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	Author	Bílá, K.; Beránková, J.; Veselý, P.; Bugnyar, T.; Schwab, C.
	Title	Responses of urban crows to con- and hetero-specific alarm calls in predator and non-predator zoo enclosures			Type	Journal Article
	Year	2017	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	20	Issue	1	Pages	43-51
	Keywords
	Abstract	Urban animals and birds in particular are able to cope with diverse novel threats in a city environment such as avoiding novel, unfamiliar predators. Predator avoidance often includes alarm signals that can be used also by hetero-specifics, which is mainly the case in mixed-species flocks. It can also occur when species do not form flocks but co-occur together. In this study we tested whether urban crows use alarm calls of conspecifics and hetero-specifics (jackdaws, Corvus monedula) differently in a predator and a non-predator context with partly novel and unfamiliar zoo animal species. Birds were tested at the Tiergarten Schönbrunn in the city of Vienna by playing back con- and hetero-specific alarm calls and control stimuli (great tit song and no stimuli) at predator (wolf, polar bear) and non-predator (eland antelope and cranes, peccaries) enclosures. We recorded responses of crows as the percentage of birds flying away after hearing the playback (out of those present before the playback) and as the number of vocalizations given by the present birds. A significantly higher percentage of crows flew away after hearing either con- or hetero-specific alarm calls, but it did not significantly differ between the predator and the non-predator context. Crows treated jackdaw calls just as crow calls, indicating that they make proper use of hetero-specific alarm calls. Responding similarly in both contexts may suggest that the crows were uncertain about the threat a particular zoo animal represents and were generally cautious. In the predator context, however, a high percentage of crows also flew away upon hearing the great tit control song which suggests that they may still evaluate those species which occasionally killed crows as more dangerous and respond to any conspicuous sound.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Bílá2017			Serial	6159
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	Author	Murray, L.M.A.; Byrne, K.; D’Eath, R.B.
	Title	Pair-bonding and companion recognition in domestic donkeys, <em>Equus asinus</em>			Type	Journal Article
	Year	2013	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	143	Issue	1	Pages	67-74
	Keywords
	Abstract	Pair and social bonding has been documented in various taxa, where pair formations are often described as being driven by kinship or sexual motivation. However, pair-bonding between unrelated individuals where sexual motivation is not a factor is not well documented. Many social relationships and pair-bonds between members of a dyad are facilitated by each individual's ability to recognise their partner using cues which are characteristic of that particular individual. The aims of this study were i) to investigate the existence of pair-bonding in domestic donkeys and ii) to determine whether members of a dyad could recognise their companion during a Y-maze recognition test. Subjects were 55 unrelated donkeys (38 gelded males, 15 females) in seven groups of mixed or same sex, comprising 4?14 individuals. Spatial proximity (nearest-neighbour) was observed three times a day over a 22-day period. Using a simulation approach based on observed data to generate randomised nearest-neighbour matrices, the statistical significance of social relationships was estimated. Of these, 42 (79.2%) were involved in significantly (p<0.05) non-random nearest-neighbour relationships, most of which were reciprocal pair relationships. Based on the spatial data, 24 of the donkeys which had shown significant reciprocal nearest-neighbour preferences for one individual (companion) were then used in a Y-maze recognition test in which they were presented with a choice of their companion and either a familiar donkey from the same group or an unfamiliar donkey from a different group. Donkeys? spatial location in the Y-maze demonstrated a preference for their companion versus familiar (one sample Wilcoxon signed rank test, W=239, p=0.002) or unfamiliar donkeys (W=222, p=0.041). These results verify anecdotal evidence from donkey handlers that donkeys often form pair-bonds, and show that reciprocal social preference and recognition are the basis of these. Pair-bond formation and companionship among donkeys have potential implications for their management, husbandry and welfare.
	Address
	Corporate Author				Thesis
	Publisher	Elsevier	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes	doi: 10.1016/j.applanim.2012.11.005			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6149
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	Author	Brooks, C.J.; Harris, S.
	Title	Directed movement and orientation across a large natural landscape by zebras, Equus burchelli antiquorum			Type	Journal Article
	Year	2008	Publication	Animal Behaviour	Abbreviated Journal	Anim. Behav.
	Volume	76	Issue	2	Pages	277-285
	Keywords	correlated random walk; directed movement; Equus burchelli antiquorum; Gps; movement path; orientation; spatial memory; spatial scale; zebra
	Abstract	We investigated how plains zebras moved across a large natural landscape by analysing the movement paths of nine zebra mares foraging out from spatially confined waterholes during the dry season in the Makgadikgadi Pans National Park, Botswana. Since it was essential to investigate directed movement over a range of spatial scales to determine the correct movement behaviour and strategy, we used Nams's scaling test for oriented movement. Zebras followed directed movement paths in the lower to medium spatial scales (10 m–3.7 km) and above their visual, and possibly olfactory, range. The spatial scale of directed movement suggests that zebras had a well-defined spatial awareness and cognitive ability. Seven zebras used directed movement paths, but the remaining two followed paths not significantly different to a correlated random walk (CRW). At large spatial scales (>3 km) no distinct movement pattern could be identified and paths could not be distinguished from a CRW. Foraging strategy affected the extent of directed movement: zebras with a confined dispersion of grazing patches around the central place directed their movements over a longer distance. Zebras may extend the distance at which they can direct their movement after improving their knowledge of the local environment.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6148
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	Author	White, A.M.; Swaisgood, R.R.; Czekala, N.
