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	Author	Moehlman, P.D.
	Title	*Behavioral patterns and communication in feral asses (Equus africanus)*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	125-169
	Keywords	Equids; Feral asses; Behavior patterns; Facial expressions; Postures; Locomotion
	Abstract	The behavior of feral populations of the African wild ass (Equus africanus) were studied in the Northern Panamint Range of Death Valley National Monument for 20 months from 1970 to 1973 [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Natl. Geogr. Soc. Res. Reports, 1970: 405-411]. Maintenance behavior is described and behavior sequences that were used in social interactions are quantified by sex and age class. Agonistic, sexual, and greeting behavior patterns are described and analyzed in conjunction with the responses they elicited. Mutual grooming mainly occurred between adult males, and between females and their offspring. Five types of vocalizations were distinguished: brays, grunts, growls, snorts, and whuffles. A second population was studied for 1 month on Ossabaw Island, GA (Moehlman, 1979). This population had more permanent social groups and had a higher rate of mutual grooming and foal social play.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2381
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	Author	Moehlman, P.D.
	Title	*Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	171-195
	Keywords	Equids; Feral asses; Social organization; Mating systems; Intraspecific variation
	Abstract	Feral asses have been studied in the arid habitats of the southwestern United States [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Reports 1970, 405-411.; Woodward, S.L., 1979. The social system of feral asses (Equus asinus). Z. Tierpsychol. 49, 304-316] and in the mesic habitat of Ossabaw Island, Georgia [Moehlman, P.D., 1979, ibid; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia. PhD dissertation, Pennsylvania State University, University Park, 219 pp]. The feral ass populations in these two locales exhibited intraspecific variation in polygynous mating systems and social organization which were consistent with the ecological classification of mating systems of Emlen and Oring (1977) [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), 215-223]. Feral asses in the arid environment have a `resource defense' polygynous mating system, and those in the mesic habitat exhibit `female (harem) defense' polygyny. The intraspecific variation observed in feral asses encompasses the interspecific variation observed in the family Equidae.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2382
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	Author	Moehlman, P.D.; Fowler, L.E.; Roe, J.H.
	Title	*Feral asses (Equus africanus) of Volcano Alcedo, Galapagos: behavioral ecology, spatial distribution, and social organization*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	197-210
	Keywords	Equids; Feral asses; Social organization; Mating systems; Intraspecific variation; Galapagos
	Abstract	Feral asses were studied on Volcano Alcedo, Galapagos Islands, Ecuador, during the wet season of 1980. On the volcano rim during March/April, two stable groups were observed to have a `female (harem) defense' polygynous mating system [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), pp. 215-223] and social behavior patterns and feeding ecology similar to feral asses living in a habitat where forage and climate are similar, e.g., Ossabaw Island, Georgia [Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Rep., 1970, pp. 405-411; Moehlman, P.D., 1997. Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats. J. Appl. Anim. Behav. Sci., this issue; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia., PhD Dissertation, Pennsylvania State University, University Park, 219 pp.].
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2383
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	Author	Boyd, L.; Bandi, N.
	Title	Reintroduction of takhi, Equus ferus przewalskii, to Hustai National Park, Mongolia: time budget and synchrony of activity pre- and post-release			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	78	Issue	2-4	Pages	87-102
	Keywords	Equids; Przewalski's horses; Takhi; Behaviour patterns; Time budgets; Behavioural synchrony; Reintroduction
	Abstract	A harem of takhi (Equus ferus przewalskii) was observed during introduction to the Hustain Nuruu Steppe Reserve of Mongolia. The goals were to examine whether the harem exhibited significant behavioural synchrony, whether release had an effect on time budget, and what the implications might be regarding acclimatisation to the wild. Behaviours were scan sampled every 10 min between the hours of 06:00 and 22:00, twice before release, twice immediately after release, and twice 2 years after reintroduction. Time budgets were constructed from these data. Considerable behavioural synchrony was evidenced both before and after release. Crepuscular grazing and midday resting were typical, regardless of the date relative to release. Upon release, the amount of time spent moving doubled for all age classes. It is suggested that this increase resulted from exploration. The amount of time spent grazing and standing remained unchanged; the increased amount of time spent moving came at the expense of resting. Two years later, the horses still spent more time moving than when captive. Somewhat less time was spent grazing, although the difference was not significant. More time was spent resting in 1996 than immediately after release. These time budgets provide evidence of successful acclimatisation to the wild. Trekking between favoured sites could account for the persistent increase in time spent moving, with concomitantly less time needed to meet nutritional needs by grazing and more time available for resting. Housing captive takhi in large enclosures is evidently insufficient to permit the amount of movement exhibited by this wild harem. The time budget of the 1- and 2-year olds was more similar to that of adults than foals, indicating approaching adulthood. That 1- and 2-year olds were nursed, without loss of body condition by the dam, provided additional evidence that the takhi achieved excellent nutritional status in the wild.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3690
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	Author	Burden, F.; Trawford, A.
