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	Author	Hemelrijk,C. K.; Wantia,J.; Gygax,L.
	Title	The construction of dominance order: comparing performance of five methods using an individual-based model			Type	Journal Article
	Year	2005	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	142	Issue	8	Pages	1043-1064
	Keywords	dominance order, ranking method, agent-based model, statistical method, aggression
	Abstract	In studies of animal behaviour investigators correlate dominance with all kinds of behavioural variables, such as reproductive success and foraging success. Many methods are used to produce a dominance hierarchy from a matrix reflecting the frequency of winning dominance interactions. These different methods produce different hierarchies. However, it is difficult to decide which ranking method is best. In this paper, we offer a new procedure for this decision: we use an individual-based model, called DomWorld, as a test-environment. We choose this model, because it provides access to both the internal dominance values of artificial agents (which reflects their fighting power) and the matrix of winning and losing among them and, in addition, because its behavioural rules are biologically inspired and its group-level patterns resemble those of real primates. We compare statistically the dominance hierarchy based on the internal dominance values of the artificial agents with the dominance hierarchy produced by ranking individuals by (a) their total frequency of winning, (b) their average dominance index, (c) a refined dominance index, the David`s score, (d) the number of subordinates each individual has and (e) a ranking method based on maximizing the linear order of the hierarchy. Because dominance hierarchies may differ depending on group size, type of society, and the interval of study, we compare these ranking methods for these conditions.We study complete samples as well as samples randomly chosen to resemble the limitations of observing real animals. It appears that two methods of medium complexity (the average dominance index and David`s score) lead to hierarchical orders that come closest to the hierarchy based on internal dominance values of the agents. We advocate usage of the average dominance index, because of its computational simplicity.
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	Call Number	refbase @ user @			Serial	445
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	Author	Hewitt, S.E.; Macdonald, D.W.; Dugdale, H.L.
	Title	Context-dependent linear dominance hierarchies in social groups of European badgers, Meles meles			Type	Journal Article
	Year	2009	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	77	Issue	1	Pages	161-169
	Keywords	David's score; directed aggression; dominance rank; European badger; grooming behaviour; linear dominance hierarchy; Meles meles; reproductive success; relatedness; social status
	Abstract	A social hierarchy is generally assumed to exist in those mammalian societies in which the costs and benefits of group living are distributed unevenly among group members. We analysed infrared closed-circuit television footage, collected over 3 years in Wytham Woods, Oxfordshire, to test whether social groups of European badgers have dominance hierarchies. Analysis of directed aggression between dyads revealed linear dominance hierarchies in three social-group-years, but patterns within social groups were not consistent across years. Dominance hierarchies were significantly steeper than random in five out of six social-group-years. In those social-group-years where a linear hierarchy was determined, there was an effect of sex on dominance rank, with females gaining significantly higher rank than males in two social-group-years. Overall, rank was not related to age, nor did it appear to affect the likelihood of an individual being wounded, or an individual's breeding status. The latter resulted from nonorthogonality between sex and breeding status, as there were only two breeding males. Overall, hierarchies were primarily dominated by breeding females, and may occur when breeding competition arises. Relatedness, unreciprocated allogrooming and sequential allomarking were not consistently related to levels of directed aggression across social-group-years. We suggest that dominance structures within European badger groups may be context dependent, with future study required to complete our understanding of where, and when, they arise.
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	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4695
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	Author	Koolhaas, J.M.; Korte, S.M.; De Boer, S.F.; Van Der Vegt, B.J.; Van Reenen, C.G.; Hopster, H.; De Jong, I.C.; Ruis, M.A.W.; Blokhuis, H.J.
