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Lloyd, A. S., Martin, J. E., Bornett-Gauci, H. L. I., & Wilkinson, R. G. (2008). Horse personality: Variation between breeds. Appl. Anim. Behav. Sci., 112(3-4), 369–383.
Abstract: Anecdotal evidence from horse owners and handlers suggests the existence of breed typical behaviour and personality in horses. This is further supported by current research on heritability of personality characteristics in a range of species. The Horse Personality Questionnaire (HPQ) is a 25-item rating method that has previously been shown to be reliable for the assessment of personality in horses. Principal component analysis on HPQ data has identified six underlying personality components in horses. These are Dominance, Anxiousness, Excitability, Protection, Sociability and Inquisitiveness. Using the HPQ a survey of 1223 horses of eight different breeds was completed. Data were analysed to explore any differences in personality between breeds across the six personality components. Breed differences in personality were identified, and it was noted that variability between breeds varied between personality components. Anxiousness and Excitability showed the most variation between breeds, whilst Dominance and Protection showed the least variance. The results identified breed typical personalities that were comparable to results from previous studies as well as anecdotal evidence provided by the popular equine literature. The results are discussed in terms of the potential selection pressures that may have resulted in these differences.
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Lloyd, A. S., Martin, J. E., Bornett-Gauci, H. L. I., & Wilkinson, R. G. (2007). Evaluation of a novel method of horse personality assessment: Rater-agreement and links to behaviour. Appl. Anim. Behav. Sci., 105(1-3), 205–222.
Abstract: The efficacy of questionnaire-based personality assessment has been shown in a variety of animal and human personality studies. There has been a recent increase in questionnaire-based studies focussing on equine personality but with a lack of comparability to studies on other species. The aim of this study was to test the reliability of an assessment method originally developed for primates and demonstrate reliability using three criteria (1) assessments by independent observers must agree with one another, (2) these assessments must predict behaviours and real-world outcomes and (3) observer ratings must be shown to reflect genuine attributes of the individuals rated, not merely the observer's implicit personality theories about how traits co-vary. The personality of 61 horses (Equus caballus) was assessed using a questionnaire constructed of 30 behaviourally defined adjectives (BDAs). Horses were each assessed by three judges, in addition to a total of 2 h behaviour observations recorded per horse. Rater agreement was demonstrated for 72.1% of the horses and 25 of the BDAs. Principal component analysis was carried out on the rating data and revealed six underlying personality components that were labelled “dominance”, “anxiousness”, “excitability”, “protection”, “sociability” and “inquisitiveness”. Component scores for horses were correlated against behavioural observations for the same horses and revealed significant correlations with 20 of the recorded behaviour variables. Correlations between specific components and their associated behaviours were logical and consistent with the types of behaviours that would be expected to be linked with such personality types. The data were shown to meet all three criteria and provided strong evidence that the assessment method was reliably measuring horse personality.
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Martin, T. I., Zentall, T. R., & Lawrence, L. (2006). Simple discrimination reversals in the domestic horse (Equus caballus): Effect of discriminative stimulus modality on learning to learn. Appl. Anim. Behav. Sci., 101(3-4), 328–338.
Abstract: The cognitive capacity of an organism, relative to that of other species, can be assessed by using a relative measure of learning. One such measure is the ability of an organism to learn about the reversal of a discrimination. The present study compared the performance of two groups of horses on a simple discrimination reversal task when the only difference between the groups was the modality of the relevant cue. For the visual group (absence or presence of a light), the spatial position was irrelevant. For the spatial group, a spatial cue (left/right) was available and the visual cue was irrelevant. Horses in the spatial group learned the original discrimination and six reversals; they also showed evidence of learning to learn. Horses in the visual group did not reach criterion during the study. As a result, there was no evidence of learning to learn.
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Chase, I. D., Tovey, C., Spangler-Martin, D., & Manfredonia, M. (2002). Individual differences versus social dynamics in the formation of animal dominance hierarchies. Proc. Natl. Acad. Sci. U.S.A., 99(8), 5744–5749.
Abstract: Linear hierarchies, the classical pecking-order structures, are formed readily in both nature and the laboratory in a great range of species including humans. However, the probability of getting linear structures by chance alone is quite low. In this paper we investigate the two hypotheses that are proposed most often to explain linear hierarchies: they are predetermined by differences in the attributes of animals, or they are produced by the dynamics of social interaction, i.e., they are self-organizing. We evaluate these hypotheses using cichlid fish as model animals, and although differences in attributes play a significant part, we find that social interaction is necessary for high proportions of groups with linear hierarchies. Our results suggest that dominance hierarchy formation is a much richer and more complex phenomenon than previously thought, and we explore the implications of these results for evolutionary biology, the social sciences, and the use of animal models in understanding human social organization.
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Bayley, P., Martin, S., & Anson, M. (1975). Temperature-jump circular dichroism: observation of chiroptical relaxation processes at millisecond time resolution. Biochem Biophys Res Commun, 66(1), 303–308.
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Herron, M. A., Martin, J. E., & Joyce, J. R. (1978). Quantitative study of the decussating optic axons in the pony, cow, sheep, and pig. Am. J. Vet .Res., 39(7), 1137–9.
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Gutierrez Rincon, J. A., Vives Turco, J., Muro Martinez, I., & Casas Vaque, I. (1992). A comparative study of the metabolic effort expended by horse riders during a jumping competition. Br J Sports Med, 26(1), 33–35.
Abstract: The three main Olympic horse riding disciplines are dressage, jumping, and three-day eventing (including dressage, cross country and jumping). In the jumping discipline (obstacle race), the 'team' (horse rider) is judged under the different conditions that might take place in a varied run. The horse is expected to show power and ability; the rider must show riding skill and good physical condition. However, the different conditions encountered by the rider during competition (duration of event, continuous isometric working level, especially in the inferior trunk, lead us to consider the need for a rider to develop different metabolic pathways to meet the high energy requirements of the competition.
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Voigtlaender-Schnabel, S., Vogel, L., Greiner, B., Wiezorek, S., Schuette, P., Solmsen, E. - H., et al. (2022). Reactions of horses to wildlife and livestock guarding dogs. CDPNews, 24, 49–58.
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Solmsen, E. - H., Bathen, M., Grüntjens, T., Hempel, E., Klose, M., Krüger, K., et al. (2021). Protecting horses against wolves in Germany. CPDnews, 23, 12–19.
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Solmsen, E. - H., Bathen, M., Grüntjens, T., Hempel, E., Klose, M., Krüger, K., et al. (2021). Protecting horses against wolves in Germany. CDPNews, 23.
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