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Klingel, H. (1998). Observations on social organization and behaviour of African and Asiatic Wild Asses (Equus africanus and Equus hemionus). Appl Anim Behav Sci, 60(2), 103–113.
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Schulte, N., & Klingel, H. (1991). Herd Structure, Leadership, Dominance and Site Attachment of the Camel, Camelus Dromedarius. Behaviour, 118(1-2), 103–114.
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Klingel, H. (1991). Tausend Zebras im Computer. Das Tier, 10, 8–16.
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Klingel, H. (1964). Zur Sozialstruktur des Steppenzebras, Equus quagga boehmi Matschie. Die Naturwissenschaften, 51(14), 347.
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Klingel, H. (1989). Odd-toed Ungulates, Horses. In B. Grzimek (Ed.), Grzimek's Encyclopedia of Mammals (Vol. 4, 550+pp. 557–594). McGraw Hill.
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Klingel, H. (). Das Verhalten der Pferde (Equidae). Handb. o. Zool., 8(10), 1–68.
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Klingel, H. (1975). Social organization and reproduction in equids. J Reprod Fertil Suppl, (23), 7–11.
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Klingel, H. (1982). Social organization of feral horses. J Reprod Fertil Suppl, 32, 89–95.
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Klingel, H. (1990). Kalameili – future home of the Przewalski horse? In S. Seifert (Ed.), Proceedings of the 5th International Symposium on the Preservation of the Przewalski Horse, Leipzig (pp. 221–224). Leipzig: Zoolog Garten.
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Klingel, H. (1991). Dix ans parmi les zèbres. Terre Sauvage, 48, 34–43.
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