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Author Smuts, M.M.S.; Penzhorn, B. L.
Title Descriptions of antomical differences between skulls and mandibles of Equus zebra and E. burchelli from southern Africa Type Journal Article
Year 1988 Publication South African Journal of Zoology Abbreviated Journal South African Journal of Zoology
Volume 23 Issue (4)3 Pages 328-336
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Notes from Professor Hans Klingels Equine Reference List Approved yes
Call Number Serial 1617
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Author Sigurjonsdottir, H.; Thorhallsdottir, A.; Hafthorsdottir, H.; Granquist S.
Title The Behaviour of Stallions in a Semiferal Herd in Iceland: Time Budgets, Home Ranges, and Interactions Type Journal Article
Year 2012 Publication International Journal of Zoology Abbreviated Journal
Volume 2012 Issue Article ID 162982 Pages
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Abstract (up) A permanent herd of Icelandic horses with four stallions and their harems was studied for a total of 316 hours in a large pasture (215 ha) in May 2007 in Iceland. Interactions between stallions of different harems and other aspects of the horses' behaviour were studied. One stallion and nine horses were introduced into the pasture prior to the study to examine the reactions of the resident stallions to a newcomer. The stallions spent significantly less time grazing than other horses and were more vigilant. Home ranges overlapped, but harems never mixed. The stallions prevented interactions between members of different harems indirectly by herding. Generally, interactions between resident stallions were nonviolent. However, encounters with the introduced stallion were more aggressive and more frequent than between the other stallions. Here, we show that four harems can share the same enclosure peacefully. The social network seems to keep aggression at a low level both within the harems and the herd as a whole. We encourage horse owners to consider the feasibility of keeping their horses in large groups because of low aggression and because such a strategy gives the young horses good opportunities to develop normally, both physically and socially.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5823
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Author Alexander, R, MCN et al
Title Fast locomotion of some african ungulates Type Journal Article
Year 1977 Publication Journal of Zoology Abbreviated Journal J Zool
Volume 183 Issue 3 Pages 291-300
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Abstract (up) ABSTRACT

Ten species of ungulate were filmed, galloping in their natural habitat. They ranged in size from Thomson's gazelle (about 20 kg) to giraffe (about 1000 kg). They were pursued to make them run as fast as possible. The films have been analysed to determine speed, stride frequency, stride and step lengths, and duty factors. The dependence of these quantities on body size is discussed.



Summary:

Fast locomotion of zebra, giraffe, warthog and seven species of Bovidae has been studied. The animals were filmed from a pursuing vehicle while galloping in their natural habitat.

Stride frequency was more closely correlated with limb length (represented by hip height) than with body mass. Mean stride frequency was proportional to (hip height)-0·51 and maximum stride frequency to (hip height) -0·63.

Maximum speed was between 10 and 14 m s -1 for all species except buffalo (7 m s -1). It was not significantly correlated with body mass.

