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George, I.; Cousillas, H.; Richard, J.-P.; Hausberger, M. |
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Title |
Song perception in the European starling: hemispheric specialisation and individual variations |
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2002 |
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Comptes Rendus Biologies |
Abbreviated Journal |
Compt. Rend. Biol. |
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325 |
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3 |
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197-204 |
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lateralisation; perception; birdsong; starling; electrophysiology; individual variations; latéralisation; perception; chant; étourneaux; électrophysiologie; variations individuelles |
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Abstract |
Hemispheric specialisation for speech in humans has been well documented. The lateralisation for song production observed in songbirds is reminiscent of this hemispheric dominance. In order to investigate whether song perception is also lateralised, we made multiunit recordings of the neuronal activity in the field L of starlings during the presentation of species-specific and artificial non-specific sounds. We observed a systematic stronger activation in one hemisphere than in the other one during the playback of species-specific sounds, with inter-subject variability in the predominant hemisphere for song perception. Such an asymmetry was not observed for artificial non-specific sounds. Thus, our results suggest that, at least at the individual level, the two hemispheres of the starlings' brain perceive and process conspecific signals differently. |
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Equine Behaviour @ team @ |
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4636 |
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Author |
Hanggi, E.B.; Ingersoll, J.F. |
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Title |
Lateral vision in horses: A behavioral investigation |
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2012 |
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Behavioural Processes |
Abbreviated Journal |
Behav. Process. |
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91 |
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1 |
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70-76 |
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Lateral vision; Horse; Equine; Stimulus discrimination; Field of view; Peripheral |
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This study investigated lateral vision in horses (Equus caballus) for the first time from a behavioral point of view. Three horses were tested using a novel experimental design to determine the range of their lateral and caudolateral vision with respect to stimulus detection and discrimination. Real-life stimuli were presented along a curvilinear wall in one of four different positions (A, B, C, D) and one of two height locations (Top, Bottom) on both sides of the horse. To test for stimulus detection, the correct stimulus was paired against a control; for stimulus discrimination, the correct stimulus was paired against another object. To indicate that the correct stimulus was detected or discriminated, the horses pushed one of two paddles. All horses scored significantly above chance on stimulus detection trials regardless of stimulus position or location. They also accurately discriminated between stimuli when objects appeared in positions A, B, and C for the top or bottom locations; however, they failed to discriminate these stimuli at position D. This study supports physiological descriptions of the equine eye and provides new behavioral data showing that horses can detect the appearance of objects within an almost fully encompassing circle and are able to identify objects within most but not all of their panoramic field of view. |
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0376-6357 |
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Equine Behaviour @ team @ |
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5621 |
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Lyn, H.; Savage-Rumbaugh, E.S. |
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Title |
Observational word learning in two bonobos (Pan paniscus): ostensive and non-ostensive contexts |
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2000 |
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Language & Communication |
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20 |
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3 |
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255-273 |
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Language acquisition; Bonobo; Fast mapping |
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Word learning has been extensively studied in humans. Children seem to be able to map new words onto objects with only one exposure to the referent. This ability has been called “fast mapping”(Carey, 1978 and Carey). Using a modified human paradigm, this paper explores two language-competent bonobos' (Pan paniscus) abilities to map new words to objects in realistic surroundings with few exposures to the referents. This paper also investigates the necessity of the apes maintaining visual contact (ostensive context) with the item to map the novel name onto the novel object. The bonobos tested in this experiment were able to map new words onto objects and could do so without visual contact with the items. |
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Equine Behaviour @ team @ |
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4675 |
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Dyson, S.; Berger, J.; Ellis, A.D.; Mullard, J. |
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Title |
Development of an ethogram for a pain scoring system in ridden horses and its application to determine the presence of musculoskeletal pain |
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2018 |
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Journal of Veterinary Behavior |
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23 |
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47-57 |
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Lameness; Equine behavior; Pain grading; Headshaking; Bucking; Rearing |
