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Author Faria, J.J.; Dyer, J.R.G.; Tosh, C.R.; Krause, J.
Title Leadership and social information use in human crowds Type Journal Article
Year 2010 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 79 Issue 4 Pages 895-901
Keywords collective animal behaviour; group; human; inadvertent social cue; information; leadership
Abstract One of the big challenges for group-living animals is to find out who in a group has pertinent information (regarding food or predators) at any moment in time, because informed individuals may not be obviously recognizable to other group members. We found that individuals in human groups were capable of identifying those with information, and this identification increased group performance: the speed and accuracy of groups in reaching a target. Using video analysis we found how informed individuals might have been identified by other group members by means of inadvertent social cues (such as starting order, time spent following and group position). Furthermore, we were able to show that at least one of these cues, the group position of informed individuals, was indeed correlated with group performance. Our final experiment confirmed that leadership was even more efficient when the group members were given the identity of the leader. We discuss the effect of information status regarding the presence and identity of leaders on collective animal behaviour.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5192
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Author Baragli, P.; Mariti, C.; Petri, L.; De Giorgio, F.; Sighieri, C.
Title Does attention make the difference? Horses' response to human stimulus after 2 different training strategies Type Journal Article
Year 2011 Publication Journal of Veterinary Behavior: Clinical Applications and Research Abbreviated Journal J Vet Behav Clin Appl Res
Volume 6 Issue 1 Pages 31-38
Keywords attention; exploration; horse; human stimulus; training
Abstract We hypothesized that in an open environment, horses cope with a series of challenges in

their interactions with human beings. If the horse is not physically constrained and is free to move

in a small enclosure, it has additional options regarding its behavioral response to the trainer. The

aim of our study was to evaluate the influence of 2 different training strategies on the horse’s behavioral

response to human stimuli. In all, 12 female ponies were randomly divided into the following 2

groups: group A, wherein horses were trained in a small enclosure (where indicators of the level of

attention and behavioral response were used to modulate the training pace and the horse’s control over

its response to the stimuli provided by the trainer) and group B, wherein horses were trained in a closed

environment (in which the trainer’s actions left no room for any behavioral response except for the one

that was requested). Horses’ behavior toward the human subject and their heart rate during 2 standardized

behavioral tests were used to compare the responses of the 2 groups. Results indicated that the

horses in group A appeared to associate human actions with a positive experience, as highlighted by

the greater degree of explorative behavior toward human beings shown by these horses during the tests.

The experience of the horses during training may have resulted in different evaluations of the person, as

a consequence of the human’s actions during training; therefore, it seems that horses evaluate human

