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Drevemo, S., Fredricson, I., Hjertén, G., & McMIKEN, D. (1987). Early development of gait asymmetries in trotting Standardbred colts. Equine. Vet. J., 19(3), 189–191.
Abstract: Summary Ten trotting Standardbred colts were recorded by high-speed cinematography at the ages of eight, 12 and 18 months. The horses were trotting on a treadmill operating at 4.0 m/secs. Five horses were subjected to a programme of intensified training from eight months of age, whereas the others were not trained and acted as controls. The films were analysed on a semi-automatic film-reading equipment and a number of variables used to demonstrate the gait symmetry were calculated and scaled by computer. Certain differences between left and right diagonal and contralateral pair of limbs, respectively, were noted, suggesting that laterality in horses may be inherited. The most pronounced systematic differences were found in 18-month old horses in the trained group. The results show the importance of careful gait examination and comprehensive coordination training at an early age.
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Goodwin, D. (1999). The importance of ethology in understanding the behaviour of the horse. Equine Veterinary Journal, 31(S28), 15–19.
Abstract: Summary Domestication has provided the horse with food, shelter, veterinary care and protection, allowing individuals an increased chance of survival. However, the restriction of movement, limited breeding opportunities and a requirement to expend energy, for the benefit of another species, conflict with the evolutionary processes which shaped the behaviour of its predecessors. The behaviour of the horse is defined by its niche as a social prey species but many of the traits which ensured the survival of its ancestors are difficult to accommodate in the domestic environment. There has been a long association between horses and man and many features of equine behaviour suggest a predisposition to interspecific cooperation. However, the importance of dominance in human understanding of social systems has tended to overemphasise its importance in the human-horse relationship. The evolving horse-human relationship from predation to companionship, has resulted in serial conflicts of interest for equine and human participants. Only by understanding the nature and origin of these conflicts can ethologists encourage equine management practices which minimise deleterious effects on the behaviour of the horse.
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Gomez Alvarez, C. B., Rhodin, M., Bobber, M. F., Meyer, H., Weishaupt, M. A., Johnston, C., et al. (2006). The effect of head and neck position on the thoracolumbar kinematics in the unridden horse. Equine Vet J Suppl, (36), 445–451.
Abstract: REASONS FOR PERFORMING STUDY: In many equestrian activities a specific position of head and/or neck is required that is dissimilar to the natural position. There is controversy about the effects of these positions on locomotion pattern, but few quantitative data are available. OBJECTIVES: To quantify the effects of 5 different head and neck positions on thoracolumbar kinematics of the horse. METHODS: Kinematics of 7 high level dressage horses were measured walking and trotting on an instrumented treadmill with the head and neck in the following positions: HNP2 = neck raised, bridge of the nose in front of the vertical; HNP3 = as HNP2 with bridge of the nose behind the vertical; HNP4 = head and neck lowered, nose behind the vertical; HNP5 = head and neck in extreme high position; HNP6 = head and neck forward and downward. HNP1 was a speed-matched control (head and neck unrestrained). RESULTS: The head and neck positions affected only the flexion-extension motion. The positions in which the neck was extended (HNP2, 3, 5) increased extension in the anterior thoracic region, but increased flexion in the posterior thoracic and lumbar region. For HNP4 the pattern was the opposite. Positions 2, 3 and 5 reduced the flexion-extension range of motion (ROM) while HNP4 increased it. HNP5 was the only position that negatively affected intravertebral pattern symmetry and reduced hindlimb protraction. The stride length was significantly reduced at walk in positions 2, 3, 4 and 5. CONCLUSIONS: There is a significant influence of head/neck position on back kinematics. Elevated head and neck induce extension in the thoracic region and flexion in the lumbar region; besides reducing the sagittal range of motion. Lowered head and neck produces the opposite. A very high position of the head and neck seems to disturb normal kinematics. POTENTIAL RELEVANCE: This study provides quantitative data on the effect of head/neck positions on thoracolumbar motion and may help in discussions on the ethical acceptability of some training methods.
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Weishaupt, M. A., Wiestner, T., von Peinen, K., Waldern, N., Roepstorff, L., van Weeren, R., et al. (2006). Effect of head and neck position on vertical ground reaction forces and interlimb coordination in the dressage horse ridden at walk and trot on a treadmill. Equine Vet J Suppl, (36), 387–392.
