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	Author	Zentall, T.R.; Clement, T.S.; Bhatt, R.S.; Allen, J.
	Title	Episodic-like memory in pigeons			Type	Journal Article
	Year	2001	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	8	Issue	4	Pages	685-690
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Memory, Short-Term/*physiology; Teaching
	Abstract	It has been proposed that memory for personal experiences (episodic memory, rather than semantic memory) relies on the conscious review of past experience and thus is unique to humans. In an attempt to demonstrate episodic-like memory in animals, we first trained pigeons to respond to the (nonverbal) question “Did you just peck or did you just refrain from pecking?” by training them on a symbolic matching task with differential responding required to the two line-orientation samples and reinforcing the choice of a red comparison if they had pecked and the choice of a green comparison if they had not pecked. Then, in Experiment 1, after providing the conditions for (but not requiring) the pigeons to peck at one new stimulus (a yellow hue) but not at another (a blue hue), we tested them with the new hue stimuli and the red and green comparisons. In Experiment 2, we tested the pigeons with novel stimuli (a circle, which they spontaneously pecked, and a dark response key, which they did not peck) and the red and green comparisons. In both experiments, pigeons chose the comparison appropriate to the response made to the test stimulus. Thus, the pigeons demonstrated that they could remember specific details about their past experiences, a result consistent with the notion that they have the capacity for forming episodic-like memories.
	Address	Department of Psychology, University of Kentucky, Lexington 40506-0044, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11848586			Approved	no
	Call Number	refbase @ user @			Serial	243
Permanent link to this record



	Author	Urcuioli, P.J.; Zentall, T.R.
	Title	Retrospective coding in pigeons' delayed matching-to-sample			Type	Journal Article
	Year	1986	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	12	Issue	1	Pages	69-77
	Keywords	Animals; Color Perception; Columbidae; Discrimination Learning; Form Perception; Memory; Mental Recall; Orientation; Pattern Recognition, Visual; Retention (Psychology)
	Abstract	In this study we examined how coding processes in pigeons' delayed matching-to-sample were affected by the stimuli to be remembered. In Experiment 1, two groups of pigeons initially learned 0-delay matching-to-sample with identical comparison stimuli (vertical and horizontal lines) but with different sample stimuli (red and green hues or vertical and horizontal lines). Longer delays were then introduced between sample offset and comparison onset to assess whether pigeons were prospectively coding the same events (viz., the correct line comparisons) or retrospectively coding different events (viz., their respective sample stimuli). The hue-sample group matched more accurately and showed a slower rate of forgetting than the line-sample group. In Experiment 2, pigeons were trained with either hues or lines as both sample and comparison stimuli, or with hue samples and line comparisons or vice versa. Subsequent delay tests revealed that the hue-sample groups remembered more accurately and generally showed slower rates of forgetting than the line-sample groups. Comparison dimension had little or no effect on performance. Together, these data suggest that pigeons retrospectively code the samples in delayed matching-to-sample.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3701260			Approved	no
	Call Number	refbase @ user @			Serial	263
Permanent link to this record