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There is evidence that more than 47% of the sports horse population in normal work may be lame, but the lameness is not recognized by owners or trainers. An alternative means of detecting pain may be recognition of behavioral changes in ridden horses. It has been demonstrated that there are differences in facial expressions in nonlame and lame horses. The purpose of this study was to develop a whole horse ethogram for ridden horses and to determine whether it could be applied repeatedly by 1 observer (repeatability study, 9 horses) and if, by application of a related pain behavior score, lame horses (n = 24) and nonlame horses (n = 13) could be differentiated. It was hypothesized that there would be some overlap in pain behavior scores among nonlame and lame horses; and that overall, nonlame horses would have a lower pain behavior score than lame horses. The ethogram was developed with 117 behavioral markers, and the horses were graded twice in random order by a trained specialist using video footage. Overall, there was a good correlation between the 2 assessments (P < 0.001; R2 = 0.91). Behavioral markers that were not consistent across the 2 assessments were omitted, reducing the ethogram to 70 markers. The modified ethogram was applied to video recordings of the nonlame horses and lame horses (ethogram evaluation). There was a strong correlation between 20 behavioral markers and the presence of lameness. The ethogram was subsequently simplified to 24 behavioral markers, by the amalgamation of similar behaviors which scored similarly and by omission of markers which showed unreliable results in relation to lameness. Following this, the maximum individual occurrence score for lame horses was 14 (out of 24 possible markers), with a median and mean score of 9 (±2 standard deviation) compared with a maximum score of 6 for nonlame horses, with a median and mean score of 2 (±1.4). For lame horses, the following behaviors occurred significantly more (P < 0.05, chi-square): ears back, mouth opening, tongue out, change in eye posture and expression, going above the bit, head tossing, tilting the head, unwillingness to go, crookedness, hurrying, changing gait spontaneously, poor quality canter, resisting, and stumbling and toe dragging. Recognition of these features as potential indicators of musculoskeletal pain may enable earlier recognition of lameness and avoidance of punishment-based training. Further research is necessary to verify this new ethogram for assessment of pain in ridden horses. |
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1558-7878 |
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Equine Behaviour @ team @ |
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6706 |
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Author |
Huebener, E. |
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Title |
Wie sich der pferdgerechte “selbsttätige Schenkel” besser vermitteln ließe; |
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Journal Article |
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2006 |
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Tierärztliche Umschau |
Abbreviated Journal |
Tierärztl. Umschau |
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8 |
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403-406 |
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Keywords |
Kultiviertes Reiten – Mensch-Pferd-Harmonie – feinfühlige, unsichtbare Hilfengebung – Schonen des Rückens von Reiter und Pferd |
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Von der Basis bis zum Spitzensport werden Pferde gewaltsam zum “Gehorsam” gebracht oder zur Ausführung von Übungen gezwungen. Aktionen gegen die “Rollkur” oder “Hyperflexion” füllen die Medien. Aber die Wurzel des Übels liegt viel tiefer. Die Grundlage kultivierten Reitens in hoher Harmonie zwischen Mensch und Pferd ist eine feinfühlige, nahezu unsichtbare Hilfengebung, für die Bewegungen des Pferderückens und des Pferderumpfes den Zeitgeber liefern. Das Wissen darum in der Reiterwelt zu verankern, ist noch immer nicht gelungen. |
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German |
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yes |
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refbase @ user @ |
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424 |
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Author |
Graf, P.; Schneider, T.; KönigvonBorstel, U.; Gauly M. |
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Kosten-Nutzen-Analyse einer objektivierten Temperamentbeurteilung bei Pferden [Economic evaluation of an objective temperament assessment in horses] |
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Journal Article |
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2013 |
Publication |
Züchtungskunde |
Abbreviated Journal |
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85 |
Issue |
2 |
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129-142 |
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Keywords |
Kosten, Nutzen, Interieurbeurteilung, Pferd, Temperament [Economy, personality assessment, horse, temperament] |
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Das Ziel der vorliegenden Studie war die Ermittlung der Kosten eines Verhaltenstests zur
objektiven Temperamentbeurteilung. Sie wurde an 1028 Pferden auf 55 Zuchtveranstaltungen
und Privatbetrieben ermittelt.
Weiterhin wurde eine Befragung zur allgemeinen Akzeptanz einer solchen Beurteilung
bei Reitpferden durchgeführt. Zusätzlich wurde mit Hilfe einer Online-Umfrage die
Meinung zu den Kosten und dem Aufwand einer solchen Beurteilung ermittelt. Die
Kosten der Einführung einer objektiven Temperamentbeurteilung entsprechen nach Einbeziehung
aller Faktoren ca. 18 Euro je Pferd. Den Kosten steht die Zahlungsbereitschaft
für eine verbesserte, da objektivierte Temperamentbeurteilung gegenüber. Insgesamt
56,7% der Befragten wären bereit, mehr als 11 Euro für eine objektive Interieurbeurteilung
auf Leistungsprüfungen im Feld zu investieren. Im Rahmen von Stationsprüfungen
wären sie sogar bereit mehr als 30 Euro aufzuwenden. Die Wertsteigerung eines im
Rahmen des Verfahrens positiv bewerteten Pferdes um 5%, die von den Teilnehmern der
Umfrage durchschnittlich angenommen wird, würde zusätzlich den Gewinn beim Pferdeverkauf
steigern. Die Ergebnisse zeigen, dass die Kosten einer objektiven Temperamentbeurteilung
durch eine erhöhte Zahlungsbereitschaft der Käufer scheinbar kompensiert
werden können, so dass die Einführung eines Temperamenttests zur objektiven Interieurbeurteilung
in Form der vorgestellten Untersuchungen grundsätzlich finanzierbar ist.