	Title	Ranging patterns in white rhinoceros, Ceratotherium simum simum: implications for mating strategies			Type	Journal Article
	Year	2007	Publication	Animal Behaviour	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	74	Issue	2	Pages	349-356
	Keywords	Ceratotherium simum simum; foraging; habitat choice; home range use; mate choice; mating systems; ranging pattern; resource distribution; white rhinoceros
	Abstract	How animals use space has important consequences for feeding ecology, social organization, mating strategies and conservation management. In white rhinoceros, female home ranges are much larger than male territories, suggesting that movement patterns are influenced by factors other than resource distribution. In this study we placed radiotransmitters on 15 female white rhinoceros, recording 1758 locations and collecting behavioural data during 1671 observation sessions, making this the largest data set of its kind in this species. We investigated how habitat variables and male territories influenced female movement and reproductive behaviour. Female home ranges were approximately 20 km2 and core areas were 5 km2, with male territories roughly the same size as female core areas. Female range size did not vary with season, but the pattern of space use did vary. Females used grassland habitat preferentially, utilizing these areas significantly more than expected based on availability. Findings relevant to the mating strategy include: (1) the amount of grassland in a male's territory predicted female use of the territory; (2) the time that a female spent in a male's territory was a significant predictor of reproductive activity with the male, indicating that females probably mate with the most familiar male; and (3) the temporal pattern of female space use suggests that females did not increase mate sampling behaviour nor did they become more choosy about which males they visited when reproductively active. These findings suggest that males may maximize reproductive success by defending areas containing more grassland habitat.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6146
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	Author	Daniel, J.C.; Mikulka, P.J.
	Title	Discrimination learning in the white rhinoceros			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	58	Issue	1–2	Pages	197-202
	Keywords	Rhinoceros; Learning
	Abstract	This study examined the ability of two adult white rhinoceroses (Ceratotherium simum simum) to develop a visual discrimination between an open circle and a triangle. These stimuli were presented as black symbols on large white cards. The cards were presented 4.6 m apart and a food reward was given if the subject approached the open circle. Ten discrimination choices were given daily until each subject reached the criterion of 80% correct responding over a block of 50 trials. The female reached the criterion over trials 151–200, while the male required considerably longer (trials 501–550). The male's discrimination was dramatically affected by a shift in the food reward. This study demonstrates that these rhinos were able to develop a successful discrimination and this protocol could be used to further examine their visual acuity.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6145
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	Author	Cinková, I.; Policht, R.
	Title	Sex and species recognition by wild male southern white rhinoceros using contact pant calls			Type	Journal Article
	Year	2016	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	19	Issue	2	Pages	375-386
	Keywords
	Abstract	Recognition of information from acoustic signals is crucial in many animals, and individuals are under selection pressure to discriminate between the signals of conspecifics and heterospecifics or males and females. Here, we first report that rhinos use information encoded in their calls to assess conspecifics and individuals of closely related species. The southern (Ceratotherium simum) and critically endangered northern (C. cottoni) white rhinos are the most social out of all the rhinoceros species and use a contact call pant. We found that southern white rhino pant calls provide reliable information about the caller’s sex, age class and social situation. Playback experiments on wild territorial southern white rhinoceros males revealed that they responded more strongly to the pant calls of conspecific females compared to the calls of other territorial males. This suggests that pant calls are more important form of communication between males and females than between territorial males. Territorial southern males also discriminated between female and territorial male calls of northern species and reacted more intensively to the calls of northern than southern males. This might be caused by a novelty effect since both species naturally live in allopatry. We conclude that white rhinos can directly benefit from assessing individuals at long distances using vocal cues especially because their eyesight is poor. Pant calls thus likely play a significant role in their social relationships and spatial organization. In addition, better understanding of vocal communication in white rhinos might be helpful in conservation management particularly because of their low reproduction in captivity.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Cinková2016			Serial	6144
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	Author	Cinková, I.; Policht, R.