	Author	Cerutti, D.T.; Staddon, J.E.R.
	Title	Immediacy versus anticipated delay in the time-left experiment: a test of the cognitive hypothesis			Type	Journal Article
	Year	2004	Publication	Journal of Experimental Psychology. Animal Behavior Processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	30	Issue	1	Pages	45-57
	Keywords	Animals; Choice Behavior/physiology; Cognition/physiology; Columbidae; Male; Models, Psychological; Psychological Theory; Reinforcement (Psychology); Reinforcement Schedule; Time Perception/*physiology
	Abstract	In the time-left experiment (J. Gibbon & R. M. Church, 1981), animals are said to compare an expectation of a fixed delay to food, for one choice, with a decreasing delay expectation for the other, mentally representing both upcoming time to food and the difference between current time and upcoming time (the cognitive hypothesis). The results of 2 experiments support a simpler view: that animals choose according to the immediacies of reinforcement for each response at a time signaled by available time markers (the temporal control hypothesis). It is not necessary to assume that animals can either represent or subtract representations of times to food to explain the results of the time-left experiment.
	Address	Department of Psychological and Brain Sciences, Duke University, Durham, NC 27708-1050, USA. cerutti@psych.duke.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14709114			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2768
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	Author	Zentall, T.R.; Weaver, J.E.; Clement, T.S.
	Title	Pigeons group time intervals according to their relative duration			Type	Journal Article
	Year	2004	Publication	Psychonomic bulletin & review	Abbreviated Journal	Psychon Bull Rev
	Volume	11	Issue	1	Pages	113-117
	Keywords	Animals; Columbidae; *Discrimination (Psychology); Reinforcement (Psychology); Time Factors; Time Perception
	Abstract	In the present research, we asked whether pigeons tended to judge time intervals not only in terms of their absolute value but also relative to a duration from which they must be discriminated (i.e., longer or shorter). Pigeons were trained on two independent temporal discriminations. In one discrimination, sample durations of 2 and 8 sec were associated with, for example, red and green hue comparisons, respectively, and in the other discrimination, sample durations of 4 and 16 sec were associated with vertical and horizontal line comparisons, respectively. If pigeons are trained on a temporal discrimination and tested with intermediate durations, the subjective midpoint typically occurs close to the geometric mean of the two trained values. The 4- and 8-sec values were selected to be the geometric mean of the two values in the other discrimination. When a 4-sec test sample was presented with the comparisons from the 2- and 8-sec discrimination, the pigeons preferred the comparison associated with the shorter sample. Similarly, when an 8-sec test sample was presented with the comparisons from the 4- and 16-sec discrimination, the pigeons preferred the comparison associated with the longer sample. Thus, a relative grouping effect was found. That is, durations that should have produced indifferent choice were influenced by their relative durations (shorter than or longer than the alternative) during training.
	Address	Department of Psychology, University of Kentucky, Lexington, Kentucky 40506-0044, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1069-9384	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15116995			Approved	no
	Call Number	refbase @ user @			Serial	231
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	Author	Dorrance, B.R.; Zentall, T.R.
	Title	Imitation of conditional discriminations in pigeons (Columba livia)			Type	Journal Article
	Year	2002	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	116	Issue	3	Pages	277-285
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Conditioning (Psychology)/physiology; Discrimination Learning/physiology; Habituation, Psychophysiologic; *Imitative Behavior; Light; Reinforcement (Psychology)
	Abstract	In the present experiments, the 2-action method was used to determine whether pigeons could learn to imitate a conditional discrimination. Demonstrator pigeons (Columba livia) stepped on a treadle in the presence of 1 light and pecked at the treadle in the presence of another light. Demonstration did not seem to affect acquisition of the conditional discrimination (Experiment 1) but did facilitate its reversal of the conditional discrimination (Experiments 2 and 3). The results suggest that pigeons are not only able to learn a specific behavior by observing another pigeon, but they can also learn under which circumstances to perform that behavior. The results have implications for proposed mechanisms of imitation in animals.
	Address	Department of Psychology, Augustana College, Rock Island, Illinois 61201, USA. psdorrance@augustana.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12234078			Approved	no
	Call Number	refbase @ user @			Serial	240
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	Author	Zentall, T.R.; Roper, K.L.; Sherburne, L.M.
	Title	Most directed forgetting in pigeons can be attributed to the absence of reinforcement on forget trials during training or to other procedural artifacts			Type	Journal Article
	Year	1995	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	63	Issue	2	Pages	127-137
	Keywords	Animals; Attention; Color Perception; Columbidae; Cues; Discrimination Learning; Mental Recall; Motivation; Pattern Recognition, Visual; Reinforcement Schedule; Retention (Psychology)
	Abstract	In research on directed forgetting in pigeons using delayed matching procedures, remember cues, presented in the delay interval between sample and comparisons, have been followed by comparisons (i.e., a memory test), whereas forget cues have been followed by one of a number of different sample-independent events. The source of directed forgetting in delayed matching to sample in pigeons was examined in a 2 x 2 design by independently manipulating whether or not forget-cue trials in training ended with reinforcement and whether or not forget-cue trials in training included a simultaneous discrimination (involving stimuli other than those used in the matching task). Results were consistent with the hypothesis that reinforced responding following forget cues is sufficient to eliminate performance deficits on forget-cue probe trials. Only when reinforcement was omitted on forget-cue trials in training (whether a discrimination was required or not) was there a decrement in accuracy on forget-cue probe trials. When reinforcement is present, however, the pattern of responding established during and following a forget cue in training may also play a role in the directed forgetting effect. These findings support the view that much of the evidence for directed forgetting using matching procedures may result from motivational and behavioral artifacts rather than the loss of memory.
	Address	Department of Psychology, University of Kentucky, Lexington 40506
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7714447			Approved	no
	Call Number	refbase @ user @			Serial	256
Permanent link to this record



	Author	Zentall, T.R.; Clement, T.S.
	Title	Memory mechanisms in pigeons: evidence of base-rate neglect			Type	Journal Article
	Year	2002	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	28	Issue	1	Pages	111-115
	Keywords	Animals; Columbidae; Discrimination Learning; Memory/*physiology; Random Allocation; Reaction Time; Reinforcement (Psychology); Retention (Psychology)
	Abstract	In delayed matching to sample, once acquired, pigeons presumably choose comparisons according to their memory for (the strength of) the sample. When memory for the sample is sufficiently weak, comparison choice should depend on the history of reinforcement associated with each of the comparison stimuli. In the present research, pigeons acquired two matching tasks in which Sample S1 was associated with one comparison from each task, C1 and C3, whereas Sample S2 was associated with Comparison C2, and Sample S3 was associated with Comparison C4. As the retention interval increased, the pigeons showed a bias to choose the comparison (C1 or C3) associated with the more frequently occurring sample (S1). Thus, pigeons were sensitive also to the (irrelevant) likelihood that each of the samples was presented. The results suggest that pigeons may allow their reference memory for the overall sample frequency to influence comparison choice, independent of the comparison stimuli present.
	Address	Department of Psychology, University of Kentucky, Lexington, Kentucky 40506-0044, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11868229			Approved	no
	Call Number	refbase @ user @			Serial	242
Permanent link to this record



