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	Author	Langbein, J.; Nurnberg, G.; Puppe, B.; Manteuffel, G.
	Title	Self-Controlled Visual Discrimination Learning of Group-Housed Dwarf Goats (Capra hircus): Behavioral Strategies and Effects of Relocation on Learning and Memory			Type	Journal Article
	Year	2006	Publication	Journal of Comparative Psychology	Abbreviated Journal	J. Comp. Psychol.
	Volume	120	Issue	1	Pages	58-66
	Keywords	dwarf goats; visual discrimination; operant learning; learning strategies; context
	Abstract	In most studies on animal learning, individual animals are tested separately in a specific learning environment and with a limited number of trials per day. An alternative approach is to test animals in a familiar environment in their social group. In this study, the authors--applying a fully automated learning device--investigated voluntary, self-controlled visual shape discrimination learning of group-housed dwarf goats (Capra hircus). The majority of the tested goats showed successful shape discrimination, which indicates the adaptive value of an effective learning strategy. However, in each group, a few individual goats developed behavioral strategies different from shape discrimination to get reward. Relocation impairs memory retrieval (probably by attention shifting) only temporarily for previously learnt shapes. The results demonstrate the usefulness of a self-controlled learning paradigm to assess learning abilities of social species in their normal social settings. This may be especially relevant for captive animals to improve their welfare.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2140
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	Author	Lacreuse, A.; Martin-Malivel, J.; Lange, H.S.; Herndon, J.G.
	Title	Effects of the menstrual cycle on looking preferences for faces in female rhesus monkeys			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	2	Pages	105-115
	Keywords	Animals; Behavior, Animal; Discrimination Learning; Estradiol/blood; Face; Female; Humans; Macaca mulatta/physiology; Male; Menstrual Cycle/blood/physiology; *Pattern Recognition, Visual
	Abstract	Fluctuations of ovarian hormones across the menstrual cycle influence a variety of social and cognitive behaviors in primates. For example, female rhesus monkeys exhibit heightened interest for males and increased agonistic interactions with other females during periods of high estrogen levels. In the present study, we hypothesized that females' preference for males during periods of high estrogen levels is also expressed at the level of face perception. We tested four intact females on two face-tasks involving neutral portraits of male and female rhesus monkeys, chimpanzees and humans. In the visual preference task (VP), monkeys had to touch a button to view a face image. The image remained on the screen as long as the button was touched, and the duration of pressing was taken as an index of the monkey's looking time for the face stimulus. In the Face-Delayed Recognition Span Test (Face-DRST), monkeys were rewarded for touching the new face in an increasing number of serially presented faces. Monkeys were tested 5 days a week across one menstrual cycle. Blood was collected every other day for analysis of estradiol and progesterone. Two of the four females were cycling at the time of testing. We did not find an influence of the cycle on Face-DRST, likely due to a floor effect. In the VP however, the two cycling individuals looked longer at conspecific male faces than female faces during the peri-ovulatory period of the cycle. Such effects were absent for human and chimpanzee faces and for the two noncycling subjects. These data suggest that ovarian hormones may influence females' preferences for specific faces, with heightened preference for male faces during the peri-ovulatory period of the cycle. Heightened interest for stimuli of significant reproductive relevance during periods of high conception risk may help guide social and sexual behavior in the rhesus monkey.
	Address	Department of Psychology, University of Massachusetts-Amherst, Tobin Hall, 135 Hicks Way, Amherst, MA 01003, USA. alacreuse@psych.umass.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16909232			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2452
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	Author	Kuroshima, H.; Fujita, K.; Adachi, I.; Iwata, K.; Fuyuki, A.
	Title	A Capuchin monkey (Cebus apella) recognizes when people do and do not know the location of food			Type	Journal Article
	Year	2003	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	6	Issue	4	Pages	283-291
	Keywords	Adaptation, Psychological; Animals; Cebus/psychology; Communication; Concept Formation; Cues; Discrimination Learning; Feeding Behavior/*psychology; Female; Intention; Male; Social Identification; Transfer (Psychology)
	Abstract	In a previous study, Kuroshima and colleagues demonstrated that capuchin monkeys (Cebus apella) learned to discriminate between a “knower” who inspected a box for food, and a “guesser” who did not. The aim of the present study was to specify whether the subjects learned a simple conditional discrimination or a causal relationship that seeing leads to knowing. In experiment 1, we introduced five types of novel containers to two subjects. Each container was of different shape and color. The subjects gradually learned to reach toward the container the knower suggested. In experiment 2, we diversified the behavior of the knower and the guesser. In experiment 3, in order to eliminate the possibility of discrimination based on differences in the magnitude and the complexity of two trainers, we equated their behaviors. One subject adapted to the novel behaviors of the knower and the guesser, successfully discriminating the two trainers. Thus this monkey clearly learned to use the inspecting action of the knower and the non-inspecting action of the guesser as a discriminative cue to recognize the baited container. This result suggests that one capuchin monkey learned to recognize the relationship between seeing and knowing.
