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().9(5), 265.
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Hendriksen, P., Elmgreen, K., & Ladewig, J. (2011). Trailer-loading of horses: Is there a difference between positive and negative reinforcement concerning effectiveness and stress-related signs? J. Vet. Behav., 6(5), 261–266.
Abstract: The traditional way to train horses is by the application of negative reinforcement (NR). In the past few years, however, the use of positive reinforcement (PR) has become more common. To evaluate the effectiveness and the possible stressor effect of the 2 training methods, 12 horses showing severe trailer-loading problems were selected and exposed to trailer-loading. They were randomly assigned to one of the 2 methods. NR consisted of various degrees of pressure (lead rope pulling, whip tapping). Pressure was removed as soon as the horse complied. PR horses were exposed to clicker training and taught to follow a target into the trailer. Heart rate (HR) was recorded every 5 seconds and behavior denoting discomfort was observed using one-zero sampling with 10 seconds sampling intervals. Training was completed when the horse could enter the trailer upon a signal, or was terminated after a maximum of 15 sessions. Of the 12 horses, 10 reached the criterion within the 15 sessions. One horse was eliminated from the study because of illness and 1 PR horse failed to enter the trailer. A Mann-Whitney U-test indicated that the horses trained with NR displayed significantly more discomfort behavior per training session than horses trained with PR (NR: 13.26 ± 3.25; PR: 3.17 ± 8.93, P < 0.0001) and that horses in the PR group spent less time (second) per session to complete the training criterion (NR: 672.9 ± 247.12; PR: 539.81 ± 166.37, P < 0.01). A Mann-Whitney U-test showed that no difference existed in mean HR (bpm) between the 2 groups (NR: 53.06 ± 11.73 bpm; PR: 55.54 ± 6.7 bpm, P > 0.05), but a Wilcoxon test showed a difference in the PR group between the baseline of HR and mean HR obtained during training sessions (baseline PR: 43 ± 8.83 bpm; PR: 55.54 ± 6.7 bpm, P < 0.05). In conclusion, the PR group provided the fastest training solution and expressed less stress response. Thus, the PR procedure could provide a preferable training solution when training horses in potentially stressing situations.
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Amici, F., Widdig, A., Lehmann, J., & Majolo, B. (2019). A meta-analysis of interindividual differences in innovation. Anim. Behav., 155, 257–268.
Abstract: The ability to innovate and the social transmission of innovations have played a central role in human evolution. However, innovation is also crucial for other animals, by allowing them to cope with novel socioecological challenges. Although innovation plays such a central role in animals' lives, we still do not know the conditions required for innovative behaviour to emerge. Here, we focused on interindividual differences in innovation by (1) extensively reviewing existing literature on innovative behaviour in animals and (2) quantitatively testing the different evolutionary hypotheses that have been proposed to explain interindividual variation in innovation propensity during foraging tasks. We ran a series of phylogenetically controlled mixed-effects meta-regression models to determine which hypotheses (if any) are supported by currently available empirical studies. Our analyses show that innovation is more common in individuals that are older and belong to the larger sex, but also in more neophilic and/or explorative individuals. Moreover, these effects change depending on the study setting (i.e. wild versus captive). Our results provide no clear support to the excess of energy or the bad competitor hypotheses and suggest that study setting and interindividual differences in traits related to personality are also important predictors of innovation.
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Lorenz, K. (1943). Die angeborenen Formen möglicher Erfahrung. Zeitschrift für Tierpsychologie, 5(2), 235–409.
Abstract: Zusammenfassung Die vorliegende Abhandlung entspringt dem Aufgabenkreis einer jungen Arbeitsgemeinschaft zwischen Geisteswissenschaft und vergleichender Psychologie, die das wichtigste Programm des neugegründeten Philosophischen Institutes der Albertus-Universität in Königsberg darstellt. Sie ist aus der einen Abteilung dieser Anstalt, dem Institut für vergleichende Psychologie hervorgegangen. Obwohl sie eine ganze Reihe unveröffentlichter neuer Beobachtungen und Versuche enthält, stehen diese nicht genug im Mittelpunkte der Untersuchung, um eine Zusammenfassung von Ergebnissen in jener Form möglich zu machen, wie sie sonst in der induktiven Naturforschung üblich ist. Immerhin aber herrscht die induktive Denkweise in der vorliegenden Arbeit so stark vor, daß eine kurze Zusammenfassung angebracht erscbeint, wenn sie auch notgedrungen die Form einer kurzen Inhaltsangabe annehmea muß.
