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	Author	Devienne, M.F.; Guezennec, C.Y.
	Title	Energy expenditure of horse riding			Type	Journal Article
	Year	2000	Publication	European Journal of Applied Physiology	Abbreviated Journal	Eur J Appl Physiol
	Volume	82	Issue	5-6	Pages	499-503
	Keywords	Adult; Animals; Energy Metabolism/physiology; Exercise/physiology; Exercise Test; Female; Hemodynamic Processes/physiology; Horses; Humans; Male; Oxygen Consumption/physiology; Respiratory Mechanics/physiology
	Abstract	Oxygen consumption (VO2), ventilation (VE) and heart rate (HR) were studied in five recreational riders with a portable oxygen analyser (K2 Cosmed, Rome) telemetric system, during two different experimental riding sessions. The first one was a dressage session in which the rider successively rode four different horses at a walk, trot and canter. The second one was a jumping training session. Each rider rode two horses, one known and one unknown. The physiological parameters were measured during warm up at a canter in suspension and when jumping an isolated obstacle at a trot and canter. This session was concluded by a jumping course with 12 obstacles. The data show a progressive increase in VO2 during the dressage session from a mean value of 0.70 (0.18) l x min(-1) [mean (SD)] at a walk, to 1.47 (0.28) l x min(-1) at a trot, and 1.9 (0.3) l x min(-1) at a canter. During the jumping session, rider VO2 was 2 (0.33) l x min(-1) with a mean HR of 155 beats x min(-1) during canter in suspension, obstacle trot and obstacle canter. The jumping course significantly enhanced VO2 and HR up to mean values of 2.40 (0.35) l x min(-1) and 176 beats x min(-1), respectively. The comparison among horses and riders during the dressage session shows differences in energy expenditure according to the horse for the same rider and between riders. During the jumping session, there was no statistical difference between riders riding known and unknown horses. In conclusion these data confirm that riding induces a significant increase in energy expenditure. During jumping, a mean value of 75% VO2max was reached. Therefore, a good aerobic capacity seems to be a factor determining riding performance in competitions. Regular riding practice and additional physical training are recommended to enhance the physical fitness of competitive riders.
	Address	Universite Paris XII-STAPS-61, Creteil, France. Mfdevienne@aol.com
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-6319	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10985607			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3731
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	Author	M, E.; östl,.; Messmann, S.; Bagu, E.; Robia, C.; Palme, R.
	Title	Measurement of Glucocorticoid Metabolite Concentrations in Faeces of Domestic Livestock			Type	Journal Article
	Year	1999	Publication	Journal of Veterinary Medicine Series A	Abbreviated Journal	J. Vet. Med. A
	Volume	46	Issue	10	Pages	621-631
	Keywords
	Abstract	After 14C-labelled cortisol infusion in ponies and pigs, faecal samples were collected. Extraction of 0.5 g faeces with 5 ml 80–90 % methanol yielded the highest radioactivity in the supernatant. Most of the metabolites were ether soluble. After high performance liquid chromatography (HPLC), the presence of immunoreactive metabolites was demonstrated by measuring each HPLC fraction using enzyme immunoassays for cortisol, corticosterone and 11-oxoaetiocholanolone. Only the assay for 11-oxoaetiocholanolone revealed peaks with co-eluting radioactivity. For biological validation of the test system, adrenocorticotrophic hormone (ACTH) and dexamethasone were injected intravenously successively in both species (n = 6). Cortisol concentration in blood and the 11-oxoaetiocholanolone immunoreactive substances in faeces were determined. In horse faeces, basal values of 2.3–35.2 nmol/kg were measured. After ACTH administration, an increase (more than 200 % above basal values) of these metabolites was seen about 1 day after ACTH administration. After dexamethasone injection the levels decreased, reaching minimum concentrations 2 days after administration. In pigs, an increase in these metabolites was measured in only three animals after ACTH; dexamethasone did not cause a decrease. The stability of the samples after defecation was tested by storing samples from cows, horses and pigs at room temperature. It was shown that there was a significant increase in the concentration of measured cortisol metabolites in bovine, equine and porcine faeces after storage for 1 h, 4 h and 24 h, respectively. In frozen samples this effect was diminished after thawing samples at 40°C; thawing the samples at 95°C prevented an increase in immunoreactive substances.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Science, Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0442	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6043
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	Author	Becker-Birck, M.; Schmidt, A.; Wulf, M.; Aurich, J.; von der Wense, A.; Möstl, E.; Berz, R.; Aurich, C.
