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Author Castley, J.G.; Knight, M.H. openurl 
  Title Population status of plains zebra, Equus burchelli, in South African National Parks. Type Journal Article
  Year 1997 Publication Scientific Services, National Parks Board Abbreviated Journal  
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  Publisher Place of Publication Kimberley, South Africa. Editor  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 2254  
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Author Dublin, H.T.; Sinclair, A.R.E.; Boutin, S.; Anderson, E.; Jago, M.; Arcese, P. doi  openurl
  Title Does competition regulate ungulate populations? Further evidence from Serengeti, Tanzania Type Journal Article
  Year 1990 Publication Oecologia Abbreviated Journal  
  Volume 82 Issue 2 Pages 283-288  
  Keywords  
  Abstract Changes in populations of several ungulate species in the Serengeti-Mara region of East Africa over the past 30 years suggest several hypotheses for their regulation and coexistence. Recent censuses in the 1980s have allowed us to test the hypotheses that: (1) there was competition between wildebeest (Connochaetes taurinus) and Thomson's gazelle (Gazella thomsoni). This predicted that gazelle numbers should have declined in the 1980s when wildebeest were food limited. Census figures show no change in gazelle numbers between 1978 and 1986, a result contrary to the interspecific competition hypothesis; (2) wildebeest and African buffalo (Syncerus caffer) populations were regulated by intraspecific competition for food. Since both populations reached food limitation in the 1970s, the hypothesis predicted that the populations should have been stable in the 1980s. The results confirm these predictions for wildebeest and the buffalo population in the Mara reserve. In the Serengeti the buffalo population declined 41% over the period 1976-1984. The decline was not evenly distributed over the park, some areas showing an 80-90% decline, others no change or an increase in numbers. The decline was associated with proximity to human habitation; (3) an outbreak of the viral disease, rinderpest, in 1982 may have been the cause of the drop in buffalo population. Blood serum samples to measure the prevalence of antibodies were collected from areas of decreasing, stable and increasing populations. If rinderpest was the cause of decrease there should be a negative relationship between the prevalence of rinderpest and the instantaneous rate of increase (r). The results showed no relationship. We conclude that rinderpest was not the major cause of the drop in buffalo numbers. Elephant (Loxodonta africana) numbers dropped 81% in Serengeti in the period 1977-1986. In the Mara there was little change. The evidence suggests that extensive poaching in northern and western Serengeti during 1979-1984 accounted for the drop in both elephant and buffalo numbers.  
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  Call Number Equine Behaviour @ team @ Serial 2281  
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Author Sigurjónsdóttir , H.; van Dierendonck, M.C.; Snorrason, S.; Thórhallsdóttir, A.G. url  doi
openurl 
  Title Social relationships in a group of horses without a mature stallion Type Journal Article
  Year 2003 Publication Behaviour Abbreviated Journal Behaviour  
  Volume 140 Issue 6 Pages 783-804  
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  Abstract 1. The social relationships in a group of Icelandic horses without a mature stallion were studied. The horses were all familiar to each other. Mutual grooming and play relationships, spatial associations, dominance-subordinate relations and the effect of kinship on these relationships were analysed.TAGSTARTBRTAGEND 2. The social structure was clearly dominated by the behaviour of the adult mares. The horses preferred to form bonds within their social class (sex/age) and they kept close proximity with their friends. The group was effectively divided into two social subgroups, adult mares as one group and adult geldings and sub-adults as another group. The sub-adults and adult geldings formed associations, which were based on mutual grooming and play, while the adult mares did not play. Differences between the sexes were evident. Males played more than the females, had more playing partners and were more popular as playmates.TAGSTARTBRTAGEND 3. Aggression rates were low. The dominance hierarchy was linear. Adult mares ranked higher than adult geldings, sub-adults and the foals. Rank was significantly correlated with age. The closer the adult mares were in rank, the more they groomed with each other. Such relationships were not found amongst the other social group.TAGSTARTBRTAGEND 4. Kinship was calculated between all pairs of animals for up to 4 or 5 generations. Allogrooming and play frequencies and proximity were all positively correlated with kinship. Adult mares, which were close in the dominance hierarchy, were on average more related than those further apart.TAGSTARTBRTAGEND 5. The social relationships in the Icelandic herd were, to some extent, different from relationships reported from unmanaged and feral horse-herds with mature stallions and bachelors. Our results suggest that adult mares groom more in groups without a stallion. Furthermore, they have more preferred partners than in natural harems and their partners are other adult mares, not their weaned offspring as seems to be the case in feral herds. The sub-adults also seem to be more socially active in the absence of stallions. Interestingly, in the Icelandic group, the adult mares showed stallion like behaviours, like mounting and protecting foals. Only by studying the behaviour and the nature of the relationships of horses in groups of different compositions, can we expect to gain a comprehensive understanding about individual social strategies and cognitive capabilities of the species. Such knowledge is valuable for management and welfare of the horse.  