	Address	Graduate School of Letters, Department of Psychology, Kyoto University, Yoshida-honmachi, Sakyo, 606-8501 Kyoto, Japan. kuroshi@psy.bun.kyoto-u.ac.jp
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12905080			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2558
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	Author	Krzeminska, W.
	Title	[The child learns about the world]			Type	Journal Article
	Year	1979	Publication	Pielegniarka i polozna	Abbreviated Journal	Pieleg Polozna
	Volume		Issue	7	Pages	24-25
	Keywords	Child; Child Development; Child, Preschool; Humans; Learning
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	Polish	Summary Language		Original Title	Dziecko poznaje swiat
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0048-4148	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:260249			Approved	no
	Call Number	refbase @ user @			Serial	43
Permanent link to this record



	Author	Krueger., K.; Farmer, K.
	Title	Social learning in Horses: Differs from individual learning only in the learning stimulus and not in the learning mechanisms			Type	Abstract
	Year	2018	Publication	14th Meeting of the Internatinoal Society for Equitation Science	Abbreviated Journal	14th Meeting ISES
	Volume		Issue		Pages
	Keywords	horse; individual learning; learning mechanisms; learning stimuli; social learning
	Abstract	Equine welfare can be enhanced by applying species specific training. This may incorporate social learning, as horses are highly social and social stimuli are of primary importance. Social learning is comparable to individual learning in its learning mechanisms, differing primarily in the way it is stimulated. Our initial study showed that horses of different breeds (N = 38) follow humans after observing other horses doing so, but only if the observed horse was familiar to and higher ranking than the observer (Fisher's exact test: N = 12, P = 0.003). A second study showed that horses and ponies (N = 25) learned to pull a rope to open a feeding apparatus after observing demonstrations by conspecifics, again, only if the demonstrating horse was older and higher ranking than the observer (Fisher's combination test, N = 3, v2 = 27.71, p = 0.006). Our third approach showed that horses and ponies (N = 24) learned to press a switch to open a feeding apparatus after observing a familiar person (GzLM: N = 24, z = 2.33, P = 0.02). Most recently, we confronted horses and ponies (N = 50) with persons demonstrating different techniques for opening a feeding apparatus. In this study we investigated whether the horses would copy the demonstrators' techniques or apply their own. Here only some horses copied the technique, and most of the successful learners used their mouths irrespective of the demonstrators' postures (Chi Square Test: N = 40, df = 2, χ2 = 31.4, p < 0.001). In all the approaches social stimuli elicited learning processes in the test horses, while only a few individuals in the control groups mastered the tasks by individual learning. The following behaviour observed in the initial study may have been facilitated by a social stimuli (social facilitation), and the opening of the feed boxes in the subsequent studies appear to be mostly the result of enhancement (social enhancement). Some horses may have used the social stimuli at first and continued their learning process by individual trial and error. However, the horses were also selective in whom and some in how to copy. This may have been conditioned (socially conditioned) or the result of simple forms of reasoning on the reliability of the particular information provided by demonstrators of certain social ranks or social positions, as high ranking and familiar horses and familiar persons were copied and some imitated exactly. Lay person message: Traditional riding instructions suggest that horses learn by observing other horses. For example, older, more experienced driving horses are used for initial training of young driving horses. We have shown that horses indeed use learning stimuli provided by other horse, as well as by humans. Horses readily accept stimuli observed in high ranking and familiar horses, and familiar persons. Such stimuli elicit learning processes which are comparable to individual learning. We suggest applying social learning whenever possible, as it is much faster and less stressful than individual learning, where learners experience negative outcomes in trial and error learning.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6405
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	Author	Krueger, K.; Farmer, K.; Heinze, J.
