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	Author	Hockenhull, J.; Creighton, E.
	Title	Unwanted oral investigative behaviour in horses: A note on the relationship between mugging behaviour, hand-feeding titbits and clicker training			Type	Journal Article
	Year	2010	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	127	Issue	3-4	Pages	104-107
	Keywords	Equine; Horse; Titbits; Food rewards; Clicker training; Mugging behaviour
	Abstract	Unwanted oral investigative in horses has been anecdotally attributed to the practice of hand-feeding. Fears over such behaviour developing as a consequence of using food rewards, for example in clicker training, have been implicated as a common reason for not employing food-based positive reinforcement training techniques. This study used data generated as part of a larger research project, and explored associations between five common oral investigative behaviours and the practices of hand-feeding and clicker training. Data were from a convenience sample of UK leisure horse owners using two self-administered Internet surveys. Ninety-one percent of respondents reported giving their horse food by hand and this practice was significantly associated with three of the five oral investigative behaviours, licking hands (P = 0.006), gently searching clothing (P < 0.001) and roughly searching clothing (P = 0.003). Nipping hands and biting clothes were not associated with hand-feeding, suggesting that risk factors for these behaviours originate outside of this practice. Clicker training techniques were employed by 14% of respondents and their use was not associated with the incidence of any of the five oral investigative behaviours. These findings suggest that horse owners should not be deterred from using food-based positive reinforcement techniques with their horses, as fears that this practice will result in unwanted oral investigative behaviours from their horses appear unfounded.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5183
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	Author	Friedrich, A.M.; Zentall, T.R.
	Title	Pigeons shift their preference toward locations of food that take more effort to obtain			Type	Journal Article
	Year	2004	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	67	Issue	3	Pages	405-415
	Keywords	Animals; Behavior, Animal; Choice Behavior; Columbidae; Exertion; Feeding Behavior; Reward
	Abstract	Although animals typically prefer to exert less effort rather than more effort to obtain food, the present research shows that requiring greater effort to obtain food at a particular location appears to increase the value of that location. In Experiment 1, pigeons' initial preference for one feeder was significantly reduced by requiring 1 peck to obtain food from that feeder and requiring 30 pecks to obtain food from the other feeder. In Experiment 2, a similar decrease in preference was not found when pigeons received reinforcement from both feeders independently of the amount of effort required. These results are consistent with the within-trial contrast effect proposed by in which the relative hedonic value of a reward depends on the state of the animal immediately prior to the reward. The greater the improvement from that prior state the greater the value of the reinforcer.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15518990			Approved	no
	Call Number	refbase @ user @			Serial	227
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	Author	Dunbar, R.I.M.; McAdam, M.R.; O'connell, S.
	Title	Mental rehearsal in great apes (Pan troglodytes and Pongo pygmaeus) and children			Type	Journal Article
	Year	2005	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	69	Issue	3	Pages	323-330
	Keywords	Algorithms; Animals; Child; Child, Preschool; Food; Frontal Lobe/anatomy & histology/physiology; Humans; Imagination; Pan troglodytes; Pongo pygmaeus; Problem Solving/physiology; Psychomotor Performance/physiology; Reward
	Abstract	The ability to rehearse possible future courses of action in the mind is an important feature of advanced social cognition in humans, and the “social brain” hypothesis implies that it might also be a feature of primate social cognition. We tested two chimpanzees, six orangutans and 63 children aged 3-7 years on a set of four puzzle boxes, half of which were presented with an opportunity to observe the box before being allowed to open it (“prior view”), the others being given without an opportunity to examine the boxes before handling them (“no prior view”). When learning effects are partialled out, puzzle boxes in the “prior view” condition were opened significantly faster than boxes given in the “no prior view” condition by the children, but not by either of the great apes. The three species differ significantly in the speed with which they opened boxes in the “no prior view” condition. The three species' performance on this task was a function of relative frontal lobe volume, suggesting that it may be possible to identify quantitative neuropsychological differences between species.
	Address	Evolutionary Psychology Research Group, School of Biological Sciences, University of Liverpool, Biosciences Building, Crown Street, Liverpool L69 7ZB, UK. rimd@liv.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15896530			Approved	no
	Call Number				Serial	2097
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	Author	Pepperberg, I.M.