	Title	Equine interspecies aggression Comment on			Type
	Year	2006	Publication	The Veterinary record	Abbreviated Journal	Vet. Rec.
	Volume	159	Issue	25	Pages	859-860
	Keywords	Aggression; Animals; Behavior, Animal; Cats; Dogs; Equidae
	Abstract
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	Language	English	Summary Language		Original Title
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	ISSN	0042-4900	ISBN		Medium
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	Notes	PMID:17172484			Approved	no
	Call Number				Serial	1777
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	Author	Moehlman, P.D.
	Title	Endangered wild equids			Type	Journal Article
	Year	2005	Publication	Scientific American	Abbreviated Journal	Sci Am
	Volume	292	Issue	3	Pages	74-81
	Keywords	Africa; Americas; Animals; Animals, Wild/physiology; Asia; Conservation of Energy Resources; Environment; *Equidae/physiology; Food Chain; Humans; Male; Reproduction
	Abstract
	Address	IUCN-The World Conservation Union/Species Survival Commission Equid Specialist Group
	Corporate Author				Thesis
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	Language	English	Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN	0036-8733	ISBN		Medium
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	Notes	PMID:15859216			Approved	no
	Call Number	refbase @ user @			Serial	551
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	Author	McDonnell, S.M.; Haviland, J.C.S.
	Title	Agonistic ethogram of the equid bachelor band			Type	Journal Article
	Year	1995	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	43	Issue	3	Pages	147-188
	Keywords	Equid; Horse; Bachelor; Agonistic; Social behavior; Ethogram
	Abstract	An ethogram of agonistic and related behaviors among equid bachelor band members was developed. Several key English-language studies on equids were reviewed to derive a preliminary inventory of specific behaviors to be included in the ethogram. A bachelor band of domestic pony stallions pastured together was observed for approximately 50 daylight hours to obtain detailed descriptions of each behavior, enable photographic and video documentation of behaviors, and identify any behaviors to be added to the preliminary inventory. An initial draft of the ethogram was sent to 65 equine researchers for review. Twenty-eight critical reviews were received and their suggestions considered for the final draft. A total of 49 elemental behaviors including five distinct vocalizations was included in the ethogram. Three complex behavioral sequences were also included. Most of the behaviors catalogued from the direct observation of pastured pony stallions were also found in the equid literature. For many, references to these behaviors specifically among males or bachelor band members were not found. The results offer a practical tool for quantitative research and other studies of equid inter-male behavior as well as for teaching of equid behavior, and should facilitate progress toward development of a complete ethogram for the horse and other equids.
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	Call Number	refbase @ user @ ; Equine Behaviour @ team @ room B 3.029			Serial	749
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	Author	Pritchard, J.C.; Lindberg, A.C.; Main, D.C.J.; Whay, H.R.
	Title	Assessment of the welfare of working horses, mules and donkeys, using health and behaviour parameters			Type	Journal Article
	Year	2005	Publication	Preventive Veterinary Medicine	Abbreviated Journal	Prev Vet Med
	Volume	69	Issue	3-4	Pages	265-283
	Keywords	Animal Welfare; Animals; Behavior, Animal; Developing Countries; Equidae; Female; Health; Male
	Abstract	Working animals provide an essential transport resource in developing countries worldwide. Many of these animals are owned by poor people and work in harsh environments, so their welfare is a cause for concern. A protocol was developed to assess the welfare of working horses, mules and donkeys in urban and peri-urban areas, using direct observation of health and behaviour parameters. In this study, 4903 animals used for draught, pack and ridden work in Afghanistan, Egypt, India, Jordan and Pakistan were assessed between December 2002 and April 2003. The data showed that donkeys were more likely than mules or horses to demonstrate avoidance or aggressive behaviour towards an observer, while horses were most likely to make a friendly approach. Fewer than 8% of working equines had abnormal mucous membranes, ectoparasites or poor coat condition. Body lesions occurred predominantly in the areas of the breast/shoulder, withers and girth in all three species, with mules having the highest prevalence of lesions in these areas (22.5, 21.3 and 28.4%, respectively). Among horses and donkeys, the prevalence of these lesions was influenced by the type of work carried out. Lesions on the head, neck, ribs, flank and tail base were seen in less than 10% of animals. Across all three species approximately 70% of animals were thin, having a body condition score (BCS) of 2 or less on a scale of 1-5 (1, very thin; 5, very fat) and more horses were in very thin condition (BCS 1) than mules or donkeys. Over 75% of animals demonstrated limb deformities and abnormalities of gait. The results of this study are being used as the initial stage of a long-term strategy to inform priorities for welfare interventions in working equines and to establish a welfare benchmark. Subsequent stages will rank the welfare concerns identified, assess the contributing risk factors and implement specific interventions to address these risks. Following intervention, success in improving welfare will be measured by repetition of this protocol and comparison with the benchmark.