	Title	The effects of age, rank and neophobia on social learning in horses			Type	Journal Article
	Year	2014	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	17	Issue	3	Pages	645-655
	Keywords	Horse; Social learning; Sociality; Ecology; Social relationships
	Abstract	Social learning is said to meet the demands of complex environments in which individuals compete over resources and co-operate to share resources. Horses (Equus caballus) were thought to lack social learning skills because they feed on homogenously distributed resources with few reasons for conflict. However, the horse’s social environment is complex, which raises the possibility that its capacity for social transfer of feeding behaviour has been underestimated. We conducted a social learning experiment using 30 socially kept horses of different ages. Five horses, one from each group, were chosen as demonstrators, and the remaining 25 horses were designated observers. Observers from each group were allowed to watch their group demonstrator opening a feeding apparatus. We found that young, low ranking, and more exploratory horses learned by observing older members of their own group, and the older the horse, the more slowly it appeared to learn. Social learning may be an adaptive specialisation to the social environment. Older animals may avoid the potential costs of acquiring complex and potentially disadvantageous feeding behaviours from younger group members. We argue that horses show social learning in the context of their social ecology, and that research procedures must take such contexts into account. Misconceptions about the horse’s sociality may have hampered earlier studies.
	Address
	Corporate Author				Thesis
	Publisher	Springer Berlin Heidelberg	Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5737
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	Author	Krueger, K. (ed)
	Title	Proceedings of the International Equine Science Meeting 2008			Type	Conference Volume
	Year	2008	Publication	IESM 2008	Abbreviated Journal
	Volume		Issue		Pages
	Keywords	Equine Ecology; Equine Sociality; Equine Learning; Equine Cognition; Equine Welfare
	Abstract	Target group: Biologists, Psychologists, Veterinarians and Professionals Meeting target: Because the last international meeting on Equine Science took place a couple years ago, there is an urgent need for equine scientists to exchange scientific knowledge, coordinate research provide knowledge for practical application, and discus research results among themselves and with professionals who work with horses. Additionally, dialog concerning the coordination of the study “Equitation Science” in Europe is urgently needed. Coordination and cooperation shall arise from the meeting, enrich the research, and advance the application of scientific knowledge for the horses` welfare.
	Address
	Corporate Author				Thesis
	Publisher	Xenophon Verlag	Place of Publication	Wald	Editor	Krueger, K.
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-3-9808134-0-2	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4508
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	Author	Krueger, K.
	Title	Behaviour of horses in the “round pen technique”			Type	Journal Article
	Year	2007	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	104	Issue	1-2	Pages	162-170
	Keywords	Learning; Dominance relationship; Horse; Human-animal relationships
	Abstract	I investigated the behavioural background of the way horses learn to follow humans in the “round pen technique” suggested by “horse whisperers” as a gentle method for initial horse training. Though the practicability of this technique has been adequately demonstrated in the past, the horses' behaviour during such training has not yet been documented in detail. In a riding arena, horses, that did not follow the trainer immediately, were chased away so that they galloped around the trainer. Galloping horses showed specific behaviour such as turning the ear to the trainer, chewing, licking, and stretching head and throat downwards. In subsequent trials horses needed to be chased for less time and finally followed immediately, even when conditions were changed or the trainer was replaced by another person. This suggests that horses learn to follow in this particular situation and also show some generalisation. However, following did not occur on a pasture even after several successful trials in the riding arena.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	280
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	Author	Krueger, K.
