meck (2005) proposed a hypothesis of timing in rats to account for the results of experiments that have used the peak procedure with gaps. According to this hypothesis, the introduction of a gap causes the animal's memory for the pregap interval to passively decay (subjectively shorten) in direct proportion to the duration and salience of the gap. Thus, animals should pause with short, nonsalient gaps but should reset their clock with longer, salient gaps. The present authors suggest that the ambiguity of the gap (i.e., the similarity between the gap and the intertrial interval in both appearance and relative duration) causes the animal to actively reset the clock and prevents adequate assessments of the fate of timed intervals prior to the gap. Furthermore, when the intertrial interval is discriminable from the gap, the evidence suggests that timed intervals prior to the gap are not lost but are retained in memory.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16248734			Approved	no
	Call Number	refbase @ user @			Serial	220
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	Author	Zentall, T.R.; Clement, T.S.
	Title	Memory mechanisms in pigeons: evidence of base-rate neglect			Type	Journal Article
	Year	2002	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	28	Issue	1	Pages	111-115
	Keywords	Animals; Columbidae; Discrimination Learning; Memory/*physiology; Random Allocation; Reaction Time; Reinforcement (Psychology); Retention (Psychology)
	Abstract	In delayed matching to sample, once acquired, pigeons presumably choose comparisons according to their memory for (the strength of) the sample. When memory for the sample is sufficiently weak, comparison choice should depend on the history of reinforcement associated with each of the comparison stimuli. In the present research, pigeons acquired two matching tasks in which Sample S1 was associated with one comparison from each task, C1 and C3, whereas Sample S2 was associated with Comparison C2, and Sample S3 was associated with Comparison C4. As the retention interval increased, the pigeons showed a bias to choose the comparison (C1 or C3) associated with the more frequently occurring sample (S1). Thus, pigeons were sensitive also to the (irrelevant) likelihood that each of the samples was presented. The results suggest that pigeons may allow their reference memory for the overall sample frequency to influence comparison choice, independent of the comparison stimuli present.
	Address	Department of Psychology, University of Kentucky, Lexington, Kentucky 40506-0044, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11868229			Approved	no
	Call Number	refbase @ user @			Serial	242
Permanent link to this record



	Author	Zentall, T.R.
	Title	Mental time travel in animals: a challenging question			Type	Journal Article
	Year	2006	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	72	Issue	2	Pages	173-183
	Keywords	Animals; Behavior, Animal; Columbidae; Concept Formation; Conditioning, Operant; Imagination; Memory; Mental Recall; Planning Techniques; Rats; Time Perception; Transfer (Psychology)
	Abstract	Humans have the ability to mentally recreate past events (using episodic memory) and imagine future events (by planning). The best evidence for such mental time travel is personal and thus subjective. For this reason, it is particularly difficult to study such behavior in animals. There is some indirect evidence, however, that animals have both episodic memory and the ability to plan for the future. When unexpectedly asked to do so, animals can report about their recent past experiences (episodic memory) and they also appear to be able to use the anticipation of a future event as the basis for a present action (planning). Thus, the ability to imagine past and future events may not be uniquely human.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16466863			Approved	no
	Call Number	refbase @ user @			Serial	218
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	Author	Zentall, T.R.; Roper, K.L.; Sherburne, L.M.
	Title	Most directed forgetting in pigeons can be attributed to the absence of reinforcement on forget trials during training or to other procedural artifacts			Type	Journal Article
	Year	1995	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	63	Issue	2	Pages	127-137
	Keywords	Animals; Attention; Color Perception; Columbidae; Cues; Discrimination Learning; Mental Recall; Motivation; Pattern Recognition, Visual; Reinforcement Schedule; Retention (Psychology)
	Abstract	In research on directed forgetting in pigeons using delayed matching procedures, remember cues, presented in the delay interval between sample and comparisons, have been followed by comparisons (i.e., a memory test), whereas forget cues have been followed by one of a number of different sample-independent events. The source of directed forgetting in delayed matching to sample in pigeons was examined in a 2 x 2 design by independently manipulating whether or not forget-cue trials in training ended with reinforcement and whether or not forget-cue trials in training included a simultaneous discrimination (involving stimuli other than those used in the matching task). Results were consistent with the hypothesis that reinforced responding following forget cues is sufficient to eliminate performance deficits on forget-cue probe trials. Only when reinforcement was omitted on forget-cue trials in training (whether a discrimination was required or not) was there a decrement in accuracy on forget-cue probe trials. When reinforcement is present, however, the pattern of responding established during and following a forget cue in training may also play a role in the directed forgetting effect. These findings support the view that much of the evidence for directed forgetting using matching procedures may result from motivational and behavioral artifacts rather than the loss of memory.
	Address	Department of Psychology, University of Kentucky, Lexington 40506
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7714447			Approved	no
	Call Number	refbase @ user @			Serial	256
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