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  Call Number Equine Behaviour @ team @ Serial 2363  
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Author VanDierendonck, M.C., de Vries, H., Schilder, M.B.H. url  doi
openurl 
  Title An Analysis of Dominance, Its Behavioural Parameters and Possible Determinants in a Herd of Icelandic horses in captivity Type Journal Article
  Year 1995 Publication Netherlands Journal of Zoology Abbreviated Journal Netherl. J. Zool.  
  Volume 45 Issue 3-4 Pages 362-385  
  Keywords Dominance; rank order; horses; Icelandic horses.  
  Abstract Feral horses are social animals, which have to rely on survival strategies centered on the formation of cohesive social bonds within their bands. Many problems in the husbandry of social animals such as horses, are due to the fact that the limits of their adaptive abilities are exceeded. Evidence suggests that the fundamental social characteristics of domestic horses have remained relatively unchanged. The social structure, social strategies and social interactions were investigated (3 non-consecutive years, 24 hr per day for several weeks) in long term established groups of domestic horses (mares and geldings of all ages) and a few small introduced groups, kept in (semi)natural environments. The general aim was to investigate the social needs of domestic horses. The social life of domestic horses was characterised by long lasting bonds with preferred partners which were established and maintained by allogrooming, play, proximity and dominance behaviours. Bonding partners were mainly found within the same sex-age group, but adult geldings also bonded with sub-adult mares and geldings. Adult mares were clustered in a group, while the other animals formed a second group. Among the adult mares, subgroups according to reproductive state were formed. Individuals regulated their social network by interfering with interactions between other members of the herd, which in itself is complex. An intervention is a behavioural action of one animal that actively interferes with an ongoing interaction between a dyad with the apparent aim of altering that interaction. This was verified by post-hoc analyses of disturbed and undisturbed interactions. Interventions in allogrooming or play were performed significantly more often when at least one member of the initial dyad was a preferred partner of, or familiar to (within the small introduced bands) the intervener. The stronger the preferred association in allogrooming between the intervener and member(s) of the initial dyad, the higher the probability the intervener would displace one initial member and continue allogrooming with the other. Just five behaviours were extracted which reliably reflected the dominance relations among horses. Aggression with the hind quarters was used both offensively and defensively and therefore not suitable as a reliable parameter. Individual dominance relationships were related to social experience. The implications of these findings for horse husbandry were assessed. It is argued that the execution of affiliative behaviours may be rewarding in itself, and therefore always will be a highly motivated behaviour. It is shown that social positive physical interactions (allogrooming, play) with other horses is an ethological need and therefore indispensable in modern husbandry systems. Ethological needs are so important for the animal that husbandry systems that lack the possibilities to execute such behaviours will cause chronic stress. It is concluded that all horses need physical social contact, and that horses, which lack appropriate social learning experiences during ontogeny, may be hampered in their social functioning later in life. Solutions for problems, including dominance problems, in individual social housing and group housing are presented.  