	Title	In search of king Solomon's ring: cognitive and communicative studies of Grey parrots (Psittacus erithacus)			Type	Journal Article
	Year	2002	Publication	Brain, behavior and evolution	Abbreviated Journal	Brain Behav Evol
	Volume	59	Issue	1-2	Pages	54-67
	Keywords	Animal Communication; Animals; Attention/physiology; Cognition/physiology; Cues; Form Perception/physiology; Humans; Intelligence; Learning/physiology; Male; Models, Psychological; Parrots/*physiology; Psychomotor Performance/physiology; Reward; Social Behavior
	Abstract	During the past 24 years, I have used a modeling technique (M/R procedure) to train Grey parrots to use an allospecific code (English speech) referentially; I then use the code to test their cognitive abilities. The oldest bird, Alex, labels more than 50 different objects, 7 colors, 5 shapes, quantities to 6, 3 categories (color, shape, material) and uses 'no', 'come here', wanna go X' and 'want Y' (X and Y are appropriate location or item labels). He combines labels to identify, request, comment upon or refuse more than 100 items and to alter his environment. He processes queries to judge category, relative size, quantity, presence or absence of similarity/difference in attributes, and show label comprehension. He semantically separates labeling from requesting. He thus exhibits capacities once presumed limited to humans or nonhuman primates. Studies on this and other Greys show that parrots given training that lacks some aspect of input present in M/R protocols (reference, functionality, social interaction) fail to acquire referential English speech. Examining how input affects the extent to which parrots acquire an allospecific code may elucidate mechanisms of other forms of exceptional learning: learning unlikely in the normal course of development but that can occur under certain conditions.
	Address	The MIT Media Lab, Cambridge, Mass. 02139, USA. impepper@media.mit.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-8977	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12097860			Approved	no
	Call Number	refbase @ user @			Serial	579
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	Author	Pollmann, U.
	Title	[Keeping of horses in circus and show businesses]			Type	Journal Article
	Year	2002	Publication	DTW. Deutsche Tierarztliche Wochenschrift	Abbreviated Journal	Dtsch Tierarztl Wochenschr
	Volume	109	Issue	3	Pages	126-129
	Keywords	Animal Husbandry/methods; Animal Welfare; Animals; Behavior, Animal; Horses; Housing, Animal; Humans; Reward
	Abstract	The conditions under which horses are kept and the performance of acts in the circus ring may give rise to animal protection-relevant aspects for circus and show horses. A number of intolerable conditions under which horses are kept and procedures adopted for the work with circus and show horses are described. In addition, attention is drawn to monitoring methods capable of exposing the deplorable shortcomings of these businesses.
	Address	Fachbereich Ethologie und Tierschutz des Chemischen und Veterinaruntersuchungsamtes Freiburg. Ursula.Pollmann@cvuafr.bwl.de
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	German	Summary Language		Original Title	Pferdehaltung in Zirkus- und Schaustellerbetrieben
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0341-6593	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11963363			Approved	no
	Call Number	refbase @ user @			Serial	1914
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	Author	Brosnan, S.F.; de Waal, F.B.M.
	Title	Socially learned preferences for differentially rewarded tokens in the brown capuchin monkey (Cebus apella)			Type	Journal Article
	Year	2004	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	118	Issue	2	Pages	133-139
	Keywords	Animals; Behavior, Animal; Cebus; Choice Behavior; Female; Learning; Male; Reward; Social Behavior
	Abstract	Social learning is assumed to underlie traditions, yet evidence indicating social learning in capuchin monkeys (Cebus apella), which exhibit traditions, is sparse. The authors tested capuchins for their ability to learn the value of novel tokens using a previously familiar token-exchange economy. Capuchins change their preferences in favor of a token worth a high-value food reward after watching a conspecific model exchange 2 differentially rewarded tokens, yet they fail to develop a similar preference after watching tokens paired with foods in the absence of a conspecific model. They also fail to learn that the value of familiar tokens has changed. Information about token value is available in all situations, but capuchins seem to pay more attention in a social situation involving novel tokens.
	Address	Living Links Center, Yerkes National Primate Research Center, and Graduate Division of Biological and Biomedical Sciences, Emory University, Atlanta, GA 30329, USA. sbrosna@emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15250800			Approved	no
	Call Number	refbase @ user @			Serial	173
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	Author	Brosnan, S.F.; De Waal, F.B.M.