beings on daily relationship experiences.
Address attention; exploration; horse; human stimulus; training
Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1558-7878 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5286
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Author Mazurek, M.; McGee, M.; Minchin, W.; Crowe, M.A.; Earley, B.
Title Is the avoidance distance test for the assessment of animals' responsiveness to humans influenced by either the dominant or flightiest animal in the group? Type Journal Article
Year 2011 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 132 Issue 3-4 Pages 107-113
Keywords Cattle; Avoidance distance; Human-animal relationship (HAR); Dominance
Abstract A previously described (Windschnurer et al., 2009) avoidance distance test was used to assess animals’ fear of humans in order to quantify the human–animal relationship (HAR). This study investigated the influence of the dominant and flightiest animals within a group on the responsiveness of animals during the avoidance distance test. Eighty-eight pregnant heifers comprised of four different genotypes were used (22 animals per genotype): Limousin × Holstein-Friesian, Limousin × Simmental, Charolais × Limousin, and Charolais × Simmental. Sixty of the 88 heifers were group housed (n = 5) into 12 pens with 3 pens per breed, while 28 heifers were singly housed (seven heifers per breed). A reactivity test was performed on days 10, 18, 25 and 30 post-housing on the singly housed heifers, and then on the group housed heifers, on the same days, to calculate a reactivity score. On days 33 and 37 flight and dominance tests, respectively, were performed to identify the flightiest and the dominant animal within each group. On day 41, an avoidance test, measuring both the avoidance distance towards a familiar and an unfamiliar human, was performed on all heifers. No difference (P > 0.05) in reactivity scores was found between the genotypes, between pens for the group housed heifers or between singly housed and group housed heifers (P = 0.28). The avoidance distance (AD) of singly (S) housed heifers towards a familiar (F) (ADSF) human was shorter (P < 0.001) than the avoidance distance of group (G) housed heifers towards an unfamiliar human (ADSU). The ADSF and ADGF were correlated with the ADSU and ADGU (R = 0.87 for singly housed heifers; R = 0.61 for group housed heifers, P < 0.001). For the singly housed heifers, no correlation was observed between reactivity score and ADSF (R = 0.36, P = 0.18), whereas the reactivity score and ADSU were correlated (R = 0.68, P = 0.004). For the group housed heifers no significant correlation was detected between the reactivity score and ADGF (R = 0.18, P = 0.22) or ADGU (R = &#8722;0.11, P = 0.39). No influence of the most dominant animal and the flightiest animals was found on the behaviour of the group in term of avoidance distance and reactivity (P > 0.05). It is concluded that the assessment of the fear of the animals towards humans using the avoidance test at the feed bunk may be useful for singly and group housed heifers and that the leaders of a group such as the flightiest animal or the dominant animal did not influence the avoidance distance test.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5376
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Author Krishnan, A.; Gandour, J.T.; Ananthakrishnan, S.; Bidelman, G.M.; Smalt, C.J.
Title Functional ear (a)symmetry in brainstem neural activity relevant to encoding of voice pitch: A precursor for hemispheric specialization? Type Journal Article
Year Publication Brain and Language Abbreviated Journal
Volume In Press, Corrected Proof Issue Pages
Keywords Auditory; Human; Brainstem; Pitch; Language; Mandarin Chinese; Fundamental frequency-following response (FFR); Functional ear asymmetry; Experience-dependent plasticity; Subcortical
Abstract Pitch processing is lateralized to the right hemisphere; linguistic pitch is further mediated by left cortical areas. This experiment investigates whether ear asymmetries vary in brainstem representation of pitch depending on linguistic status. Brainstem frequency-following responses (FFRs) were elicited by monaural stimulation of the left and right ear of 15 native speakers of Mandarin Chinese using two synthetic speech stimuli that differ in linguistic status of tone. One represented a native lexical tone (Tone 2: T2); the other, T2', a nonnative variant in which the pitch contour was a mirror image of T2 with the same starting and ending frequencies. Two 40-ms portions of f0 contours were selected in order to compare two regions (R1, early; R2 late) differing in pitch acceleration rate and perceptual saliency. In R2, linguistic status effects revealed that T2 exhibited a larger degree of FFR rightward ear asymmetry as reflected in f0 amplitude relative to T2'. Relative to midline (ear asymmetry = 0), the only ear asymmetry reaching significance was that favoring left ear stimulation elicited by T2'. By left- and right-ear stimulation separately, FFRs elicited by T2 were larger than T2' in the right ear only. Within T2', FFRs elicited by the earlier region were larger than the later in both ears. Within T2, no significant differences in FFRS were observed between regions in either ear. Collectively, these findings support the idea that origins of cortical processing preferences for perceptually-salient portions of pitch are rooted in early, preattentive stages of processing in the brainstem.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0093-934x ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5391
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Author Birke, L.; Hockenhull, J.; Creighton, E.; Pinno, L.; Mee, J.; Mills, D.