	Title	Coping styles in animals: current status in behavior and stress-physiology			Type	Journal Article
	Year	1999	Publication	Neuroscience & Biobehavioral Reviews	Abbreviated Journal
	Volume	23	Issue	7	Pages	925-935
	Keywords	Coping; Aggression; Stress; Disease; Corticosterone
	Abstract	This paper summarizes the current views on coping styles as a useful concept in understanding individual adaptive capacity and vulnerability to stress-related disease. Studies in feral populations indicate the existence of a proactive and a reactive coping style. These coping styles seem to play a role in the population ecology of the species. Despite domestication, genetic selection and inbreeding, the same coping styles can, to some extent, also be observed in laboratory and farm animals. Coping styles are characterized by consistent behavioral and neuroendocrine characteristics, some of which seem to be causally linked to each other. Evidence is accumulating that the two coping styles might explain a differential vulnerability to stress mediated disease due to the differential adaptive value of the two coping styles and the accompanying neuroendocrine differentiation.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4416
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	Author	Feuerstein, N.; Terkel, J.
	Title	Interrelationships of dogs (Canis familiaris) and cats (Felis catus L.) living under the same roof			Type	Journal Article
	Year	2008	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	113	Issue	1-3	Pages	150-165
	Keywords	Cats; Dogs; Shared home; Interrelationship; Aggression; Amicability; Indifference; Adaptation
	Abstract	In the process of domestication, dogs (Canis familiaris) and cats (Felis catus) have undergone thousands of years of genetic changes that have adapted them to the human environment. Both species have acquired a global distribution and it has become quite common to find homes with the two living side by side. Nevertheless, there is widespread belief that interspecific communication between dogs and cats is problematic, stemming from their separate evolutionary development and different social structures. Consequently, many people considering possible adoption of both species are concerned about their ability to get along. Interrelationships of dogs and cats living together were studied here in an attempt to determine the main factors influencing the type of relationship likely to develop between the two species. Two approaches were used: (1) a questionnaire completed by owners of both dog(s) and cat(s), which provided a broad database of the animals' behaviors; and (2) observations carried out in participants' homes on their dog-cat interactions. Two separate ethograms for dogs and cats served for analyses of their body language. The findings revealed the following: Both species showed a similar ability to establish a relatively amicable relationship with the other species; the animals' gender had little influence on the nature of their interrelationship; and adoption of the cat prior to the dog appears to conduce to establishing an amicable relationship, as does their first encounter taking place at an early age (up to 6 months of age in cats and up to 1 year in dogs). The findings also suggest that the majority of these dogs and cats understood the particular body language displayed by one animal that has an opposite meaning for the other species; and that the earlier the age of first encounter between the two, the better this understanding. It can be concluded that exposure of both species at an early age to the presence of the other facilitates the learning of each other's body language, and the consequent establishment of an amicable relationship. A better understanding of the various factors that contribute to determining the two species' relationship should not only improve the quality of life of these pets, but also reassure and encourage more people to adopt both cat and dog.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4757
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	Author	Barber, J.A.; Crowell-Davis, S.L.
	Title	Maternal behavior of Belgian (Equus caballus) mares			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	41	Issue	3-4	Pages	161-189
	Keywords	Belgian; Draft horse; Ontogeny; Maternal aggression; Maternal behavior; Nursing behavior; Recumbency response
	Abstract	The relationship between ten Belgian mares and their offspring was studied from the first day of foal life to 17 weeks of age. Mares and foals spent more time at greater distances from each other as foals matured. Mares exhibited the recumbency response, being in closer proximity to their foals when foals were recumbent than when they were upright. Foals initiated the majority of nursing bouts. Frequency and duration of nursing bouts and percentage of time resting recumbently declined as foals matured. Foals also terminated the their foals, and they were most likely to do so in the first month of foal life. Maternal initiation of nursing. There was usually no discernible foal response to maternal aggression. Little difference between maternal behavior directed towards colts and fillies was found for all aspects of the study. Maternal behavior in the Belgian draft horse was similar to that reported for other equid breeds.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2268
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	Author	Austin, N.P.; Rogers, L.J.