	Title	Cortisol release, heart rate and heart rate variability, and superficial body temperature, in horses lunged either with hyperflexion of the neck or with an extended head and neck position			Type	Journal Article
	Year	2013	Publication	Journal of Animal Physiology and Animal Nutrition	Abbreviated Journal
	Volume	97	Issue	2	Pages	322-330
	Keywords	animal welfare; equitation; stress; training
	Abstract	Bringing the head and neck of ridden horses into a position of hyperflexion is widely used in equestrian sports. In our study, the hypothesis was tested that hyperflexion is an acute stressor for horses. Salivary cortisol concentrations, heart rate, heart rate variability (HRV) and superficial body temperature were determined in horses (n = 16) lunged on two subsequent days. The head and neck of the horse was fixed with side reins in a position allowing forward extension on day A and fixed in hyperflexion on day B. The order of treatments alternated between horses. In response to lunging, cortisol concentration increased (day A from 0.73 ± 0.06 to 1.41 ± 0.13 ng/ml, p < 0.001; day B from 0.68 ± 0.07 to 1.38 ± 0.13 ng/ml, p < 0.001) but did not differ between days A and B. Beat-to-beat (RR) interval decreased in response to lunging on both days. HRV variables standard deviation of RR interval (SDRR) and RMSSD (root mean square of successive RR differences) decreased (p < 0.001) but did not differ between days. In the cranial region of the neck, the difference between maximum and minimum temperature was increased in hyperflexion (p < 0.01). In conclusion, physiological parameters do not indicate an acute stress response to hyperflexion of the head alone in horses lunged at moderate speed and not touched with the whip. However, if hyperflexion is combined with active intervention of a rider, a stressful experience for the horse cannot be excluded.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0396	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6182
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	Author	Bamford, A.J.; Monadjem, A.; Hardy, I.C.W.
	Title	Associations of Avian Facial Flushing and Skin Colouration with Agonistic Interaction Outcomes			Type	Journal Article
	Year	2010	Publication	Ethology	Abbreviated Journal	Ethology
	Volume		Issue		Pages	no-no
	Keywords
	Abstract	Abstract Facial flushing, a colour change caused by variation of blood flow through highly vascularized skin, has been observed in taxonomically diverse bird species but the function of the behaviour has not been assessed. Lappet-faced vultures, Aegypius tracheliotos, have unfeathered heads that can rapidly flush from pink to dark red, and this has been hypothesized to indicate contest ability in vulture gatherings. We show that adults with flushed heads won most interactions against those with pale heads. A previously unnoticed colour variation of the throat, visible only when the head is flushed, was also related to the outcome of interactions: blue-throated adults participated in, and won, more interactions than red-throated adults. We suggest that the non-fixed groups of which lappet-faced vulture populations consist promote the evolution of signals of dominance that can be adjusted extremely rapidly.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5180
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	Author	Collier, T.C.; Blumstein, D.T.; Girod, L.; Taylor, C.E.
	Title	Is Alarm Calling Risky? Marmots Avoid Calling from Risky Places			Type	Journal Article
	Year	2010	Publication	Ethology	Abbreviated Journal	Ethology
	Volume		Issue		Pages	no-no
	Keywords
	Abstract	Abstract Alarm calling is common in many species. A prevalent assumption is that calling puts the vocalizing individual at increased risk of predation. If calling is indeed costly, we need special explanations for its evolution and maintenance. In some, but not all species, callers vocalize away from safety and thus may be exposed to an increased risk of predation. However, for species that emit bouts with one or a few calls, it is often difficult to identify the caller and find the precise location where a call was produced. We analyzed the spatial dynamics of yellow-bellied marmot (Marmota flaviventris) alarm calling using an acoustic localization system to determine the location from where calls were emitted. Marmots almost always called from positions close to the safety of their burrows, and, if they produced more than one alarm call, tended to end their calling bouts closer to safety than they started them. These results suggest that for this species, potential increased predation risk from alarm calling is greatly mitigated and indeed calling may have limited predation costs.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5181
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	Author	Hemelrijk, C.K.
	Title	Understanding Social Behaviour with the Help of Complexity Science (Invited Article)			Type	Journal Article
	Year	2002	Publication	Ethology	Abbreviated Journal	Ethology
	Volume	108	Issue	8	Pages	655-671
	Keywords
	Abstract	Abstract In the study of complexity, a new kind of explanation has been developed for social behaviour. It shows how patterns of social behaviour can arise as a side-effect of the interaction of individuals with their social or physical environment (e.g. by self-organization). This development may influence our ideas about the direct causation and evolution of social behaviour. Furthermore, it may influence our theories about the integration of different traits. This new method has been made possible by the increase in computing power. It is now applied in many areas of science, such as physics, chemistry, sociology and economics. However, in zoology and anthropology it is still rare. The major aim of this paper is to make this method more generally accepted among behavioural scientists.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Verlag, GmbH	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5200
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	Author	Hemelrijk, C.K.; Hildenbrandt, H.
	Title	Self-Organized Shape and Frontal Density of Fish Schools			Type	Journal Article
	Year	2008	Publication	Ethology	Abbreviated Journal	Ethology
	Volume	114	Issue	3	Pages	245-254
	Keywords
	Abstract	Abstract Models of swarming (based on avoidance, alignment and attraction) produce patterns of behaviour also seen in schools of fish. However, the significance of such similarities has been questioned, because some model assumptions are unrealistic [e.g. speed in most models is constant with random error, the perception is global and the size of the schools that have been studied is small (up to 128 individuals)]. This criticism also applies to our former model, in which we demonstrated the emergence of two patterns of spatial organization, i.e. oblong school form and high frontal density, which are supposed to function as protection against predators. In our new model we respond to this criticism by making the following improvements: individuals have a preferred ‘cruise speed’ from which they can deviate in order to avoid others or to catch up with them. Their range of perception is inversely related to density, with which we take into account that high density limits the perception of others that are further away. Swarm sizes range from 10 to 2000 individuals. The model is three-dimensional. Further, we show that the two spatial patterns (oblong shape and high frontal density) emerge by self-organization as a side-effect of coordination at two speeds (of two or four body lengths per second) for schools of sizes above 20. Our analysis of the model leads to the development of a new set of hypotheses. If empirical data confirm these hypotheses, then in a school of real fish these patterns may arise as a side-effect of their coordination in the same way as in the model.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5202
Permanent link to this record



	Author	Noë, R.; van Schaik, C.P.; van Hooff, J.A.R.A.M.