Title Horses' responses to variation in human approach Type Journal Article
Year Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume In Press, Corrected Proof Issue Pages
Keywords Horse; Flight response; Human approach; Body posture; Approach speed; Natural horsemanship
Abstract The behaviour of humans around horses is thought to have a substantial impact on how people are perceived in subsequent interactions and many horse trainers give detailed advice on how handlers should behave when initially approaching a loose horse. Here we report on three studies designed to explore the effect of different human approach styles on the behaviour of naïve and experienced horses. In the first study, the change in flight distance (distance at which horses started to avoid an approaching human) of twelve semi-feral Dartmoor ponies, undergoing training to allow handling, was assessed. Over the 10 handling sessions median flight distance decreased significantly (p < 0.001) from 2.38 m to 0.00 m and there was a significant positive shift in the ponies' behaviour following the appearance of the researcher (p = 0.002). In a second study the effect of a direct (vigorous, swinging a lead rope and with eye contact) versus indirect (relaxed, no rope swinging and without eye contact) approach style was assessed on six adult experienced riding horses. The mean flight distance during a direct approach style (6.87 m) was significantly greater than that which occurred during an indirect approach style (2.32 m). Direction of approach was not found to significantly affect flight distance. In a third study, the effect of the rope was removed and a similar method to the second study applied to a group of naïve, feral ponies. The effect of different components of approach style, speed of approach, handler body posture and direction of gaze, which might contribute to observed differences in behavioural responses, were then examined systematically in this population. This revealed no significant difference in mean flight distance between the two approach styles (2.28 m indirect versus 2.37 m direct approach), but ponies were significantly more likely to move off in trot (p = 0.025) and to travel further (p = 0.001) when a direct approach was used. Speed of approach was the most salient factor, with a fast approach increasing both the tendency to move off in trot (p < 0.001) and distance travelled (p < 0.001). Body posture (relaxed or tense) had no effect, while flight distance was significantly greater when the person was looking away (p = 0.045). These results suggest horses may have an important egocentric spatial barrier, which perhaps relates to personal space and triggering of the flight response. Contrary to popular belief, body posture did not appear to be very important in the contexts examined unless accompanied by extraneous aids, while the speed of approach is particularly significant. These results are of important practical relevance in reducing the risk of injury, and the effective management of horses with minimal stress.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5401
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Author Sankey, C.; Richard-Yris, M.-A.; Leroy, H.; Henry, S.; Hausberger, M.
Title Positive interactions lead to lasting positive memories in horses, Equus caballus Type Journal Article
Year 2010 Publication Animal Behaviour Abbreviated Journal Anim. Behav.
Volume 79 Issue 4 Pages 869-875
Keywords Equus caballus; horse; human–animal relationship; learning; memory; positive interaction; social cognition
Abstract Social relationships are important in social species. These relationships, based on repeated interactions, define each partner's expectations during the following encounters. The creation of a relationship implies high social cognitive abilities which require that each partner is able to associate the positive or negative content of an interaction with a specific partner and to recall this association. In this study, we tested the effects of repeated interactions on the memory kept by 23 young horses about humans, after 6 and 8 months of separation. The association of a reward with a learning task in an interactional context induced positive reactions towards humans during training. It also increased contact and interest, not only just after training, but also several months later, despite no further interaction with humans. In addition, this ‘positive memory’ of humans extended to novel persons. Overall, positive reinforcement enhanced learning and memorization of the task itself. These findings suggest remarkable social cognitive abilities that can be transposed from intraspecific to interspecific social contexts.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5418
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Author Krueger, K; Farmer, K.
Title Laterality in the Horse [Lateralität beim Pferd ] Type Journal Article
Year 2011 Publication mensch & pferd international Abbreviated Journal mup
Volume 4 Issue Pages 160-167
Keywords Laterality, horse, information processing, training, welfare, human-animal interaction
Abstract Horses are one-sided, not only on a motor level, but they also prefer to use one eye, ear or nostril over the other under particular circumstances. Horses usually prefer using the left eye to observe novel objects and humans. This preference is more marked in emotional situations and when confronted with unknown persons. Thus the horse’s visual laterality provides a good option for assessing its mental state during training or in human-horse interactions. A strong preference for the left eye may signal that a horse cannot deal with certain training situations or is emotionally affected by a particular person.