[The aim of the present study was to assess costs as well as riders’ acceptance of an
objective temperament evaluation in riding horses. Costs were determined based on a
novel object test conducted in 1028 horses tested on 65 occasions during performance
tests or in private stables. In addition, an online survey was used to identify riders’
opinion about the costs and benefits of such an assessment. Based on the conditions
assumed in the present study the costs for temperament testing have amount 18 Euro per
horse. More than 50% of the respondents were willing to pay more than 11 Euro for an
objective temperament assessment in their horses during performance tests in field.
Within performance tests on station they would spend more than 30 Euro for an objective temperament assessment. Participants further assumed a rise in value of favourably
assessed horses by 5%, leading to increased profits when selling the horse. In conclusion,
riders appear to be willing to cover the additional costs accrued from the temperament
test. Therefore, the introduction of an objective temperament assessment is likely to pay
off.] |
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0044-5401 |
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Equine Behaviour @ team @ |
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5866 |
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Author |
Kienapfel, K. |
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Title |
Und was meinen die Pferde dazu? – Über das Ausdrucksverhalten von Pferden bei verschiedenen Halsstellungen |
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Journal Article |
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Year |
2011 |
Publication |
Pferdeheilkunde |
Abbreviated Journal |
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27 |
Issue |
4(Juli/August) |
Pages |
372-380 |
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Keywords |
Kopf-Hals-Position, „Rollkur“, Tierschutz, Ausdrucksverhalten |
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Abstract |
Die aktuellen Diskussionen in der Reiterwelt, welche Art und Weise des Reitens, besonders welche Kopf-Hals-Stellung zu erstreben ist, werfen
die Frage auf, ob und wie die Pferde selbst ihr Befinden zum Ausdruck bringen. Über die Empfindungen der Pferde in verschiedenen
Kopfhaltungen ist bisher sehr wenig bekannt. Deswegen wurde zunächst an stehenden Pferden das Ausdrucksverhalten beobachtet. Missfallensäußerungen
häuften sich (mit 49,7% aller gezeigten Verhaltensauffälligkeiten wie Sperren, Rückwärtsgehen und Kopfschlagen) in der
aufgerollten, hyperflektierten Stellung des Halses. An zweiter Stelle folgten Unmutsäußerungen in der absolut aufgerichteten Haltung
(34,9%). Auch die beigezäumte Haltung wurde nicht ohne Unmutsbekundungen hingenommen, hier war deren Anzahl aber wesentlich
geringer (11,2%). Die hohe Kopfstellung (0,17%) und die Dehnungshaltung (0,23%) bereiteten den Tieren kaum Unbehagen. Auch das
Ausdrucksverhalten der Pferde unter dem Reiter wurde untersucht. Hierfür wurden, unter Berücksichtigung der schriftlich fixierten Regeln für
das Turnierwesen der FN, je 30 Pferde in zwei Kategorien beobachtet: mit der Stirnlinie vor der Senkrechten und Pferde mit der Stirnlinie
hinter der Senkrechten. Die Beobachtungen wurden unbemerkt von den Reitern auf den Abreiteplätzen von Turnieren durchgeführt. Die
Anzahl an Verhaltensauffälligkeiten der Pferde mit der Stirnlinie hinter der Senkrechten war deutlich (89,3 %) erhöht im Gegensatz zu der
anderen Gruppe (10,7 %). Die Pferde mit der Stirnlinie hinter der Senkrechten zeigten 8 Mal mehr Unmutsäußerungen als die mit der Stirnlinie
vor der Senkrechten. Entgegen den Regeln der FN zeigten die durchgeführten Scans, dass unmittelbar vor den Prüfungen auf Turnieren
92,8% der Pferde mit der Stirnnasen-Linie hinter der Senkrechten geritten wurden. Ein Befund dieser Studie ist die Feststellung, dass die
Praxis deutlich von den Regeln abweicht. Das Reiten mit der Stirnlinie hinter der Senkrechten ist nach diesen Befunden abzulehnen, da die
Pferde deutliches Unwohlsein in dieser Haltung signalisieren. |
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Equine Behaviour @ team @ |
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5957 |
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Author |
Gardner, A., West, S. A. |
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Title |
Cooperation and Punishment, Especially in Humans |
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Journal Article |
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Year |
2004 |
Publication |
The American Naturalist |
Abbreviated Journal |
Americ. Natur. |
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Volume |
164 |
Issue |
6 |
Pages |
753-764 |
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Keywords |
kin selection, neighbor-modulated fitness, repression of |
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Abstract |
Explaining altruistic cooperation is one of the greatest
challenges faced by sociologists, economists, and evolutionary biologists.