Since the small species ran at least as fast as the large ones they attained higher Froude numbers. Relative stride length was approximately 1·8 (Froude number)0·39 for all species, irrespective of size. Relative step length was approximately 0·65 (Froude number)0·2, both for the fore feet and for the hind ones. The vertical forces exerted by the feet are proportional to (body weight)×(Froude number)0·2 so the forces at maximum speed are larger multiples of body weight for small species than for large ones.
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Notes from Prof. Hans Klingels Equine Reference List Approved no
Call Number refbase @ user @ Serial 130
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Author King, S.R.B.; Gurnell, J.
Title Scent-marking behaviour by stallions: an assessment of function in a reintroduced population of Przewalski horses (Equus ferus przewalskii) Type Journal Article
Year 2007 Publication Journal of Zoology Abbreviated Journal J Zool
Volume 272 Issue 1 Pages 30-36
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Abstract (up) Abstract Scent marking is a common form of intraspecific communication in mammal species, and using faeces or urine is a cost-effective way of signalling competitive ability and resource holding power. Marking is ritually performed by male equids, and here we assess the function of male scent-marking behaviour in a recently introduced population of Przewalski horses Equus ferus przewalskii in Mongolia. Two forms of scent marking were observed: defecation on stud piles formed from repeated dunging in the same place, and overmarking of faeces and urine of mares. Stud piles were marked with dung by the harem holder and sniffed before and after dung was deposited. They were not found specifically at the periphery of harem ranges but occurred for the most part along routes used by the horses, and were more common in the core parts of harem ranges or where harem ranges overlapped. Thus, rather than being used to defend range boundaries, stud piles were placed predominantly where they would be encountered by male intruders. Mare excreta were covered with urine by the stallion, but were only sniffed before they were marked, not after. These marks appear to advertise to the mare and other, intruding stallions that the harem holder was the mare's consort and that the interloper should not risk trying to steal the mare or sneak a mating. Thus, the two forms of marking by harem holders appear to combine as first and second lines of defence of paternity rights in male intrasexual competition.
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Notes Approved no
Call Number Equine Behaviour @ team @ King2007 Serial 2315
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Author Geisbauer, G.; Griebel, U.; Schmid, A.; Timney, B
Title Brightness discrimination and neutral point Type Journal Article
Year 2004 Publication Canadian Journal of Zoology Abbreviated Journal Can. J. Zool
Volume 82 Issue 4 Pages 660-670
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Abstract (up) Abstract: Equine brightness discrimination ability and color discrimination were measured using a two-choice discrimination

task. Two Haflinger horses (Equus caballus L., 1758) were trained to discriminate 30 different shades of grey

varying from low to high relative brightness. Their ability to distinguish shades of grey was poor, with calculated

Weber fractions of 0.42 and 0.45. In addition, a “neutral point” test to determine the dimensionality of color vision

was carried out. Three hues of blue-green were tested versus a range of grey targets with brightnesses similar to those

of the blue-green targets. A neutral point was found at about 480 nm. Thus, we can conclude that horses possess

