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Benson-Amram, S., Weldele, M. L., & Holekamp, K. E. (2013). A comparison of innovative problem-solving abilities between wild and captive spotted hyaenas, Crocuta crocuta. Animal Behaviour, 85(2), 349–356.
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Smith, J. E., Kolowski, J. M., Graham, K. E., Dawes, S. E., & Holekamp, K. E. (2008). Social and ecological determinants of fission-fusion dynamics in the spotted hyaena. Anim. Behav., 76(3), 619–636.
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