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von Borstel, U. U., Duncan, I. J. H., Shoveller, A. K., Merkies, K., Keeling, L. J., & Millman, S. T. (2009). Impact of riding in a coercively obtained Rollkur posture on welfare and fear of performance horses. Appl. Anim. Behav. Sci., 116(2-4), 228–236.
Abstract: Rollkur, the usually coercively obtained hyperflexion of the horse's neck, is employed as a training method by some dressage riders; however, its use is controversial as it may cause discomfort and adversely affect the horse's welfare. The objectives of this study were to determine: (1) if horses showed differences in stress, discomfort and fear responses as measured by heart rate and behaviour when ridden in Rollkur (R) obtained by pressure on the reins compared to regular poll flexion (i.e. with the nose-line being at or just in front of the vertical; N), and (2) if they showed a preference between the two riding styles when given the choice. Fifteen riding horses were ridden 30 times through a Y-maze randomly alternating between sides. Riding through one arm of the Y-maze was always followed by a short round ridden in R, whereas riding through the other arm was followed by a short round ridden in N. Immediately after the conditioning phase, horses were again repeatedly ridden into the maze; however, riders left it to the horse to decide which arm of the maze to enter. During R, horses moved slower and showed more often behavioural signs of discomfort, such as tail-swishing, head-tossing or attempted bucks (P < 0.05), and 14 of the 15 horses chose significantly (P < 0.05) more often the maze-arm associated with N rather than R. Subsequently, eight of the horses were also subjected to two fear tests following a short ride in N as well as a ride in R. During R, horses tended to react stronger (P = 0.092) to the fear stimuli and to take longer (P = 0.087) to approach them. These findings indicate that a coercively obtained Rollkur position may be uncomfortable for horses and that it makes them more fearful and therefore potentially more dangerous to ride. Further studies need to assess horses' reaction to gradual training of Rollkur, as opposed to a coercively obtained hyperflexion, in order to decide whether the practice should be banned.
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Baragli, P., Vitale, V., Paoletti, E., Sighieri, C., & Reddon, A. R. (2011). Detour behaviour in horses (Equus caballus). J. Ethol., 29(2), 227–234.
Abstract: The objective of this study was to investigate the ability of horses (Equus caballus) to detour around symmetric and asymmetric obstacles. Ten female Italian saddle horses were each used in three detour tasks. In the first task, the ability to detour around a symmetrical obstacle was evaluated; in the second and third tasks subjects were required to perform a detour around an asymmetrical obstacle with two different degrees of asymmetry. The direction chosen to move around the obstacle and time required to make the detour were recorded. The results suggest that horses have the spatial abilities required to perform detour tasks with both symmetric and asymmetric obstacles. The strategy used to perform the task varied between subjects. For five horses, lateralized behaviour was observed when detouring the obstacle; this was consistently in one direction (three on the left and two on the right). For these horses, no evidence of spatial learning or reasoning was found. The other five horses did not solve this task in a lateralized manner, and a trend towards decreasing lateralization was observed as asymmetry, and hence task difficulty, increased. These non-lateralized horses may have higher spatial reasoning abilities.
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Irving-Pease, E. K., Ryan, H., Jamieson, A., Dimopoulos, E. A., Larson, G., & Frantz, L. A. F. (2019). Paleogenomics of Animal Domestication. In C. Lindqvist, & O. P. Rajora (Eds.), Paleogenomics: Genome-Scale Analysis of Ancient DNA (pp. 225–272). Cham: Springer International Publishing.
Abstract: Starting with dogs, over 15,000 years ago, the domestication of animals has been central in the development of modern societies. Because of its importance for a range of disciplines – including archaeology, biology and the humanities – domestication has been studied extensively. This chapter reviews how the field of paleogenomics has revolutionised, and will continue to revolutionise, our understanding of animal domestication. We discuss how the recovery of ancient DNA from archaeological remains is allowing researchers to overcome inherent shortcomings arising from the analysis of modern DNA alone. In particular, we show how DNA, extracted from ancient substrates, has proven to be a crucial source of information to reconstruct the geographic and temporal origin of domestic species. We also discuss how ancient DNA is being used by geneticists and archaeologists to directly observe evolutionary changes linked to artificial and natural selection to generate a richer understanding of this fascinating process.
