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	Author	Mejdell, C.M.; Buvik, T.; Jørgensen, G.H.M.; Bøe, K.E.
	Title	Horses can learn to use symbols to communicate their preferences			Type	Journal Article
	Year	2016	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	184	Issue		Pages	66-73
	Keywords	Operant conditioning; Blanket; Rug; Thermoregulation; Cognition; Clicker training
	Abstract	Abstract This paper describes a method in which horses learn to communicate by touching different neutral visual symbols, in order to tell the handler whether they want to have a blanket on or not. Horses were trained for 10–15 min per day, following a training program comprising ten steps in a strategic order. Reward based operant conditioning was used to teach horses to approach and touch a board, and to understand the meaning of three different symbols. Heat and cold challenges were performed to help learning and to check level of understanding. At certain stages, a learning criterion of correct responses for 8–14 successive trials had to be achieved before proceeding. After introducing the free choice situation, on average at training day 11, the horse could choose between a “no change” symbol and the symbol for either “blanket on” or “blanket off” depending on whether the horse already wore a blanket or not. A cut off point for performance or non-performance was set to day 14, and 23/23 horses successfully learned the task within this limit. Horses of warm-blood type needed fewer training days to reach criterion than cold-bloods (P < 0.05). Horses were then tested under differing weather conditions. Results show that choices made, i.e. the symbol touched, was not random but dependent on weather. Horses chose to stay without a blanket in nice weather, and they chose to have a blanket on when the weather was wet, windy and cold (χ2 = 36.67, P < 0.005). This indicates that horses both had an understanding of the consequence of their choice on own thermal comfort, and that they successfully had learned to communicate their preference by using the symbols. The method represents a novel tool for studying preferences in horses.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6022
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	Author	Keeling, L.J.; Bøe, K.E.; Christensen, J.W.; Hyyppä, S.; Jansson, H.; Jørgensen, G.H.M.; Ladewig, J.; Mejdell, C.M.; Särkijärvi, S.; Søndergaard, E.; Hartmann, E.
	Title	Injury incidence, reactivity and ease of handling of horses kept in groups: a matched case control study in four Nordic countries			Type	Journal Article
	Year		Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl Anim Behav Sci
	Volume		Issue		Pages
	Keywords	horse; behaviour; reactivity; injury; welfare; management
	Abstract	Abstract There is increasing interest in keeping horses in groups, but progress is hampered by a lack of knowledge about which horses can and should be kept together. Therefore, our objective was to investigate the effect of group composition on the occurrence of injuries among horses, the ease of removing horses from groups and horses’ reactivity to a fearful stimulus. Using a matched case control design, 61 groups of horses were studied in Denmark, Norway, Finland and Sweden. They were allocated into groups of similar or different age and sex or where membership changed regularly or remained stable. Injuries were recorded before mixing the horses into treatment groups, the day after mixing and four weeks later. Reactivity of horses to a moving novel object and the behaviour of a horse being removed from its group and the reactions of other group members towards this horse and the handler were evaluated. It was hypothesized that a more socially variable group composition has beneficial effects on behaviour, ease of handling and reducing reactivity whereas frequent changes in group composition has negative consequences, resulting in more injuries. We found that differences in treatment effects were mainly related to breed, rather than group composition. Icelandic horses reacted less to the movement of the novel object (P = 0.007) and approached it more afterwards (P = 0.04). They also had fewer new injuries than warmbloods following mixing (P < 0.001) and fewer than all other groups 4 weeks later (P < 0.01). Most new injuries after mixing were minor and recorded on the horse’s head, chest, hind legs and rump. In conclusion, variations in sex and age composition of the group had little effect on injury level, reactivity and ease of handling compared to the general effect of breed. Concerns about the risk of severe injuries associated with keeping horses in groups are probably overestimated. Thus, we propose that horses can be successfully kept in groups of different sex and age composition.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6020
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	Author	Krueger, K.; Hollenhorst, H.; Schuetz, A.; Weil, S.
	Title	Social learning and innovative learning in horses.			Type	Conference Article
	Year	2015	Publication	Proceedings of the 3. International Equine Science Meeting	Abbreviated Journal
	Volume	3	Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Xenophon Publishing	Place of Publication	Wald	Editor	K. Krueger
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-3-95625-000-2	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5956
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	Author	Gorodnichenko, Y.; Roland, G.
	Title	Individualism, innovation, and long-run growth			Type	Journal Article
	Year	2011	Publication	Proceedings of the National Academy of Sciences	Abbreviated Journal	Proc. Natl. Acad. Sci. U.S.A.
	Volume	108	Issue	Supplement 4	Pages	21316-21319
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes	10.1073/pnas.1101933108			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5941
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	Author	Young, H.P.
	Title	The dynamics of social innovation			Type	Journal Article
	Year	2011	Publication	Proceedings of the National Academy of Sciences	Abbreviated Journal	Proc. Natl. Acad. Sci. U.S.A.
	Volume	108	Issue	Supplement 4	Pages	21285-21291
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes	10.1073/pnas.1100973108			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5940
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	Author	Whiten, A.; Byrne, R.W.
	Title	Tactical deception in primates			Type	Journal Article
	Year	1988	Publication	Behavioral and Brain Sciences	Abbreviated Journal	Behav. Brain Sci.
	Volume	11	Issue	02	Pages	233-244
	Keywords
	Abstract	ABSTRACT Tactical deception occurs when an individual is able to use an “honest” act from his normal repertoire in a different context to mislead familiar individuals. Although primates have a reputation for social skill, most primate groups are so intimate that any deception is likely to be subtle and infrequent. Published records are sparse and often anecdotal. We have solicited new records from many primatologists and searched for repeating patterns. This has revealed several different forms of deceptive tactic, which we classify in terms of the function they perform. For each class, we sketch the features of another individual's state of mind that an individual acting with deceptive intent must be able to represent, thus acting as a “natural psychologist.” Our analysis will sharpen attention to apparent taxonomic differences. Before these findings can be generalized, however, behavioral scientists must agree on some fundamental methodological and theoretical questions in the study of the evolution of social cognition.