	Title	The Market Effect: an Explanation for Pay-off Asymmetries among Collaborating Animals			Type	Journal Article
	Year	1991	Publication	Ethology	Abbreviated Journal	Ethology
	Volume	87	Issue	1-2	Pages	97-118
	Keywords
	Abstract	Abstract * 1Animals can derive leverage over others from (a) resource holding power, based for instance on fighting ability or dominance, and (b) the possession of commodities, such as special skills and resources that cannot be taken away by force. * 2We contend that power based on the possession of commodities strongly depends on the level of supply and demand for that commodity, a phenomenon we call the ‘market effect’. * 3Several theoretical and empirical examples are given of social systems in which animals belong to two distinct classes that offer two different kinds of commodities. * 4The relative frequency of occurrence of the two classes is shown to determine the relative power of their members. * 5We consider the theoretical properties of bargaining processes by which relative power is converted into corresponding pay-off distributions. * 6We propose coalition games, a class of games with more than two players and in which bargaining is possible, as suitable paradigms for collaboration among members of social units.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Publishing Ltd	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5413
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	Author	Mettke-Hofmann, C.; Winkler, H.; Leisler, B.
	Title	The Significance of Ecological Factors for Exploration and Neophobia in Parrots			Type	Journal Article
	Year	2002	Publication	Ethology	Abbreviated Journal
	Volume	108	Issue	3	Pages	249-272
	Keywords
	Abstract	Exploratory behaviour plays an important role in most animals for gathering information about their environment. If it constitutes an adaptation to different environmental conditions exploratory behaviour should differ between species. This has been tested with several hypotheses. Sixty-one parrot species (Psittacidae) from eight tribes with different diets and habitat preferences were investigated in aviaries. Two tests were carried out. First, a novel object (wooden ring) in the familiar aviary was presented on two test days in the exploration test. Latencies until first contact with the object and the duration of exploration were recorded. Secondly, in the neophobia test, novel objects were placed beside the feeding dish and latencies until first food intake were recorded. The exploration and neophobia data were related to 12 (13) ecological variables using multiple regression analyses. Phylogenetic relationships were considered. Species that inhabit complex habitats, such as forest edges, or that feed on buds or species from islands showed the shortest latencies in the exploration test. In contrast, long latencies were related to a diet including a great amount of seeds and/or flowers. The longest duration of exploration occurred in species eating nuts or originating from islands, whereas short durations were related to feeding on seeds. Neophobia was positively related to a diet consisting of insects, and negatively to a diet of leaves. There was no relationship between measures of exploration and neophobia. Exploration and neophobia seem to be tightly related to the ecology of a species.
	Address
	Corporate Author				Thesis
	Publisher	Blackwell Wissenschafts-Verlag	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5617
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	Author	Briard, L.; Dorn, C.; Petit, O.
	Title	Personality and Affinities Play a Key Role in the Organisation of Collective Movements in a Group of Domestic Horses			Type	Journal Article
	Year	2015	Publication	Ethology	Abbreviated Journal	Ethology
	Volume	121	Issue	9	Pages	888-902
	Keywords	decision-making; equids; hierarchy; leadership; social network
	Abstract	Understanding how groups of individuals with different motives come to daily decisions about the exploitation of their environment is a key question in animal behaviour. While interindividual differences are often seen only as a threat to group cohesion, growing evidence shows that they may to some extent facilitate effective collective action. Recent studies suggest that personality differences influence how individuals are attracted to conspecifics and affect their behaviour as an initiator or a follower. However, most of the existing studies are limited to a few taxa, mainly social fish and arthropods. Horses are social herbivores that live in long-lasting groups and show identifiable personality differences between individuals. We studied a group of 38 individuals living in a 30-ha hilly pasture. Over 200 h, we sought to identify how far individual differences such as personality and affinity distribution affect the dynamic of their collective movements. First, we report that individuals distribute their relationships according to similar personality and hierarchical rank. This is the first study that demonstrates a positive assortment between unrelated individuals according to personality in a mammal species. Second, we measured individual propensity to initiate and found that bold individuals initiated more often than shy individuals. However, their success in terms of number of followers and joining duration did not depend on their individual characteristics. Moreover, joining process is influenced by social network, with preferred partners following each other and bolder individuals being located more often at the front of the movement. Our results illustrate the importance of taking into account interindividual behavioural differences in studies of social behaviours.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1439-0310	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6153
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