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  Call Number Equine Behaviour @ team @ Serial 2368  
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Author Moehlman, P.D.; Kebede, F.; Yohannes, H. url  doi
openurl 
  Title The African wild ass (Equus africanus): conservation status in the horn of Africa Type Journal Article
  Year 1998 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 60 Issue 2-3 Pages 115-124  
  Keywords Equus africanus; Critically endangered; Extinction  
  Abstract From 1989 to 1996, surveys were made in most of the historic range of African wild asses in Somalia, Ethiopia, and Eritrea. From the 1970s to the mid 1990s populations of African wild asses (Equus africanus, Fitzinger, 1857) in Somalia and Ethiopia have declined from approximately 6 to 30 per 100 km2 to 1 or 2 per 100 km2. Given the current IUCN criteria, they are Critically Endangered (CR) and face extremely high risk of extinction in the wild in the immediate future, as their populations have been reduced by at least 80% over the last 10+ years (IUCN, 1994). Basic research is needed on this species as scientific information on its reproductive biology, behavior, ecology, and genetics is very limited. Improved support needs to be provided to existing parks and reserves and new multiple use reserves need to be established.  
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  Call Number Equine Behaviour @ team @ Serial 2380  
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Author Moehlman, P.D.; Fowler, L.E.; Roe, J.H. url  doi
openurl 
  Title Feral asses (Equus africanus) of Volcano Alcedo, Galapagos: behavioral ecology, spatial distribution, and social organization Type Journal Article
  Year 1998 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 60 Issue 2-3 Pages 197-210  
  Keywords Equids; Feral asses; Social organization; Mating systems; Intraspecific variation; Galapagos  
  Abstract Feral asses were studied on Volcano Alcedo, Galapagos Islands, Ecuador, during the wet season of 1980. On the volcano rim during March/April, two stable groups were observed to have a `female (harem) defense' polygynous mating system [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), pp. 215-223] and social behavior patterns and feeding ecology similar to feral asses living in a habitat where forage and climate are similar, e.g., Ossabaw Island, Georgia [Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Rep., 1970, pp. 405-411; Moehlman, P.D., 1997. Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats. J. Appl. Anim. Behav. Sci., this issue; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia., PhD Dissertation, Pennsylvania State University, University Park, 219 pp.].  
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  Call Number Equine Behaviour @ team @ Serial 2383  
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Author Monard, A.-M.; Duncan, P.; Fritz, H.; Feh, C. doi  openurl
  Title Variations in the birth sex ratio and neonatal mortality in a natural herd of horses Type Journal Article
  Year 1997 Publication Behavioral Ecology and Sociobiology Abbreviated Journal Behav. Ecol. Sociobiol.  
  Volume 41 Issue 4 Pages 243-249  
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  Abstract Variations in birth sex ratios and sex differences in juvenile mortality occur in a number of mammalian species, and in many cases have been linked to resource availability. Most of these biases in offspring sex ratios concern polygynous species with pronounced sexual dimorphism, and where females only are philopatric. Data on species with unusual life-history strategies, such as slight sexual dimorphism or dispersal by both sexes, are of particular interest. In this study of a natural herd of horses (Equus caballus) which experienced an eruptive cycle, and therefore a period of nutritional stress, male offspring had higher neonatal mortality rates in nutritionally poor years than in good ones, whereas “year quality” had no effect on the mortality of female offspring; year quality could therefore be used by mares as predictor of sex-specific offspring survival. We show that the environmental conditions that predicted lower survival of males were negatively related to their production: the birth sex ratio the following year was female-biased; and mares were less likely to produce a son when they had produced a son the preceding year. There was no significant effect of mother's parity, age or rank, or the timing of conception or birth on offspring sex ratios. The mechanism leading to biases in the birth sex ratio could have been the loss of male embryos by mares that did not foal. As there was no evidence for selective abortion of male foetuses in females that did foal the next year, it is not necessary to invoke maternal adjustment, though this remains a possibility. Finally, there was a suggestion that male offspring were more costly to raise than females, since mothers that reared a son in poor years tended to experience an increase in the interbirth interval between their two subsequent offspring.  