The problem is determining why an individual would carry
out a costly behavior that benefits another. Possible solutions to this
problem include kinship, repeated interactions, and policing. Another
solution that has recently received much attention is the threat
of punishment. However, punishing behavior is often costly for the
punisher, and so it is not immediately clear how costly punishment
could evolve. We use a direct (neighbor-modulated) fitness approach
to analyze when punishment is favored. This methodology reveals
that, contrary to previous suggestions, relatedness between interacting
individuals is not crucial to explaining cooperation through punishment.
In fact, increasing relatedness directly disfavors punishing
behavior. Instead, the crucial factor is a positive correlation between
the punishment strategy of an individual and the cooperation it
receives. This could arise in several ways, such as when facultative
adjustment of behavior leads individuals to cooperate more when
interacting with individuals who are more likely to punish. More
generally, our results provide a clear example of how the fundamental
factor driving the evolution of social traits is a correlation between
social partners and how this can arise for reasons other than genealogical
kinship. |
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University of Edinburgh, West Mains Road, Edinburgh EH9 3JT, |
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no |
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refbase @ user @ |
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341 |
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Author |
Asa, C.S. |
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Title |
Male reproductive success in free-ranging feral horses |
Type |
Journal Article |
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Year |
1999 |
Publication |
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Abbreviated Journal |
Behavioral Ecology and Sociobiology |
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Volume |
47 |
Issue |
1-2 |
Pages |
89-93 |
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Keywords |
Key words Reproductive success; Alternative mating strategies; Mating systems; Paternity |
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Abstract |
In the social organization of feral horses, adult males compete to monopolize groups or bands of females, sometimes called harems. Alternative male strategies are to remain alone or with other bachelors or, less commonly, to accept subordinate status within a harem. The hypothesis that dominant harem stallion status confers a reproductive advantage was tested in free-ranging feral horses. The presence of foals in harems headed by vasectomized (VSX) versus intact stallions was used to assess the ability of these stallions to control reproduction in their harems. Of harems headed by VSX stallions, 17 and 33% contained foals during years 2 and 3 post-treatment, respectively. In contrast, 86 and 80% of harems headed by non-VSX stallions contained foals in those years. Acquisition of pregnant mares appeared more likely than sneak copulations by bachelor stallions to account for foals in harems with a single stallion. However, most foals were born into harems that included a subordinate stallion, an occurrence that was undoubtedly exacerbated by the extended breeding season resulting from the sterility of the harem stallion. Thus, in comparing alternative reproductive tactics, bachelors appeared less successful than subordinate stallions within a harem. However, the highest reproductive success was achieved by the harem stallion, further demonstrating that alternative tactics are not equally profitable. |
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Springer-Verlag |
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English |
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0340-5443 |
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no |
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Equine Behaviour @ team @ |
Serial |
5786 |
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Author |
Barton, R.A.; Byrne, R.W.; Whiten, A. |
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Title |
Ecology, feeding competition and social structure in baboons |
Type |
Journal Article |
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Year |
1996 |
Publication |
Behavioral Ecology and Sociobiology |
Abbreviated Journal |
Behav. Ecol. Sociobiol. |
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Volume |
38 |
Issue |
5 |
Pages |
321-329 |
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Keywords |
Key words Ecology – Competition – Group size – Baboons |
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Abstract |
Predictions of the model of van Schaik (1989) of female-bonding in primates are tested by systematically comparing the ecology, level of within-group contest competition for food (WGC), and patterns of social behaviour found in two contrasting baboon populations. Significant differences were found in food distribution (percentage of the diet from clumped sources), feeding supplant rates and grooming patterns. In accord with the model, the tendencies of females to affiliate and form coalitions with one another, and to be philopatric, were strongest where ecological conditions promoted WGC. Group fission in the population with strong WGC was “horizontal” with respect to female dominance rank, and associated with female-female aggression during a period of elevated feeding competition. In contrast, where WGC was low, females' grooming was focused on adult males rather than other females. Recent evidence suggests that group fission here is initiated by males, tends to result in the formation of one-male groups, and is not related to feeding competition but to male-male competition for mates. An ecological model of baboon social structure is presented which incorporates the effects of female-female competition, male-male competition, and predation pressure. The model potentially accounts for wide variability in group size, group structure and social relationships within the genus Papio. Socio-ecological convergence between common baboons and hamadryas baboons, however, may be limited in some respects by phylogenetic inertia. |
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