dichromatic color vision.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 3649
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Author Weckerly, Floyd W.
Title Social bonding and aggression in female Roosevelt elk Type Journal Article
Year 1999 Publication Canadian Journal of Zoology Abbreviated Journal Can J Zool
Volume 77 Issue 9 Pages 1379-1384
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Abstract (up) Abstract: The relationship between degree of social bonding (extent of association among individuals) and level of aggression in ruminants is unclear. I examined social bonding and aggression in three groups of female Roosevelt elk (Cervus elaphus roosevelti) over 2 years. I hypothesized that when animals are socially bonded, bouts of aggression will be won by the individual initiating the aggression, occur quickly, and involve little physical contact, and the level of aggression does not correlate with group size. The degree of social bonding was high among individuals in all groups. Dyads of known individuals were together >80% of the time. A permutation analysis indicated that groups with the observed sizes had <0.001 chance of random association, except on one occasion when the probability was 0.72 for one group. Using focal-animal sampling, aggressive interactions were won 72% of the time by the initiator, occurred quickly (<5 s), and involved little physical contact, and the level of aggression was not correlated with group size. The level of aggression was, however, significantly lower in one of the groups. This group may have had access to more abundant food resources than the other groups. Socially bonded elk conducted aggressive interactions in a fashion that did not disrupt social stability. Résumé : La relation entre le degré de liaison sociale (importance des associations entre individus) et l`agressivité n`est pas claire chez les ruminants. J`ai étudié les liaisons sociales et l`agressivité chez trois groupes de femelles du Cerf de Roosevelt (Cervus elaphus roosevelti) pendant 2 ans. J`ai posé en hypothèse que, chez les animaux liés socialement, la victoire devrait être emportée par l`individu qui entreprend l`agression, l`agression devrait être de courte durée, se faire avec peu de contacts physiques et la fréquence des agressions ne devrait pas être liée à la taille du groupe. Des paires d`individus passaient plus de 80% de leur temps ensemble. Une analyse des permutations a démontré que, chez les groupes des tailles observées, la probabilité d`une association aléatoire était de moins de 0,001, sauf en un cas où cette probabilité a été évaluée à 0,72 chez un groupe. Par échantillonnage directionnel, j`ai observé que les interactions agressives étaient gagnées par l`individu attaquant 72% du temps, étaient de courte durée (<5 s), se faisaient avec peu de contacts physiques et leur fréquence n`était pas reliée à la taille du groupe. Il y avait cependant moins d`agressivité chez l`un des groupes. Il se peut que ce groupe ait eu accès à plus de ressources alimentaires que les autres. Chez les cerfs liés par des liens sociaux, l`agressivité ne se manifeste pas de façon à déséquilibrer la stabilité sociale.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4268
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Author VanDierendonck, M.C., de Vries, H., Schilder, M.B.H.
Title An Analysis of Dominance, Its Behavioural Parameters and Possible Determinants in a Herd of Icelandic horses in captivity Type Journal Article
Year 1995 Publication Netherlands Journal of Zoology Abbreviated Journal Netherl. J. Zool.
Volume 45 Issue 3-4 Pages 362-385
Keywords Dominance; rank order; horses; Icelandic horses.
Abstract (up) Feral horses are social animals, which have to rely on survival strategies centered on the formation of cohesive social bonds within their bands. Many problems in the husbandry of social animals such as horses, are due to the fact that the limits of their adaptive abilities are exceeded. Evidence suggests that the fundamental social characteristics of domestic horses have remained relatively unchanged. The social structure, social strategies and social interactions were investigated (3 non-consecutive years, 24 hr per day for several weeks) in long term established groups of domestic horses (mares and geldings of all ages) and a few small introduced groups, kept in (semi)natural environments. The general aim was to investigate the social needs of domestic horses. The social life of domestic horses was characterised by long lasting bonds with preferred partners which were established and maintained by allogrooming, play, proximity and dominance behaviours. Bonding partners were mainly found within the same sex-age group, but adult geldings also bonded with sub-adult mares and geldings. Adult mares were clustered in a group, while the other animals formed a second group. Among the adult mares, subgroups according to reproductive state were formed. Individuals regulated their social network by interfering with interactions between other members of the herd, which in itself is complex. An intervention is a behavioural action of one animal that actively interferes with an ongoing interaction between a dyad with the apparent aim of altering that interaction. This was verified by post-hoc analyses of disturbed and undisturbed interactions. Interventions in allogrooming or play were performed significantly more often when at least one member of the initial dyad was a preferred partner of, or familiar to (within the small introduced bands) the intervener. The stronger the preferred association in allogrooming between the intervener and member(s) of the initial dyad, the higher the probability the intervener would displace one initial member and continue allogrooming with the other. Just five behaviours were extracted which reliably reflected the dominance relations among horses. Aggression with the hind quarters was used both offensively and defensively and therefore not suitable as a reliable parameter. Individual dominance relationships were related to social experience. The implications of these findings for horse husbandry were assessed. It is argued that the execution of affiliative behaviours may be rewarding in itself, and therefore always will be a highly motivated behaviour. It is shown that social positive physical interactions (allogrooming, play) with other horses is an ethological need and therefore indispensable in modern husbandry systems. Ethological needs are so important for the animal that husbandry systems that lack the possibilities to execute such behaviours will cause chronic stress. It is concluded that all horses need physical social contact, and that horses, which lack appropriate social learning experiences during ontogeny, may be hampered in their social functioning later in life. Solutions for problems, including dominance problems, in individual social housing and group housing are presented.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 2368