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  Call Number Equine Behaviour @ team @ Serial 2388  
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Author Munthali, S.M.; Banda, H.M. doi  openurl
  Title Distribution and abundance of the common ungulates of Nyika National Park Type Journal Article
  Year 1992 Publication Afr. J. Ecol Abbreviated Journal Afr. J. Ecol  
  Volume 30 Issue Pages 203-212  
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  Call Number Equine Behaviour @ team @ Serial 2390  
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Author Mason, W.A.; Hollis, J.H. url  doi
openurl 
  Title Communication between young rhesus monkeys Type Journal Article
  Year 1962 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 10 Issue 3-4 Pages 211-221  
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  Abstract 1. 1. The communication performance of 12 rhesus monkeys was investigated in a situation in which the rewards of both members of a pair of monkeys could not exceed chance levels unless the operator monkey responded to cues provided by the informant monkey which indicated the location of food. Each member of the pair was trained in both operator and informant roles in different phases of the experiment. Communication performance improved progressively to levels consistently above chance. However, communication learning appeared to be specific to the role in which the individual was trained, and when roles were reversed no evidence of transfer was obtained. Tests of foodsharing behaviour showed a substantial increase in the tendency to share food with the partner following communication training. This occurred however, only when the partner was the only social stimulus present; if another monkey was also present there was no evidence of preferential responses to the partner. In all phases of communication training, monkeys which were housed together performed more efficiently than did monkeys housed individually.2. 2. The acquisition of stimulus-producing responses was investigated by causing an opaque screen to remain in front of the informant unless the operator monkey pulled a vertical lever at the front of its restraining cage. Initially, operators responded immediately to the foodcarts, but with further testing there was a steady increase in the tendency to defer the response to the food-carts until the lever had been pulled, revealing the informant monkey.3. 3. Transfer of communication training was tested with new monkey informants, and with two inanimate stimuli, a mechanical puppet, and a stationary plaque. The latter two objects were placed behind the rewarded food-carts before each trial. There was clear evidence of positive transfer to each of these conditions, but marked differences among conditions were obtained. Performance with the monkeys averaged 76 per cent. correct, as compared with 62 and 40 per cent., with the puppet and the plaque, respectively.4. 4. To test the ability of trained operator monkeys to select the appropriate informant on the basis of behavioural cues, the communication situation was arranged so that two informant monkeys were present on all trials. However, on any trial only one of these informants could be rewarded, and the operator's rewards were contingent upon delivering food to this informant. Efficiency of discrimination began at approximately 45 per cent, (chance = 25 per cent. and improved progressively to levels above 75 per cent.  
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  Call Number Equine Behaviour @ team @ Serial 3017  
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Author Diekamp, B.; Prior, H.; Güntürkün, O. doi  openurl
  Title Functional lateralization, interhemispheric transfer and position bias in serial reversal learning in pigeons (Columba livia) Type Journal Article
  Year 1999 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 2 Issue 4 Pages 187-196  
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  Abstract In the present study we investigated lateralization of color reversal learning in pigeons. After monocular acquisition of a simple color discrimination with either the left or right eye, birds were tested in a serial reversal procedure. While there was only a slight and non-significant difference in choice accuracy during original color discrimination, a stable superiority of birds using the right eye emerged in serial reversals. Both groups showed a characteristic 'learning-to-learn' effect, but right-eyed subjects improved faster and reached a lower asymptotic error rate. Subsequent testing for interocular transfer demonstrated a difference between pre- and post-shift choice accuracy in pigeons switching from right to left eye but not vice versa. This can be accounted for by differences in maximum performance using either the left or right eye along with an equally efficient but incomplete interocular transfer in both directions. Detailed analysis of the birds' response patterns during serial reversals revealed a preference for the right of two response keys in both groups. This bias was most pronounced at the beginning of a session. It decreased within sessions, but became more pronounced in late reversals, thus indicating a successful strategy for mastering the serial reversal task. Interocular transfer of response patterns revealed an unexpected asymmetry. Birds switching from right to left eye continued to prefer the right side, whereas pigeons shifting from left to right eye were now biased towards the left side. The results suggest that lateralized performance during reversal learning in pigeons rests on a complex interplay of learning about individual stimuli, stimulus dimensions, and lateralized response strategies.  
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  Call Number Equine Behaviour @ team @ Serial 3223  
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