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	Author	Parker, G.A.
	Title	Assessment strategy and the evolution of fighting behaviour			Type	Journal Article
	Year	1974	Publication	Journal of Theoretical Biology	Abbreviated Journal	J. Theor. Biol.
	Volume	47	Issue	1	Pages	223-243
	Keywords
	Abstract	The view is examined that the adaptive value of conventional aspects of fighting behaviour is for assessment of relative RHP (resource holding power) of the combatants. Outcomes of aggressive disputes should be decided by each individual's fitness budget available for expenditure during a fight (determined by the fitness difference between adoption of alternative strategies, escalation or withdrawal without escalation) and on the rate of expenditure of the fitness budget if escalation occurs (determined by the RHPs of the combatants). Thus response thresholds for alternative strategies (“assessments”) will be determined by natural selection on a basis of which opponent is likely to expend its fitness budget first, should escalation occur. This “loser” should retreat (before escalation) and the winner should stay in possession of the resource. Many aggressive decisions depend on whether one is a resource holder, or an attacker. Assuming the RHP of the combatants to be equal, there are many instances of fitness pay-off imbalances between holder and attacker which should weight the dispute outcome in favour of one or other opponent by allowing it a greater expendable fitness budget. Usually the weighting favours the holder; the attacker therefore needs a correspondingly higher RHP before it may be expected to win. This is not invariably the case, and much observed data fits the predictions of this sort of model. If assessments are perfect and budget expenditure rates exactly predictable, then there would never seem to be any case for escalation. Escalation can be explained in terms of injury inflictions (expenditures) occurring as discrete events; i.e. as “bouts” won or lost during fighting. Assessment can give only a probabilistic prediction of the outcome of a bout. A simple model is developed to investigate escalation situations. Each combatant assesses relative RHP; this correlates with an absolute probability of winning the next bout (cabs). The stake played for is infliction of loss of RHP and is determined by the fitness budgets of the opponents. (Each individual plays for the withdrawal of its opponent.) This defines a critical probability of winning (ccrit) for each combatant, above which escalation is the favourable strategy (cabs > ccrit) and below which withdrawal is favourable (cabs < ccrit). Escalation should occur only where cabs-ccrit is positive for both combatants. This model gives predictions compatible with the observations, indicating that RHP loss alone can be adequate to explain withdrawal: escalation behaviour. Withdrawal tendency will be increased by low searching costs. Escalations should be restricted to closely matched RHP opponents if RHP disparity is the major imbalance. Outside the “escalation range” of a given individual, the higher RHP individual wins and the lower one loses (i.e. it should withdraw after conventional display). RHP disparity and holder: attacker imbalance should both interact to shape the observed pattern, though their relative importances will depend on species and situation. In some instances selection may favour immediate withdrawal from an occupied territory even without assessment of RHP.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4935
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	Author	Syme, G.J.
	Title	Competitive orders as measures of social dominance			Type	Journal Article
	Year	1974	Publication		Abbreviated Journal
	Volume	22	Issue	Part 4	Pages	931-940
	Keywords
	Abstract	The use of competitive orders as measures of social dominance is examined, the conclusion being that such use is based on the assumption of the unidimensionality of social dominance. Evidence is presented to show that this is not always the case. Consequently it is suggested that each competitive order must be validated in terms of its measurement of priority of access and response requirements (internal validity) as well as its generality (external validity) before it can be regarded as a dominance measure. Problems of the validity of aggression orders as measures of social dominance are also examined along with their relationship to competitive orders.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2188
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	Author	Jarman, P.J .
	Title	The social behaviour of antelope in relation to their ecology			Type	Journal Article
	Year	1974	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	48	Issue	1-4	Pages	213-267
	Keywords
	Abstract	The types of social organisation displayed by the African antelope species have been assigned in this paper to five classes, distinguished largely by the strategies used by the reproductively active males in securing mating rights, and the effects of those strategies on other social castes. The paper attempts to show that these strategies are appropriate to each class because of the effects of other, ecological, aspects of their ways of life. The paper describes different feeding styles among antelope, in terms of selection of food items and coverage of home ranges. It argues that these feeding styles bear a relationship to maximum group size of feeding animals through the influence of dispersion of food items upon group cohesion. The feeding styles also bear a relationship to body size and to habitat choice, both of which influence the antelope species' antipredator behaviour. Thus feeding style is related to anti-predator behaviour which, in many species, influences minimum group size. Group size and the pattern of movement over the annual home range affect the likelihood of females being found in a given place at a given time, and it is this likelihood which, to a large extent, determines the kind of strategy a male must employ to achieve mating rights. The effects of the different strategies employed by males can be seen in such aspects of each species' biology as sexual dimorphism, adult sex ratio, and differential distribution of the sexes.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4264
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	Author	Syme, G.J.; Pollard, J.S.; Syme, L.A.; Reid, R.M.