	Title	Lateralization of agonistic and vigilance responses in Przewalski horses (Equus przewalskii)			Type	Journal Article
	Year	2014	Publication	Applied Animal Behaviour Science	Abbreviated Journal
	Volume	151	Issue		Pages	43-50
	Keywords	Behavioural lateralization: Eye preference; Limb preference; Aggression; Vigilance; Reactivity; Przewalski horses
	Abstract	tEye and limb preferences were scored in the closest undomesticated relative of Equuscaballus using the same methods as used previously to study laterality in feral horses.Observations were made of 33 Przewalski horses (Equus ferus przewalskii) (male N = 20,female N = 13) living under natural social conditions on a large reserve in France. Signifi-cant left-eye/side biases were found in agonistic interactions within harem bands (M ± SEbias to left 58% ± 0.01 for threats, P < 0.001; 68% ± 0.05 for attacks; P < 0.001) and in stallionfights (threats, 52% ± 0.01 left, P < 0.001; attacks, 63% ± 0.02 left, P < 0.001): as many as 80%of the horses were significantly lateralized in attack responses within harem bands. Lat-erality of vigilance was measured as lifting up the head from grazing and turning it to theleft or right side: a directional bias to the left was found (M ± SE 53% ± 0.02 left, P < 0.001).Side bias in reactivity was calculated as the percent of head lifts above the level of thewithers on the left or right side and this was also left side biased (M ± SE 73% ± 0.03 left,P < 0.001). These results indicate right-hemisphere specialization for control of aggressionand responses to novelty. The left bias in attack scores within harem bands was strongerin males than females (P = 0.024) and in immature than adult horses (P = 0.032). Immaturehorses were also more strongly lateralized than adults in vigilance scores (P = 0.022), whichmay suggest that experience reduces these side biases. Our results show that Przewalskihorses exhibit left eye preferences, as do feral horses, and do so even more strongly thanferal horses. Considering feral and Przewalski horses together, we deduce that ancestralhorses had similar lateral biases. Also similar to feral horses, the Przewalski horses showedno significant forelimb preference at the group level or in the majority of horses at theindividual level, confirming the hypothesis that previously reported limb preferences indomestic breeds are entrained or generated by breed-specific selection.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
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	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5768
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	Author	Cloutier, S.; Newberry, R.C.; Honda, K.
	Title	Comparison of social ranks based on worm-running and aggressive behaviour in young domestic fowl			Type	Journal Article
	Year	2004	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	65	Issue	1	Pages	79-86
	Keywords	Aggression; Social behaviour; Dominance; Play; Chickens; Animal welfare
	Abstract	Worm-running is behaviour in which a chick runs carrying a worm-like object while flock mates follow and attempt to grab the object from its beak. We hypothesised that social ranks based on worm-running frequency are stable over time and are positively correlated with social ranks based on success in aggressive interactions when older. At 8-12 days of age, we scored worm-running in 17 groups of 12 female White Leghorn chicks during three 10-min tests. Based on instantaneous scans at 5-s intervals, the bird carrying the `worm' most often was placed in rank one and so on down the rank order. These tests were repeated at 68-70 days of age. An aggression index for each bird was calculated as the number of aggressive acts given, divided by the number given and received, during three 1-h observation periods when the birds were 68-70 days. Ranks obtained in worm-running tests were positively correlated over the two age periods (P<0.05) but were not correlated with ranks based on the aggression index (P>0.05). Our results indicate that worm-running ranks are not predictive of success in aggressive interactions. Instead, worm-running fits some criteria for play.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number				Serial	2090
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	Author	Araba, B.D.; Crowell-Davis, S.L.
	Title	Dominance relationships and aggression of foals (Equus caballus)			Type	Journal Article
	Year	1994	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	41	Issue	1-2	Pages	1-25
	Keywords	aggression; dominance; horse; Equus caballus
	Abstract	Studied a herd of 15 Belgian brood-mares and 10 foals. Specific aspects of social structure studied were dominance-subordinance relationships, preferred associates, social spacing, aggression rates, the frequency of aggressions administered down the dominance hierarchy, and interactive play bouts. The rank order of the foals, both before and after weaning, was positively correlated with the rank order of their dams. There was also a significant relationship between a foal's rank and its total aggression or aggression rate per subordinate post-weaning. Higher ranking foals had higher rates of aggression. Over 80% of threats were directed down the dominance hierachy. The play-rank order of the foals, scored by the number of times foal left a play bout, was not significantly correlated with the rank order as scored by agonistic interactions. -from Authors
	Address	Dept Anatomy and Radiology, Univ of Georgia, Athens, GA, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	01681591 (Issn)	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	790
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	Author	Jennings, D.J.
