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Hunt, G. R., Gray R.D., & Taylor, A. H. (2013). Why is tool use rare in animals? (Boesch C C. J. anz C, Ed.). Cambridge, MA.: Cambridge University Press.
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Krueger, K., Gruentjens, T., & Hempel, E. (2023). Wolf contact in horses at permanent pasture in Germany. Plos One, 18(8), e0289767.
Abstract: Wolves returned to Germany in 2000, leading to fear in German horse owners that their horses could be in danger of wolf attacks or panic-like escapes from pastures when sighting wolves. However, reports from southern European countries indicate that wolf predation on horses diminishes with increasing presence of wildlife. Therefore, we conducted a long-term, filed observation between January 2015 and July 2022 on 13 non breeding riding horses, mares and geldings, kept permanently on two pastures within the range of wildlife and a stable wolf pack with annual offspring. Wildlife cameras at the fences of the pastures made 984 times recordings of wolves and 3151 times recordings of wildlife in and around the pastures. Between 1 January 2022 and 23 March 2022 we observed two stable horse groups. Pasture 1 was grazed by five horses of mixed breed, four mares and one gelding, with the median age of 8 years (min. = 6y, max. = 29y). Pasture 2 was grazed by eight heavy warmbloods and draught horses, three mares and five geldings, with the median age of 16 years (min. = 13y, max. = 22y). During this period no wolf was recorded at pasture 2, but wild boar several times, whereas at pasture 1, wolves were recorded 89 times, and for the wildlife mostly hare. Wolves may have avoided pasture 2 because of the presence of wild boar or because the large group of older, heavy breed horses may have formed a stable, protective group. The latter needs to be confirmed in a follow-up field observation, which records anti-predator behavior and welfare indicators in horses. In conclusion, wolves did not attack the mature horses on pastures with plenty of wildlife and the horses did not respond to the presence of wolves with visible signs of reduced welfare or panic. This indicates that wolves may prefer to prey on easily accessible wildlife around and at horse pastures and that Central European horses become accustom to the presence of non-hunting wolves.
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Capitani, C., Chynoweth, M., Kusak, J., Çoban, E., & Sekercioglu, Ç. H. (2016). Wolf diet in an agricultural landscape of north-eastern Turkey. Mammalia, 80(3), 329–334.
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Reinhardt, I., Kluth, G., Balzer, S., & Steyer, K. (2022). Wolfsverursachte Schäden, Präventions- und Ausgleichszahlungen in Deutschland 2021 (Markus Ritz, Ed.) (Vol. 41). Görlitz, Deutschland: DBBW-Dokumentations- und Beratungsstelle des Bundes zum Thema Wolf.
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Wotschikowsky, U. (2007). Wölfe und Jäger in der Oberlausitz. Broschüre, Freundeskreis freilebender Wölfe, .
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Wallner, B., Palmieri, N., Vogl, C., Rigler, D., Bozlak, E., Druml, T., et al. (2017). Y Chromosome Uncovers the Recent Oriental Origin of Modern Stallions. Current Biology, 27(13), 2029–2035.e5.
Abstract: The Y chromosome directly reflects male genealogies, but the extremely low Y chromosome sequence diversity in horses has prevented the reconstruction of stallion genealogies [1, 2]. Here, we resolve the first Y chromosome genealogy of modern horses by screening 1.46 Mb of the male-specific region of the Y chromosome (MSY) in 52 horses from 21 breeds. Based on highly accurate pedigree data, we estimated the de novo mutation rate of the horse MSY and showed that various modern horse Y chromosome lineages split much later than the domestication of the species. Apart from few private northern European haplotypes, all modern horse breeds clustered together in a roughly 700-year-old haplogroup that was transmitted to Europe by the import of Oriental stallions. The Oriental horse group consisted of two major subclades: the Original Arabian lineage and the Turkoman horse lineage. We show that the English Thoroughbred MSY was derived from the Turkoman lineage and that English Thoroughbred sires are largely responsible for the predominance of this haplotype in modern horses.
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Holzapfel, M., Wagner, C., & Kluth, G. et al. (2011). Zur Nahrungsökologie der Wölfe (Canis lupus) in Deutschland. Beiträge zur Jagd- und Wildforschung, 36, 117–128.
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