	Address
	Corporate Author				Thesis
	Publisher	Cambridge Journals Online	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1469-1825	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5937
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	Author	Ayala, I.; Martos, N.F.; Silvan, G.; Gutierrez-Panizo, C.; Clavel, J.G.; Illera, J.C.
	Title	Cortisol, adrenocorticotropic hormone, serotonin, adrenaline and noradrenaline serum concentrations in relation to disease and stress in the horse			Type	Journal Article
	Year	2012	Publication	Research in Veterinary Science	Abbreviated Journal
	Volume	93	Issue	1	Pages	103-107
	Keywords	Horse; Disease; Cortisol; Acth; Serotonin; Catecholamines; Stress
	Abstract	No detailed comparative data are available on the hormonal parameters of horses suffering from a number of diseases. The aim of our study was to measure concentrations of cortisol, adrenocorticotropic hormone (ACTH), serotonin, adrenaline and noradrenaline in horses with various diseases and following surgery, to assess the response of the HPA axis and adrenal medulla. Blood samples were obtained from six groups of horses comprising a total of 119 animals as follows: laminitis, acute abdominal syndrome (AAS), castration surgery, acute diseases, chronic diseases and healthy controls. Serum hormonal concentrations were determined for each group for comparison. Statistically significant differences between all groups and controls were found for cortisol, ACTH (except for castration), serotonin and adrenaline concentrations but only in horses with laminitis and AAS for noradrenaline. No statistically significant differences were found between males and females. The largest changes in the pituitary–adrenal axis activity occurred mainly in acute diseases, laminitis and in the AAS group.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0034-5288	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5935
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	Author	Legare, C.H.; Nielsen, M.
	Title	Imitation and Innovation: The Dual Engines of Cultural Learning			Type	Journal Article
	Year		Publication	Trends in Cognitive Sciences	Abbreviated Journal	Trends in Cognitive Sciences
	Volume	19	Issue	11	Pages	688-699
	Keywords
	Abstract	Imitation and innovation work in tandem to support cultural learning in children and facilitate our capacity for cumulative culture. Here we propose an integrated theoretical account of how the unique demands of acquiring instrumental skills and cultural conventions provide insight into when children imitate, when they innovate, and to what degree. For instrumental learning, with an increase in experience, high fidelity imitation decreases and innovation increases. By contrast, for conventional learning, imitative fidelity stays high, regardless of experience, and innovation stays low. We synthesize cutting edge research on the development of imitative flexibility and innovation to provide insight into the social learning mechanisms underpinning the uniquely human mind.
	Address
	Corporate Author				Thesis
	Publisher	Elsevier	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1364-6613	ISBN		Medium
	Area		Expedition		Conference
	Notes	doi: 10.1016/j.tics.2015.08.005			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5931
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	Author	Rørvang, M.V.; Ahrendt, L.P.; Christensen, J.W.
	Title	A trained demonstrator has a calming effect on naïve horses when crossing a novel surface			Type	Journal Article
	Year	2015	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	171	Issue		Pages	117-120
	Keywords	Fear; Habituation; Social learning; Social transmission; Heart rate
	Abstract	Abstract Habituated horses have been found to have a calming effect on conspecifics in fear-eliciting situations. In practice, experienced horses are often used as companions when young horses are introduced to potentially frightening situations, like loading onto a trailer. However, studies of social transmission of habituation in horses are scarce. This study investigated if demonstration by a habituated demonstrator horse influenced the willingness of young Icelandic horses (n = 22, 3 years old) to cross a novel surface. Observer horses (n = 11) were allowed to observe the similarly aged demonstrator horse being led five times across a novel surface. Immediately afterwards the observer horses were given the opportunity to cross the novel surface themselves to obtain food on the other side. Controls (n = 11) were allowed to observe the demonstrator eating on the opposite side of the novel surface but not the demonstration of crossing the novel surface. All observers and controls succeeded the task, but observers had significantly lower average and maximum heart rate, compared to controls. This result suggests a calming effect of the demonstration, which could be exploited for habituation training of horses in fear-eliciting situations.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5922
Permanent link to this record



	Author	Krützen, M.; Mann, J.; Heithaus, M.R.; Connor, R.C.; Bejder, L.; Sherwin, W.B.
	Title	Cultural transmission of tool use in bottlenose dolphins			Type	Journal Article
	Year	2005	Publication	Proceedings of the National Academy of Sciences of the United States of America	Abbreviated Journal	Proc. Natl. Acad. Sci. U.S.A.
	Volume	102	Issue	25	Pages	8939-8943
	Keywords
	Abstract	In Shark Bay, wild bottlenose dolphins (Tursiops sp.) apparently use marine sponges as foraging tools. We demonstrate that genetic and ecological explanations for this behavior are inadequate; thus, “sponging” classifies as the first case of an existing material culture in a marine mammal species. Using mitochondrial DNA analyses, we show that sponging shows an almost exclusive vertical social transmission within a single matriline from mother to female offspring. Moreover, significant genetic relatedness among all adult spongers at the nuclear level indicates very recent coancestry, suggesting that all spongers are descendents of one recent “Sponging Eve.” Unlike in apes, tool use in this population is almost exclusively limited to a single matriline that is part of a large albeit open social network of frequently interacting individuals, adding a new dimension to charting cultural phenomena among animals.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes	10.1073/pnas.0500232102			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5916
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