	Title	Limited evidence that visual lateralization is associated with fitness in rutting male fallow deer			Type	Journal Article
	Year	2014	Publication	Animal Behaviour	Abbreviated Journal	Anim. Behav.
	Volume	92	Issue		Pages	85-91
	Keywords	aggression; contest behaviour; Dama dama; fallow deer; lateral display; lateralization; mating success; third-party intervention behaviour
	Abstract	Under certain models of animal competition, individuals are expected to gather information about opponent quality in order to determine whether they should fight or withdraw. However, the ability to process complex information differs between individuals and across brain hemispheres: a feature of vertebrate cognition known as lateralization that is not anticipated by contest models. I investigated the relationship between aggressive behaviour and mating success during the fallow deer, Dama dama, rut and a measure of lateralization derived from eye preference during parallel walking. Results show that there was no relationship between the tendency to escalate to fighting or predictability in the tendency to engage in fighting and lateralization. Conversely, there was a quadratic relationship between third-party intervention behaviour and lateralization: the greater the tendency to intervene in ongoing fights the lower the degree of lateralization. However, individuals that showed lateralization for right-eye use were least likely to be targeted by the intervening male; thus lateralization is beneficial in this context because targeted males are highly likely to lose this subsequent encounter. The relationship between lateralization and mating success was also nonlinear: males that showed little evidence for an eye bias during lateral displays had the greatest mating success. Taken together, individuals that showed lateralization benefited from avoiding being targeted after third-party intervention; conversely, individuals that showed little evidence for lateralization actively intervened during ongoing fights and had higher mating success. These results suggest that, although lateralization does appear to confer a fitness advantage on individuals, this is not as extensive as anticipated.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5802
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	Author	Jonart, L.M.; Hill, G.E.; Badyaev, A.V.
	Title	Fighting ability and motivation: determinants of dominance and contest strategies in females of a passerine bird			Type	Journal Article
	Year	2007	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	74	Issue	6	Pages	1675-1681
	Keywords	aggression; Carpodacus mexicanus; communication; house finch; passerines; resource holding potential
	Abstract	The communication of aggressive motivation or fighting ability has important fitness consequences for competing animals. Selection should favour rapid and honest communication between opponents to settle dominance relationships while avoiding prolonged and intense fighting. We investigated factors that influence fighting strategies and contest outcomes in female house finches, Carpodacus mexicanus, specifically focusing on the following questions. (1) What social contexts trigger an aggressive response? (2) Does body size and condition contribute to female fighting ability? (3) Do contextual factors, such as mate presence, nest status, nest proximity, and site experience contribute to fighting motivation? (4) Does contest intensity and duration increase as the differences in fighting ability between opponents decrease? (5) What is the relative contribution of fighting ability and aggressive motivation to the outcome of a contest? We found that aggression was triggered most frequently by female intrusions in the vicinity of nest and by extrapair female intrusions on an established pair. Female fighting and contest outcomes were strongly influenced by body condition and body size, and females were more motivated to initiate fights and won more contests when their mates were present. Females at the later breeding stages and those fighting closer to their nests were dominant and won more fights compared to females at earlier breeding stages or further from their nests. Females initiated a greater proportion of contests against opponents with similar local familiarity and breeding history. Escalated and prolonged contests, while rare, occurred exclusively between females of the most similar body size and condition. When differences in body condition between opponents are not easily perceived, contestants might escalate contests for more reliable assessments of relative fighting ability. Finally, body condition was not a strong determinant of contest outcome in the contexts with easily assessed differences in the resource value (e.g. mate presence), but without these motivational differences, body condition was the ultimate determinant of contest outcomes.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4317
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