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Klingel, H. (). Das Verhalten der Pferde (Equidae). Handb. o. Zool., 8(10), 1–68.
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Klingel, H. (1964). Zur Sozialstruktur des Steppenzebras, Equus quagga boehmi Matschie. Die Naturwissenschaften, 51(14), 347.
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Klingel, H. (1967). Soziale Organisation und Verhalten freilebender Steppenzebras (Equus quagga). Z. Tierpsychol., 24(5), 580–624.
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Klingel, H. (1968). Sozial Organisation und Verhaltensweisen von Hartmann- und Bergzebras (Equus zebra hartmannae und E. z. zebra). Z. Tierpsychol., 25, 76–88.
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Klingel, H. (1975). Social organization and reproduction in equids. J Reprod Fertil Suppl, (23), 7–11.
Abstract: There are two distinct types of social organization and, accordingly, two types of mating systems in equids. In the horse, Plains zebra and Mountain zebra, the adults live in non-territorial and cohesive one-male groups and in stallion groups. The family stallions have exclusive mating rights which are respected by all others. In Grevy's zebra and in the African and Asiatic wild asses, the stallions are permanently territorial and have exclusive mating rights within their territories. Ecological and evolutionary aspects are discussed.
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Klingel, H. (1982). Social organization of feral horses. J Reprod Fertil Suppl, 32, 89–95.
Abstract: The basic social unit in feral horses is the family group consisting of one stallion, one to a few unrelated mares and their foals. Surplus stallions associate in bachelor groups. Stallions are instrumental in bringing mares together in a unit which then persists even without a stallion. The similarity of social organization in populations living in a variety of different habitats indicates that feral horses have reverted to the habits of their wild ancestors, and that domestication has had no influence on this basic behavioural feature.
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Klingel, H. (1989). Odd-toed Ungulates, Horses. In B. Grzimek (Ed.), Grzimek's Encyclopedia of Mammals (Vol. 4, 550+pp. 557–594). McGraw Hill.
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Klingel, H. (1990). Kalameili – future home of the Przewalski horse? In S. Seifert (Ed.), Proceedings of the 5th International Symposium on the Preservation of the Przewalski Horse, Leipzig (pp. 221–224). Leipzig: Zoolog Garten.
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Klingel, H. (1991). Dix ans parmi les zèbres. Terre Sauvage, 48, 34–43.
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Klingel, H. (1991). Tausend Zebras im Computer. Das Tier, 10, 8–16.
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