Abstract: REASONS FOR PERFORMING STUDY: Little is known in quantitative terms about the influence of different head-neck positions (HNPs) on the loading pattern of the locomotor apparatus. Therefore it is difficult to predict whether a specific riding technique is beneficial for the horse or if it may increase the risk for injury. OBJECTIVE: To improve the understanding of forelimb-hindlimb balance and its underlying temporal changes in relation to different head and neck positions. METHODS: Vertical ground reaction force and time parameters of each limb were measured in 7 high level dressage horses while being ridden at walk and trot on an instrumented treadmill in 6 predetermined HNPs: HNP1 – free, unrestrained with loose reins; HNP2 – neck raised, bridge of the nose in front of the vertical; HNP3 – neck raised, bridge of the nose behind the vertical; HNP4 – neck lowered and flexed, bridge of the nose considerably behind the vertical; HNP5 – neck extremely elevated and bridge of the nose considerably in front of the vertical; HNP6 – neck and head extended forward and downward. Positions were judged by a qualified dressage judge. HNPs were assessed by comparing the data to a velocity-matched reference HNP (HNP2). Differences were tested using paired t test or Wilcoxon signed rank test (P<0.05). RESULTS: At the walk, stride duration and overreach distance increased in HNP1, but decreased in HNP3 and HNP5. Stride impulse was shifted to the forehand in HNP1 and HNP6, but shifted to the hindquarters in HNP5. At the trot, stride duration increased in HNP4 and HNP5. Overreach distance was shorter in HNP4. Stride impulse shifted to the hindquarters in HNP5. In HNP1 peak forces decreased in the forelimbs; in HNP5 peak forces increased in fore- and hindlimbs. CONCLUSIONS: HNP5 had the biggest impact on limb timing and load distribution and behaved inversely to HNP1 and HNP6. Shortening of forelimb stance duration in HNP5 increased peak forces although the percentage of stride impulse carried by the forelimbs decreased. POTENTIAL RELEVANCE: An extremely high HNP affects functionality much more than an extremely low neck.
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Krange, O., & Skogen, K. (2011). When the lads go hunting: The 'Hammertown mechanism' and the conflict over wolves in Norway. Ethnography, 12(4), 466–489.
Abstract: Rural communities are changing. Depopulation and unemployment is accompanied by the advance of new perspectives on nature, where protection trumps resource extraction. These developments are perceived as threatening by rural working-class people with close ties to traditional land use ? a situation they often meet with cultural resistance. Cultural resistance is not necessarily launched against institutionalized power, nor does it necessarily imply a desire for fundamental social change. It should rather be seen as a struggle for autonomy. However, autonomy does not entail influence outside the cultural realm. Struggles to uphold traditional rural lifestyles ? for example by denouncing the current nature conservation regime ? could be understood in much the same conceptual framework as Willis employed in ?Learning to labour?. Based on an ethnographic study of the conflicts over wolf protection, we demonstrate that ?the Hammertown mechanism? is of a more general nature than often implied in the discussion of Willis? work.
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Zaccaroni, M., Passilongo, D., Buccianti, A., Dessi-Fulgheri, F., Facchini, C., & Gazzola, A. (2012). Group specific vocal signature in free- ranging wolf packs. Ethol Ecol Evol, 24.
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Rutberg, A. T. (1987). Horse Fly Harassment and the Social Behavior of Feral Ponies. Ethology, 75(2), 145–154.
Abstract: Abstract Horse flies (Tabanidae) on and around feral ponies in harem groups were counted at Assateague Island National Seashore, Maryland, U.S.A., between June and August 1985. Harem stallions attracted the most flies; adult mares showed intermediate fly numbers, while few flies landed on foals under any circumstances. The use of thermal and chemical cues by flies selecting a host may have helped create this disparity. When flies were abundant, ponies reduced spacing within the group. Ponies in larger groups suffered from fewer flies than ponies in smaller groups. There was, however, no evidence that ponies merged into larger groups in response to fly harassment, suggesting that biting flies play little role in structuring pony social organization.
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Meriggi, A., Dagradi, V., Dondina, O., Perversi, M., Milanesi, P., Lombardini, M., et al. (2014). Short-term responses of wolf feeding habits to changes of wild and domestic ungulate abundance in Northern Italy. Ethology Ecology & Evolution, 27(4), 389–411.
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Iliopoulos, Y., Youlatos, D., & Sgardelis, S. (2013). Wolf pack rendezvous site selection in Greece is mainly affected by anthropogenic landscape features. Eur J Wildl Res, 60.
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Galaverni, M., Palumbo, D., Fabbri, E., Caniglia, R., Greco, C., & Randi, E. (2012). Monitoring wolves (Canis lupus) by non-invasive genetics and camera trapping: A small-scale pilot study. Eur J Wildl Res, 58.
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