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Author de Villiers, M.S.; Richardson, P.R.K.; van Jaarsveld, A.S.
Title Patterns of coalition formation and spatial association in a social carnivore, the African wild dog (Lycaon pictus) Type Journal Article
Year 2003 Publication Journal of Zoology Abbreviated Journal J Zool
Volume 260 Issue 4 Pages 377-389
Keywords coalitions; social systems; Carnivora; Lycaon pictus
Abstract (up) In many social species, relationships within groups seem to be non-random but related to variables such as rank, kinship or sexual attractiveness. The endangered African wild dog Lycaon pictus is a social carnivore that lives in large, stable packs, and intra-pack associations might be expected to display similar patterns. We investigated patterns of coalition formation (support during dominance interactions, and partnership interactions) and resting associations between members of a captive pack of 19 wild dogs. The social organization of the captive pack was similar to that of free-ranging packs in many respects. Polyadic (group) incidents of coalition support were also observed in a free-ranging pack. Patterns of coalition formation in the captive pack were related to rank. Most aggressive interactions involved high-ranking individuals (particularly the alpha, beta and third-ranking males) and coalitionary support tended to reinforce the existing hierarchy. However, there was at least one example of support influencing a successful rank challenge. Support was affected by potential risks and benefits, the latter including dominance through association and revolutionary alliances. An even stronger pattern overlaid associations between pack members: coalitions and resting associations were strongest between members of the same age–sex cohort, and may have enabled the eventual dominance of younger pack members over adults. Among adults, coalitionary associations were sometimes overridden by intersexual relationships. The results from this captive pack suggest that wild dogs are sensitive to differences in competitive ability. This information, in conjunction with strong affiliative bonds between littermates, is used to manoeuvre for position in the social hierarchy. It may also be important during dispersal, in encounters with other dispersing groups of the same sex. Although most features of the social structure of the captive pack were comparable to those of free-ranging packs, aspects such as the influence of relatedness on coalition formation still need to be explored.
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Publisher Blackwell Publishing Ltd Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN 1469-7998 ISBN Medium
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5249
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Author Mace, G.M.; Harvey, P.H.; Clutton-Brock, T.H.
Title Brain size and ecology in small mammals Type Journal Article
Year 1981 Publication Journal of Zoology Abbreviated Journal J Zool
Volume 193 Issue 3 Pages 333-354
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Abstract (up) Relative brain size (measured as gross brain size after body size effects are removed) differs systematically between families of rodents, insectivores and lagomorphs. The Sciuridae have the largest relative brain size, the Soricidae and Bathyergidae the smallest. These results are discussed and compared with previous analyses of relative brain sizes among primates and bats. These differences complicate comparisons between relative brain size across phylogenetically diverse species and attempts to relate differences in relative brain size to ecological variables. To overcome these problems, best fit relationships were estimated for each family, and values for each genus were expressed as deviations from the lines of best fit. We refer to these values as Comparative Brain Size (CBS). Differences in CBS are related to differences in habitat type (forest-dwelling genera have larger CBS' than grassland forms), in diet (folivores have smaller CBS' than generalists or insectivores, frugivores and granivores), in zonation (arboreal genera have larger CBS' than terrestrial ones) and in activity timing (nocturnal genera have larger CBS' than dirurnal ones). However, these ecological categories are interrelated and, when the effects of other ecological differences are taken into account using analyses of variance, only the differences associated with diet, and possibly habitat remain.
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Publisher Blackwell Publishing Ltd Place of Publication Editor
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ISSN 1469-7998 ISBN Medium
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5455
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Author Leadbeater, E.
Title What evolves in the evolution of social learning? Type Journal Article
Year 2015 Publication Journal of Zoology Abbreviated Journal J Zool
Volume 295 Issue 1 Pages 4-11
Keywords social learning; associative learning; social information use
Abstract (up) Social learning is fundamental to social life across the animal kingdom, but we still know little about how natural selection has shaped social learning abilities on a proximate level. Sometimes, complex social learning phenomena can be entirely explained by Pavlovian processes that have little to do with the evolution of sociality. This implies that the ability to learn socially could be an exaptation, not an adaptation, to social life but not that social learning abilities have been left untouched by natural selection. I discuss new empirical evidence for associative learning in social information use, explain how natural selection might facilitate the associative learning process and discuss why such studies are changing the way that we think about social learning.
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ISSN 1469-7998 ISBN Medium
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6015
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