	Author	Clement, T.S.; Weaver, J.E.; Sherburne, L.M.; Zentall, T.R.
	Title	Simultaneous discrimination learning in pigeons: value of S- affects the relative value of its associated S+			Type	Journal Article
	Year	1998	Publication	The Quarterly journal of experimental psychology. B, Comparative and physiological psychology	Abbreviated Journal	Q J Exp Psychol B
	Volume	51	Issue	4	Pages	363-378
	Keywords	Animals; Attention; Color Perception; Columbidae; Discrimination Learning; Female; Male; *Motivation; Orientation; Transfer (Psychology)
	Abstract	In a simple simultaneous discrimination involving a positive stimulus (S+) and a negative stimulus (S-), it has been hypothesized that positive value can transfer from the S+ to the S- (thus increasing the relative value of the S-) and also that negative value can transfer from the S- to the S+ (thus diminishing the relative value of the S+; Fersen, Wynne, Delius, & Staddon, 1991). Evidence for positive value transfer has been reported in pigeons (e.g. Zentall & Sherburne, 1994). The purpose of the present experiments was to determine, in a simultaneous discrimination, whether the S- diminishes the value of the S+ or the S- is contrasted with the S+ (thus enhancing the value of the S+). In two experiments, we found evidence for contrast, rather than value transfer, attributable to simultaneous discrimination training. Thus, not only does the S+ appear to enhance the value of the S-, but the S- appears to enhance rather than reduce the value of the S+.
	Address	Department of Psychology, University of Kentucky, Lexington 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0272-4995	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:9854439			Approved	no
	Call Number	refbase @ user @			Serial	252
Permanent link to this record



	Author	Martin, T.I.; Zentall, T.R.
	Title	Post-choice information processing by pigeons			Type	Journal Article
	Year	2005	Publication	Animal cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	4	Pages	273-278
	Keywords	Animals; Behavior, Animal; Choice Behavior; *Columbidae; Discrimination Learning
	Abstract	In a conditional discrimination (matching-to-sample), a sample is followed by two comparison stimuli, one of which is correct, depending on the sample. Evidence from previous research suggests that if the stimulus display is maintained following an incorrect response (the so-called penalty-time procedure), acquisition by pigeons is facilitated. The present research tested the hypothesis that the penalty-time procedure allows the pigeons to review and learn from the maintained stimulus display following an incorrect choice. It did so by including a penalty-time group for which, following an incorrect choice, the sample changed to match the incorrect comparison, thus providing the pigeons with post-choice 'misinformation.' This misinformation group acquired the matching task significantly slower than the standard penalty-time group (that had no change in the sample following an error). Furthermore, acquisition of matching by a control group that received no penalty time fell midway between the other two groups, suggesting that the pigeons did not merely take more care in making choices because of the aversiveness of penalty-time. Thus, it appears that in the acquisition of matching-to-sample, when the stimulus display is maintained following an incorrect choice, the pigeons can review or acquire information from the display. This is the first time that such an effect has been reported for a nonhuman species.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15744507			Approved	no
	Call Number	refbase @ user @			Serial	225
Permanent link to this record



	Author	Urcuioli, P.J.; DeMarse, T.B.; Zentall, T.R.
	Title	Transfer across delayed discriminations: II. Differences in the substitutability of initial versus test stimuli			Type	Journal Article
	Year	1998	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	24	Issue	1	Pages	47-59
	Keywords	Animals; Behavior, Animal; Columbidae/physiology; Discrimination Learning/physiology; Pattern Recognition, Visual/physiology; Reinforcement (Psychology); Retention (Psychology)/physiology
	Abstract	In 2 experiments, pigeons were trained on, and then transferred to, delayed simple discriminations in which the initial stimuli signalled reinforcement versus extinction following a retention interval. Experiment 1 showed that discriminative responding on the retention test transferred to novel test stimuli that had appeared in another delayed simple discrimination but not to stimuli having the same reinforcement history off-baseline. By contrast, Experiment 2 showed that performances transferred to novel initial stimuli whether they had been trained on-baseline or off-baseline. These results suggest that the test stimuli in delayed simple discriminations acquire control over responding only in the memory task itself. On the other hand, control by the initial stimuli, if coded as outcome expectancies, does not require such task-specific training.
	Address	Department of Psychological Sciences, Purdue University, West Lafayette, Indiana 47907-1364, USA. uche@psych.purdue.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:9438965			Approved	no
	Call Number	refbase @ user @			Serial	253
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