	Title	An analysis of the limited access measure of social dominance in rats			Type	Journal Article
	Year	1974	Publication		Abbreviated Journal
	Volume	22	Issue	2	Pages	486-500
	Keywords
	Abstract	The limited access situation in which only one of two or more subjects can gain access to a reward during a restricted time-period is an accepted measure of dominance in the rat. This study attempts to validate the technique by establishing the relationship between individual and competitive performance in order to determine whether `priority of access' has been measured. The generality of the competitive orders is examined by altering the competitive response while retaining the same reward. In view of the data collected for both time and weight-gain measures in food and water competition it is doubtful whether the limited access competitive technique can be considered a valid measure of dominance for the laboratory rat.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2187
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	Author	Dunbar, R.I.M.
	Title	Observations on the ecology and social organization of the green monkey,Cercopithecus sabaeus, in Senegal			Type	Journal Article
	Year	1974	Publication	Primates	Abbreviated Journal	Primates
	Volume	15	Issue	4	Pages	341-350
	Keywords
	Abstract	The green monkey,Cercopithecus sabaeus, has not been studied in its natural habitat in West Africa. This paper reports observations made during a 3-month study in Senegal. Green monkeys live in multimale groups averaging some 12 individuals. Information is given on home range size, use of habitat, daily activity patterns, diet and birth seasonality. Social organization is discussed and data are given on the relationships between age-sex classes, aggression and leadership. Inter-group relations are discussed and it is suggested that groups defend their ranges as territories. The ecology and social organization of green monkeys is compared with that of populations ofC. aethiops studied in East Africa and they are found to be similar.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2062
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	Author	Richards, S.M.
	Title	The concept of dominance and methods of assessment			Type	Journal Article
	Year	1974	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	22	Issue	Part 4	Pages	914-930
	Keywords
	Abstract	The arrangement of a social group of individuals into a dominance hierarchy is useful in studies of social behaviour only if a wide variety of social interactions can then be predicted. However, definitions of dominance commonly used are numerous and confused. To assess the usefulness of the concept of dominance, studies were made on six breeding groups of rhesus macaques (Macaca mulata) to determine whether different measures of dominance agreed with each other. The measures tested in this study were found to agree. It is therefore suggested that dominance is a useful intervening variable. Possible reasons for the reported lack of correlation between some measures used by other authors are discussed.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2154
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	Author	Altmann, J.
	Title	Observational Study of Behavior: Sampling Methods			Type	Journal Article
	Year	1974	Publication	Behaviour	Abbreviated Journal	Behaviour
	Volume	49	Issue	3-4	Pages	227-266
	Keywords
	Abstract	Seven major types of sampling for observational studies of social behavior have been found in the literature. These methods differ considerably in their suitability for providing unbiased data of various kinds. Below is a summary of the major recommended uses of each technique: In this paper, I have tried to point out the major strengths and weaknesses of each sampling method. Some methods are intrinsically biased with respect to many variables, others to fewer. In choosing a sampling method the main question is whether the procedure results in a biased sample of the variables under study. A method can produce a biased sample directly, as a result of intrinsic bias with respect to a study variable, or secondarily due to some degree of dependence (correlation) between the study variable and a directly-biased variable. In order to choose a sampling technique, the observer needs to consider carefully the characteristics of behavior and social interactions that are relevant to the study population and the research questions at hand. In most studies one will not have adequate empirical knowledge of the dependencies between relevant variables. Under the circumstances, the observer should avoid intrinsic biases to whatever extent possible, in particular those that direcly affect the variables under study. Finally, it will often be possible to use more than one sampling method in a study. Such samples can be taken successively or, under favorable conditions, even concurrently. For example, we have found it possible to take Instantaneous Samples of the identities and distances of nearest neighbors of a focal individual at five or ten minute intervals during Focal-Animal (behavior) Samples on that individual. Often during Focal-Animal Sampling one can also record All Occurrences of Some Behaviors, for the whole social group, for categories of conspicuous behavior, such as predation, intergroup contact, drinking, and so on. The extent to which concurrent multiple sampling is feasible will depend very much on the behavior categories and rate of occurrence, the observational conditions, etc. Where feasible, such multiple sampling can greatly aid in the efficient use of research time.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4684
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	Author	Geist, V.