	Title	Monkeys reject unequal pay			Type	Journal Article
	Year	2003	Publication	Nature	Abbreviated Journal	Nature
	Volume	425	Issue	6955	Pages	297-299
	Keywords	Aging; Animals; Cebus/psychology; Choice Behavior; Cooperative Behavior; Female; Male; *Reward; Social Justice
	Abstract	During the evolution of cooperation it may have become critical for individuals to compare their own efforts and pay-offs with those of others. Negative reactions may occur when expectations are violated. One theory proposes that aversion to inequity can explain human cooperation within the bounds of the rational choice model, and may in fact be more inclusive than previous explanations. Although there exists substantial cultural variation in its particulars, this 'sense of fairness' is probably a human universal that has been shown to prevail in a wide variety of circumstances. However, we are not the only cooperative animals, hence inequity aversion may not be uniquely human. Many highly cooperative nonhuman species seem guided by a set of expectations about the outcome of cooperation and the division of resources. Here we demonstrate that a nonhuman primate, the brown capuchin monkey (Cebus apella), responds negatively to unequal reward distribution in exchanges with a human experimenter. Monkeys refused to participate if they witnessed a conspecific obtain a more attractive reward for equal effort, an effect amplified if the partner received such a reward without any effort at all. These reactions support an early evolutionary origin of inequity aversion.
	Address	Living Links, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia 30329, USA. sbrosna@emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1476-4687	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:13679918			Approved	no
	Call Number	refbase @ user @			Serial	179
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	Author	de Waal, F.B.; Berger, M.L.
	Title	Payment for labour in monkeys			Type	Journal Article
	Year	2000	Publication	Nature	Abbreviated Journal	Nature
	Volume	404	Issue	6778	Pages	563
	Keywords	Animals; Cebus/physiology; Cooperative Behavior; Evolution; *Feeding Behavior; Female; Male; Reward
	Abstract
	Address	Living Links, Yerkes Regional Primate Research Center, and Department of Psychology, Emory University, Atlanta, Georgia 30329, USA. dewaal@emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0028-0836	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10766228			Approved	no
	Call Number	refbase @ user @			Serial	190
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	Author	Johnson, D.D.P.; Stopka, P.; Knights, S.
	Title	Sociology: The puzzle of human cooperation			Type	Journal Article
	Year	2003	Publication	Nature	Abbreviated Journal	Nature
	Volume	421	Issue	6926	Pages	911-2; discussion 912
	Keywords	Altruism; Cooperative Behavior; Evolution; Humans; Models, Biological; Punishment; Reward; Risk
	Abstract
	Address	Olin Institute for Strategic Studies, Harvard University, Cambridge, Massachusetts 02138, USA. dominic@post.harvard.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0028-0836	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12606989			Approved	no
	Call Number	refbase @ user @			Serial	467
Permanent link to this record



	Author	Brosnan, S.F.; de Waal, F.B.M.
	Title	Responses to a simple barter task in chimpanzees, Pan troglodytes			Type	Journal Article
	Year	2005	Publication	Primates	Abbreviated Journal	Primates
	Volume	46	Issue	3	Pages	173-182
	Keywords	Analysis of Variance; Animals; Choice Behavior/physiology; Conditioning (Psychology)/physiology; Learning/physiology; Pan troglodytes/physiology; Reward; Sex Factors; Social Behavior; *Token Economy
	Abstract	Chimpanzees (Pan troglodytes) frequently participate in social exchange involving multiple goods and services of variable value, yet they have not been tested in a formalized situation to see whether they can barter using multiple tokens and rewards. We set up a simple barter economy with two tokens and two associated rewards and tested chimpanzees on their ability to obtain rewards by returning the matching token in situations in which their access to tokens was unlimited or limited. Chimpanzees easily learned to associate value with the tokens, as expected, and did barter, but followed a simple strategy of favoring the higher-value token, regardless of the reward proffered, instead of a more complex but more effective strategy of returning the token that matched the reward. This response is similar to that shown by capuchin monkeys in our previous study. We speculate that this response, while not ideal, may be sufficient to allow for stability of the social exchange system in these primates, and that the importance of social barter to both species may have led to this convergence of strategies.
	Address	Living Links, Yerkes National Primate Research Center, Atlanta, GA 30322, USA. sbrosna@emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0032-8332	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15824938			Approved	no
	Call Number	refbase @ user @			Serial	167
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