	Title	Discrimination of familiarity and sex from chemical cues in the dung by wild southern white rhinoceros			Type	Journal Article
	Year	2015	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	18	Issue	1	Pages	385-392
	Keywords
	Abstract	Communication in rhinos is primarily mediated by the vocal and olfactory signals as they have relatively poor eyesight. White rhinos are the most social of all the rhinoceros species, they defecate at common dungheaps and the adult bulls use dung and urine to mark their territory. Chemical communication may therefore be particularly important in the social interactions of white rhinos, and its knowledge could be very helpful in their management and conservation. However, no studies have investigated up until now the olfactory discrimination in any rhinoceros species in the wild. We have experimentally studied the reactions of the wild southern white rhinos (Ceratotherium simum) to the dung of familiar and unfamiliar adult females and adult territorial males. We registered the number of sniffing events, the duration of sniffing and the latency of the vigilance posture from the onset of sniffing. The dung of unfamiliar rhinos was sniffed longer than that of familiar rhinos. The rhinos showed a shorter latency of vigilance posture to the familiar dung of males than that of females. For unfamiliar dung, they displayed a shorter latency of vigilance posture to female than male dung. Our results indicate that the rhinos are able to discriminate the familiarity and sex of conspecifics from the smell of their dung. Olfactory cues could therefore play an important role in the social relationships and spatial organization of the southern white rhinoceros.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Cinková2015			Serial	6143
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	Author	Heberlein, M.T.E.; Manser, M.B.; Turner, D.C.
	Title	Deceptive-like behaviour in dogs (Canis familiaris)			Type	Journal Article
	Year	2017	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	20	Issue	3	Pages	511-520
	Keywords
	Abstract	Deception, the use of false signals to modify the behaviour of the receiver, occurs in low frequencies even in stable signalling systems. For example, it can be advantageous for subordinate individuals to deceive in competitive situations. We investigated in a three-way choice task whether dogs are able to mislead a human competitor, i.e. if they are capable of tactical deception. During training, dogs experienced the role of their owner, as always being cooperative, and two unfamiliar humans, one acting ‘cooperatively’ by giving food and the other being ‘competitive’ and keeping the food for themselves. During the test, the dog had the options to lead one of these partners to one of the three potential food locations: one contained a favoured food item, the other a non-preferred food item and the third remained empty. After having led one of the partners, the dog always had the possibility of leading its cooperative owner to one of the food locations. Therefore, a dog would have a direct benefit from misleading the competitive partner since it would then get another chance to receive the preferred food from the owner. On the first test day, the dogs led the cooperative partner to the preferred food box more often than expected by chance and more often than the competitive partner. On the second day, they even led the competitive partner less often to the preferred food than expected by chance and more often to the empty box than the cooperative partner. These results show that dogs distinguished between the cooperative and the competitive partner, and indicate the flexibility of dogs to adjust their behaviour and that they are able to use tactical deception.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Heberlein2017			Serial	6136
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	Author	Ringhofer, M.; Yamamoto, S.
	Title	Erratum to: Domestic horses send signals to humans when they are faced with an unsolvable task			Type	Journal Article
	Year	2017	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	20	Issue	3	Pages	407-407
	Keywords
	Abstract	Some domestic animals are thought to be skilled at social communication with humans due to the process of domestication. Horses, being in close relationship with humans, similar to dogs, might be skilled at communication with humans. Previous studies have indicated that they are sensitive to bodily signals and the attentional state of humans; however, there are few studies that investigate communication with humans and responses to the knowledge state of humans. Our first question was whether and how horses send signals to their potentially helpful but ignorant caretakers in a problem-solving situation where a food item was hidden in a bucket that was accessible only to the caretakers. We then examined whether horses alter their behaviours on the basis of the caretakers’ knowledge of where the food was hidden. We found that horses communicated to their caretakers using visual and tactile signals. The signalling behaviour of the horses significantly increased in conditions where the caretakers had not seen the hiding of the food. These results suggest that horses alter their communicative behaviour towards humans in accordance with humans’ knowledge state.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Ringhofer2017			Serial	6135
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	Author	Ringhofer, M.; Yamamoto, S.
	Title	Domestic horses send signals to humans when they face with an unsolvable task			Type	Journal Article
	Year	2017	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	20	Issue	3	Pages	397-405
	Keywords
	Abstract	Some domestic animals are thought to be skilled at social communication with humans due to the process of domestication. Horses, being in close relationship with humans, similar to dogs, might be skilled at communication with humans. Previous studies have indicated that they are sensitive to bodily signals and the attentional state of humans; however, there are few studies that investigate communication with humans and responses to the knowledge state of humans. Our first question was whether and how horses send signals to their potentially helpful but ignorant caretakers in a problem-solving situation where a food item was hidden in a bucket that was accessible only to the caretakers. We then examined whether horses alter their behaviours on the basis of the caretakers’ knowledge of where the food was hidden. We found that horses communicated to their caretakers using visual and tactile signals. The signalling behaviour of the horses significantly increased in conditions where the caretakers had not seen the hiding of the food. These results suggest that horses alter their communicative behaviour towards humans in accordance with humans’ knowledge state.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9456	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Ringhofer2017			Serial	6134
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