	Address	Department of Clinical Veterinary Science, University of Bristol, Langford, Bristol BS40 5DU, UK. joy.pritchard@bristol.ac.uk
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	Language	English	Summary Language		Original Title
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	ISSN	0167-5877	ISBN		Medium
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	Notes	PMID:15907574			Approved	no
	Call Number				Serial	1889
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	Author	Cameron, E.Z.; Linklater, W.L.; Stafford, K.J.; Minot, E.O.
	Title	Aging and improving reproductive success in horses: declining residual reproductive value or just older and wiser?			Type	Journal Article
	Year	2000	Publication	Behavioral Ecology and Sociobiology	Abbreviated Journal	Behav. Ecol. Sociobiol.
	Volume	47	Issue	4	Pages	243-249
	Keywords	Maternal investment – Equidae – Equus caballus
	Abstract	In many mammalian species, female success in raising offspring improves as they age. The residual reproductive value hypothesis predicts that each individual offspring will be more valuable to the mother as she ages because there is less conflict between the current and potential future offspring. Therefore, as mothers age, their investment into individual offspring should increase. Empirical evidence for an influence of declining residual reproductive value on maternal investment is unconvincing. Older mothers may not invest more, but may be more successful due to greater experience, allowing them to target their investment more appropriately (targeted reproductive effort hypothesis). Most studies do not preclude either hypothesis. Mare age significantly influenced maternal investment in feral horses living on the North Island of New Zealand. Older mares, that were more successful at raising foals, were more protective for the first 20 days of life, but less diligent thereafter. Total maternal input by older mothers did not seem to be any greater, but was better targeted at the most critical period for foal survival and a similar pattern was observed in mares that had lost a foal in the previous year. In addition, older mothers were more likely to foal in consecutive years, supporting the hypothesis that they are investing less than younger mares in individual offspring. Therefore, older mothers seem to become more successful by targeting their investment better due to experience, not by investing more in their offspring.
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	Notes				Approved	no
	Call Number				Serial	2019
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	Author	Blunden, A.S.; Smith, K.C.; Whitwell, K.E.; Dunn, K.A.
	Title	Systemic infection by equid herpesvirus-1 in a Grevy's zebra stallion (Equus grevyi) with particular reference to genital pathology			Type	Journal Article
	Year	1998	Publication	Journal of Comparative Pathology	Abbreviated Journal	J Comp Pathol
	Volume	119	Issue	4	Pages	485-493
	Keywords	Animals; Animals, Zoo; Epididymis/pathology/virology; Equidae/virology; Herpesviridae Infections/diagnosis/pathology/veterinary; Herpesvirus 1, Equid/isolation & purification/pathogenicity; Lymph Nodes/pathology/virology; Male; Nasal Mucosa/pathology/virology; Pulmonary Edema/pathology; Spleen/virology; Testis/pathology/virology
	Abstract	A severe multi-systemic form of equid herpesvirus-1 infection is described in an adult zebra stallion. There was multifocal necrotizing rhinitis, marked hydrothorax and pulmonary oedema, with viral antigen expression in degenerating epithelial cells, local endothelial cells and intravascular leucocytes of the nasal mucosa and lung. Specific localization of EHV-1 infection was seen in the testes and epididymides, including infection of Leydig cells and germinal epithelium, which would have facilitated venereal shedding of virus in life. The case provided a unique opportunity to study hitherto undescribed aspects of the pathogenesis of naturally occurring EHV-1 infection in the male equine genital tract. Restriction digests of the isolate demonstrated a pattern similar to that of EHV-1 isolates previously recovered from aborted zebra and onager fetuses.
	Address	Animal Health Trust Centre for Preventive Medicine, Newmarket, Suffolk, UK
	Corporate Author				Thesis
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	Language	English	Summary Language		Original Title
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	ISSN	0021-9975	ISBN		Medium
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	Notes	PMID:9839210			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2239
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