Pferde benutzen für die Begutachtung von Objekten und Menschen bevorzugt eine bestimmte Nüster, ein Ohr oder ein Auge. So betrachten die meisten Pferde Objekte und Menschen mit dem linken Auge. Die Lateralitätsforschung erklärt diese sensorische Lateralität mit der Verarbeitung von Informationen unterschiedlicher Qualität in verschiedenen Gehirnhälften und zeigt auf, dass positive und negative emotionale Informationen sowie soziale Sachverhalte mit dem linken Auge aufgenommen und vorwiegend an die rechte Gehirnhälfte weitergegeben werden. In diesem Zusammenhang ermöglicht die visuelle Lateralität, den Gemütszustand des Pferdes im Training und im therapeutischen Fördereinsatz zu erkennen und zu berücksichtigen.
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Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5444
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Author Call, J.; Tomasello, M.
Title Use of social information in the problem solving of orangutans (<em>Pongo pygmaeus</em>) and human children (<em>Homo sapiens</em>) Type Journal Article
Year 1995 Publication Journal of Comparative Psychology Abbreviated Journal J. Comp. Psychol.
Volume 109 Issue 3 Pages 308-320
Keywords cross species imitative learning in problem solving, human 3–4 yr olds vs orangutans
Abstract Fourteen juvenile and adult orangutans and 24 3- and 4-yr-old children participated in 4 studies on imitative learning in a problem-solving situation. In all studies a simple to operate apparatus was used, but its internal mechanism was hidden from subjects to prevent individual learning. In the 1st study, orangutans observed a human demonstrator perform 1 of 4 actions on the apparatus and obtain a reward; they subsequently showed no signs of imitative learning. Similar results were obtained in a 2nd study in which orangutan demonstrators were used. Similar results were also obtained in a 3rd study in which a human encouraged imitation from an orangutan that had previously been taught to mimic arbitrary human actions. In a 4th study, human 3- and 4-yr-old children learned the task by means of imitation. (PsycINFO Database Record (c) 2010 APA, all rights reserved)
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Corporate Author Thesis
Publisher US: American Psychological Association Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1939-2087 (Electronic); 0735-7036 (Print) ISBN Medium
Area Expedition Conference
Notes Approved yes
Call Number Equine Behaviour @ team @ 1995-42883-001 Serial 5448
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Author Menke, C.; Waiblinger, S.; Foelsch, D.W.; Wiepkema, P.R.
Title Social Behaviour and Injuries of Horned Cows in Loose Housing Systems Type Journal Article
Year 1999 Publication Animal Welfare Abbreviated Journal Anim Welfare
Volume 8 Issue 3 Pages 243-258
Keywords Animal Welfare; Horned Dairy Cows; Human-Animal Relationship; Injuries; Loose Housing; Management; Social Behaviour
Abstract The relationship between social behaviour and skin injuries (caused by horns) of loose housed horned cows was investigated on 35 dairy farms. While the frequencies of two agonistic behaviour elements (push and chase away) were positively correlated with the occurrence of skin injuries, the frequencies of butting and homing were not. Butting appears to have an ambivalent motivation, in that its occurrence is correlated positively both with agonistic behaviour and with social licking. Horning showed a positive correlation with social licking only. Four groups of husbandry conditions that may be associated with the occurrence of social behaviour and of injuries were distinguished: i) herd management, with variables including problem solving management by the farmer, integration of new cows, and dealing with periparturient and oestrus cows; ii) human-animal relationship, with variables including ability to identify individual cows, frequency of brushing the cows, number of milkers, and frequency of personnel changes; iii) animal characteristics, with the variable of herd size; and iv) stable characteristics, with the variable of space per cow (m2). The relevance of the husbandry variables investigated here had been confirmed in a previous stepwise regression analysis (Menke 1996). The variables for herd management and human-animal relationship conditions correlated in a consistent way with the occurrence of agonistic behaviour and/or of injuries, while most of them also correlated in the opposite direction with the occurrence of social licking. Herd size correlated positively with agonistic behaviour, but negatively with social licking. Space per cow correlated negatively with agonistic behaviour and injuries. In more than 70 per cent of the herds investigated, the levels of agonistic behaviour and of skin injuries were low, implying that horned dairy cows can be kept with less risk than is often assumed. We argue that such risks strongly depend on management factors that can be improved.
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Language (up) Summary Language Original Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5480
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Author Uchiyama, H.; Ohtani, N.; Ohta, M.
Title Three-dimensional analysis of horse and human gaits in therapeutic riding Type Journal Article
Year 2011 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 135 Issue 4 Pages 271-276
Keywords Equine-facilitated interventions; Horse gait; Human gait; Three-dimensional analysis; Horse riding
Abstract Therapeutic horse riding or hippotherapy is used as an intervention for treating individuals with mental and physical disabilities. Equine-assisted interventions are based on the hypothesis that the movement of the horse's pelvis during horseback riding resembles human ambulation, and thus provides motor and sensory inputs similar to those received during human walking. However, this hypothesis has not been investigated quantitatively and qualitatively. This study aimed to verify the hypothesis by conducting a three-dimensional analysis of the horse's movements while walking and human ambulation. Using four sets of equipments, we analysed the acceleration patterns of walking in 50 healthy humans and 11 horses. In addition, we analysed the exercise intensity by comparing the heart rate, breathing rate, and blood pressure of 127 healthy individuals before and after walking and horse riding. The acceleration data series of the stride phase of horse walking were compared with those of human walking, and the frequencies (in Hz) were analysed by Fast Fourier transform. The acceleration curves of human walking overlapped with those of horse walking, with the frequency band of human walking corresponding with that of horse walking. Exercise intensity, as measured by the heart rate and breathing rate, was not significantly different between horse riding and human walking. The levels of diastolic blood pressure were slightly higher during horse riding than during walking, but were lower during both conditions compared with those in normal conditions (P < 0.01). The present study shows that, although not completely matched, the accelerations of the horse and human walking are comparable quantitatively and qualitatively. Horse riding at a walking gait could generate motor and sensory inputs similar to those produced by human walking, and thus could provide optimum benefits to persons with ambulatory difficulties.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language (up) Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5488
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