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Byrne R.W. (1994). The evolution of intelligence. In P.J.B. Slater and T.R. Halliday (Ed.), Behaviour and Evolution (pp. 223–265). Cambridge,UK: Cambridge University Press.
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Genov, P. W., & Kostava, V. (1993). Untersuchungen zur zahlenmäßigen Stärke des Wolfes und seiner Einwirkung auf die Haustierbestände in Bulgarien. Zeitschrift für Jagdwissenschaft, 39(4), 217–223.
Abstract: Die Untersuchung wurde in der Zeitspanne von 1984 bis 1988 durchgeführt. Es wurden die Protokolle des Staatlichen Versicherungsinstituts benutzt, die Angaben für Raubüberfälle von Wölfen auf Haustiere beinhalten (Tabelle 1). Außerdem wurden Angaben über die während dieser Zeitspanne erlegten Wölfe zusammengefaßt. Die Abschußzahlen lauten: 1984 – 163, 1985 – 147, 1986 – 179, 1987 – 211 und 1988 – 220 Tiere. Die Anzahl der in den einzelnen Gebirgen lebenden Wölfe wurde nach einer Umfrage festgestellt. Für die in Betracht kommenden Gebirge werden folgende Bestandszahlen angenommen: Rhodopen -- 60-80 Individuen, 189 bis 264 km2 pro Tier, Rila- und Piringebirge -- 60-80 Tiere, 109 bis 145 km2 pro Tier, Ossogowo-Belassiza Gebirgssystem -- 40-50 Individuen, 57-70 km2 pro Tier, West- und Mittelbalkan -- 35-38 Wölfe, 200 km2 pro Tier. Dazu kommen noch 10-15 Wölfe im Flußbecken von Beli Lom und etwa 20 Exemplare in Strandscha- und Sakargebirge. Insgesamt lebten in Bulgarien im Jahre 1988 etwa 260-330 Wölfe (Abb. 1).
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Christensen, J. W., Søndergaard, E., Thodberg, K., & Halekoh, U. (2011). Effects of repeated regrouping on horse behaviour and injuries. Appl. Anim. Behav. Sci., 133(3), 199–206.
Abstract: Domestic horses are faced with social challenges throughout their lives due to limitations in social contact, space restrictions and frequent changes in social companionship. This is in contrast to natural conditions where horses live in relatively stable harem bands. Currently, little is known about how repeated regrouping affect horse behaviour and welfare, and it is unknown whether horses may adapt to regrouping. In this study, we aimed to investigate the effects of an unstable group structure, caused by weekly regroupings, on behaviour and frequency of injuries in young horses. Forty-five horses were included in the study and were randomly assigned to the treatments; Stable (S; seven groups of three horses) or Unstable (U; eight groups of three horses). The experimental period lasted 7 weeks, during which horses in Stable groups remained in the same group, whereas one horse was exchanged between Unstable groups every week. The groups were kept in 80m×80m grass-covered enclosures and were fed additional roughage on the ground daily. Social interactions were recorded in Unstable groups immediately after each regrouping (30min), and in both Stable and Unstable groups on day 1, 3 and 6 after each regrouping (2×20min/group/day). Injuries were scored by the end of the experimental period. The level of aggression shown by horses in Unstable groups immediately after regrouping was not affected by week (F5,35=0.42, P=0.83), indicating that horses neither habituated, nor sensitized, to repeated regrouping. Compared to horses in Stable groups, more agonistic behaviour was shown by horses in Unstable groups (i.e. non-contact agonistic; F1,65=5.60, P=0.02), whereas there was no treatment effect on other variables. The level of play behaviour appeared, however, to be more variable in Unstable groups. There was a significant effect of week on the level of contact agonistic interactions as well as greeting behaviour, due to a high occurrence in weeks 4-6. Non-contact agonistic interactions constituted the major part of agonistic interactions (66%). Possibly as consequence, no serious injuries were registered and there was no treatment effect (U=184; P=0.11). We conclude that the behaviour of young horses is affected by group management, and that horses appear not to adapt to weekly regroupings.
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