	Title	Social learning and innovative behaviour in horses			Type	Conference Article
	Year	2015	Publication	Proceedings of the 3. International Equine Science Meeting	Abbreviated Journal	Proc. 3. Int. Equine. Sci. Mtg
	Volume		Issue		Pages
	Keywords	social learning, innovative behaviour, Equus caballus, cognitive capacities
	Abstract	The evaluation of important parameters for measuring the horses’ cognitive capacities is one of the central topics of the equine behaviour team at Nürtingen-Geislingen University. Social complexity has been said to be one of the settings in which needs for cognitive capacities arise in animals. A variety of studies throughout the last two decades proved the horses’ social complexity to be far more elaborate than previously assumed. Horses form social bonds for the protection of offspring, intervene in encounters of others, identify group mates individually and easily orientate in a fission fusion society. In such socially complex societies, animals will benefit from learning socially. In many bird and primate species the degree of social complexity correlates nicely with the species abilities for social learning. Social learning was, therefore, argued to be an indicator for elaborate mental capacities in animals. We were delighted to prove that horses actually copy social behaviour and techniques for operating a feeding apparatus from older and higher ranking group members. In a recent study we found young horses, at the age of 3 to 12, to copy the operation of a feeding apparatus from a human demonstrator. Social learning seems to work nicely in horses when the social background of the animals is considered. The degree to which individual animals adapt to changes in their social or physical environment by finding innovative solution appears to be the other side of the coin, of whether animals adjust to challenges by social learning. It is not very astonishing, that along with the animals’ social complexity and their ability to learn socially also the degree to which they show innovative behaviour was claimed to be one of the most important demonstrations of advanced cognitive capacities. In a recent approach, we started to ask horse owners and horse keepers in many countries to tell us about unusual behaviour of their horses via a web site (http://innovative-behaviour.org). To date, we received 204 cases of innovative behaviour descriptions from which six cases were clear examples of tool use or borderline tool use. We categorized the innovative behaviours into the classes, a) innovations to gain food, b) innovations to gain freedom, c) social innovations, d) innovations to increase maintenance, and e) innovations that could not be clearly assigned to a category. About 20% of the innovative horses showed more than one innovation. These animals could be termed “true innovators”. Again, young horses were more innovative than older ones with the age group 5 – 9 showing the highest number of innovative behaviour descriptions. In a nutshell, the horses’ cognitive capacities appear to be underestimated throughout the last decades. The horses’ social complexity is far more elaborate than previously assumed, horses learn socially from conspecific and humans, some of them demonstrate innovative behaviour adaptations to their environment and even simple forms of tool use.
	Address
	Corporate Author	Krueger, K.			Thesis
	Publisher	Xenophon Publishing	Place of Publication	Wald	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume	in prep	Series Issue		Edition
	ISSN		ISBN	978-3-95625-000-2	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5848
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	Author	Koba, Y.; Tanida, H.
	Title	How do miniature pigs discriminate between people?: Discrimination between people wearing coveralls of the same colour			Type	Journal Article
	Year	2001	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	73	Issue	1	Pages	45-58
	Keywords	Pigs; Learning; Recognition; Human-animal relationships
	Abstract	Seven experiments were conducted on four miniature pigs to determine: (1) whether the pigs can discriminate between people wearing the same coloured clothing; (2) what cues they rely on if they could discriminate. For 2 weeks before the experiments began, the pigs were conditioned in a Y-maze to receive raisins from the rewarder wearing dark blue coveralls. They were then given the opportunity to choose the rewarder or non-rewarder in these experiments. Each session consisted of 20 trials. Successful discrimination was that the pig chose the rewarder at least 15 times in 20 trials (P<0.05: by χ2-test). In Experiment 1, both rewarder and non-rewarder wore dark blue coveralls. By 20 sessions, all pigs successfully identified the rewarder. In Experiment 2: (1) both wore coveralls of the same new colours or (2) one of them wore coveralls of new colours. They significantly preferred the rewarder even though the rewarder and/or non-rewarder wore coveralls of new colours. In Experiment 3, both wore dark blue coveralls but olfactory cues were obscured and auditory cues were not given. The pigs were able to identify the rewarder successfully irrespective of changing auditory and olfactory cues. In Experiment 4, both wore dark blue coveralls but covered part of their face and body in different ways. The correct response rate decreased when a part of the face and the whole body of the rewarder and non-rewarder were covered. In Experiment 5, both wore dark blue coveralls and changed their apparent body size by shifting sitting position. The correct response rate increased as the difference in body size between the experimenters increased. In Experiment 6, the distance between the experimenters and the pig was increased by 30 cm increments. The correct response rate of each pig decreased as the experimenters receded from the pig, but performance varied among the pigs. In Experiment 7, the light intensity of the experimental room was reduced from 550 to 80 lx and then to 20 lx. The correct response rate of each pig decreased with the reduction in light intensity, but all the pigs discriminated the rewarder from the non-rewarder significantly even at 20 lx. In conclusion, the pigs were able to discriminate between people wearing coveralls of the same colour after sufficient reinforcement. These results indicate that pigs are capable of using visual cues to discriminate between people.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	839
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