	Title	On the Relationship of Social Evolution and Ecology in Ungulates			Type	Journal Article
	Year	1974	Publication	Amer. Zool.	Abbreviated Journal
	Volume	14	Issue	1	Pages	205-220
	Keywords
	Abstract	Much of the social behavior and organization of ungulates can be related to ecological parameters such as fiber content of forage, plant productivity, plant biomass, plant species diversity, productivity gradients, temporal and spatial fluctuations in productivity, habitat stability, food dispersion, three-dimensional structure of habitat, colonization, and predator density and diversity. These ecological variables can be linked via individual natural selection with the species' anti-predator strategies, emphasis on different channels of communication, relative frequency of damaging and non-damaging overt aggression, gregariousness and group structure, juvenile dispersal, home-range traditions, monogamy and polygamy, sexual dimorphism, territoriality, hierarchical rank structure, and plasticity of social structures. The ecological variables have primary manifestations which are behavior or which affect behavior, as well as secondary manifestations affecting behavior. There are logical links between the hypothesis linking ecology and behavior discussed here with some principles from bioenergetics, zoogeography, and paleontology. Although links do exist between ecology and behavior, they nevertheless represent distinct realms of natural selection in which social behavior appears as the more conservative element. The theoretical basis for this is discussed.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4261
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	Author	Andrew, R.J.
	Title	Changes in visual responsiveness following intercollicular lesions and their effects on avoidance and attack			Type	Journal Article
	Year	1974	Publication	Brain, Behavior and Evolution	Abbreviated Journal	Brain Behav Evol
	Volume	10	Issue	4-5	Pages	400-424
	Keywords	Animals; Chickens; Humans; Male; Mutism; Superior Colliculi/*physiology; Tectum Mesencephali; Testosterone; Visual Fields; Vocalization, Animal
	Abstract	In the normal chick, conspicuous visual stimuli induce targetting and pecking together, with vocalization. All three are abolished by lesion of the intercollicular area (ICo) or of connections passing through its medial margin. After such lesions, chicks also cease to treat significant visual stimuli as if they were startling and exciting, and may delay response as a result. However, they are still able to recognise, orient accurately to, and respond appropriately to, a variety of complex visual stimuli (e.g. food grains, copulation object). In addition, they are little affected by strange surroundings. Lesion evidence suggests the mammalian subcollicular area to have similar functions to the ICo and to be homologous with it. A route (present in bird), which is well-known in mammals for its association with threat, defense and escape evoked by strange and frightening objects (amygdala-diencephalic periventricular system-central mesencephalic grey, A-DPS-CMG) is stimuli via the 2 ICo (subcollicular area). Two different mechanisms may be involved caudal to the ICo. One consists of tectal afferents which might modulate the evocation of targetting, pecking and other responses via the tectum. The other is the predorsal system of tectal efferents which may mediate such responses. Classical syndromes of tameness and unresponsiveness produced by various interruptions of the A-DPS-CMG route may depend on interruption of connections to these midbrain mechanisms. Attack is depressed by ICo lesions as one aspect of reduced responsiveness to conspicuous and startling visual stimuli. Avoidance, which is apparently mediated by a separate system, much as in Anura, is facilitated.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-8977	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:1169102			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4626
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	Author	Powell, G.V.N.
	Title	Experimental analysis of the social value of flocking by starlings (Sturnus vulgaris) in relation to predation and foraging			Type	Journal Article
	Year	1974	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	22	Issue	2	Pages	501-505
	Keywords
	Abstract	In groups of ten, indidual starlings, Sturnus vulgaris, spent significantly less time in surveillance than did individuals in smaller groups and responded more quickly than single birds to a flying model hawk. Captive starlings in flocks reduce their individual surveillance efforts, but their combined efforts still enable them to be more effective than single birds in the detection of predators. Foraging behaviour of flocks was observed by placing single starlings with groups of tricoloured blackbirds, Agelaius tricolor; the starlings reduced the time they devoted to surveillance at the same rate as if they were with other starlings.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2147
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