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Houpt, K. A. (2006). Why horse behaviour is important to the equine clinician. Equine Vet J, 38(5), 386–387.
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Houpt, K. A., & Smith, R. (1993). Animal behavior case of the month. J Am Vet Med Assoc, 203(3), 377–378.
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Sweeting, M. P., Houpt, C. E., & Houpt, K. A. (1985). Social Facilitation of Feeding and Time Budgets in Stabled Ponies. J. Anim Sci., 60(2), 369–374.
Abstract: Eight pairs of pony mares were observed. Members of a pair were housed in adjacent stalls and fed hay ad libitum. The behavior of both ponies was recorded simultaneously in the morning (1000 to 1200 h) and afternoon (1400 to 1600 h) for a total of 117 h. The time budget was: 70.1 {+/-} 8.6% eating; 17.8 {+/-} 7.4% standing (including stand rest, stand alert and stand nonajert); 5.2 {+/-} 7.0% pushing hay; 2.9 {+/-} 1.2% walking; 1.9 {+/-} 2.9% drinking; 1.3 {+/-} 1.1% self-grooming; .2 {+/-} .3% defecating; .06 {+/-} .1% chewing nonfood items; .06 {+/-} .03% urination; .06 {+/-} .1% licking salt; .07 {+/-} .1% pawing hay; .6 {+/-} .7% lying and .07 {+/-} .08% stretching the neck over the stall wall dividing the ponies. While eating, the ponies lifted their heads 25.4 {+/-} 11.0 times/h. In less than one-half of the occasions when urination or defecation was observed, the ponies walked away from the spot where they had been eating to eliminate. During one-half of the observations, visual contact between the ponies was prevented by a solid partition between the stalls. The ponies spent significantly more time standing nonalert when the partition prevented visual contact (12 {+/-} 7%) than when visual contact could take place (6 {+/-} 3%, P<.05). When fresh hay was supplied in the mornings, the ponies spent similar amounts of time eating whether visual contact was allowed or not, but in the afternoon significantly more time was spent feeding when visual contact was allowed (73 {+/-} 4%) than when it was not (60 {+/-} 7%). Less time was spent eating, in the absence of visual contact, despite the presence of auditory and olfactory contact. Apparently social facilitation is important in maintaining feeding behavior in ponies. N1 -
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Houpt, K. A., Eggleston, A., Kunkle, K., & Houpt, T. R. (2000). Effect of water restriction on equine behaviour and physiology. Equine Vet J, 32(4), 341–344.
Abstract: Six pregnant mares were used to determine what level of water restriction causes physiological and/or behavioural changes indicative of stress. Nonlegume hay was fed ad libitum. During the first week of restriction, 5 l water/100 kg bwt was available, during the second week 4 l/100 kg bwt and, during the third week, 3 l/100 kg bwt. Ad libitum water intake was 6.9 l/100 kg bwt; at 3 l/100 kg bwt water intake was 42% of this. Daily hay intake fell significantly with increasing water restriction from 12.9 +/- 0.75 kg to 8.3 +/- 0.54 kg; bodyweight fell significantly for a total loss of 48.5 +/- 8.3 kg in 3 weeks. Daily blood samples were analysed; osmolality rose significantly with increasing water restriction from 282 +/- 0.7 mosmols/kg to 293.3 +/- 0.8 mosmols/kg bwt, but plasma protein and PCV did not change significantly. Cortisol concentrations fell from 8.1 ng/ml to 6.4 ng/ml over the 3 week period. Aldosterone fell from 211.3 +/- 74.2 pg/ml to 92.5 +/- 27.5 pg/ml at the end of the first week. The behaviour of 4 of the 6 mares was recorded 24 h/day for the duration of the study. The only significant difference was in time spent eating, which decreased with increasing water restriction from 46 +/- 3% to 30 +/- 3%. It is concluded that water restriction to 4 l/100 kg bwt dehydrates pregnant mares and may diminish their welfare, but is not life- or pregnancy-threatening.
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Houpt, K. A. (1981). Equine behavior problems in relation to humane management. Int. J. Stud. Anim. Prob., 2(6), 329–337.
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Haag, E. L., Rudman, R., & Houpt, K. A. (1980). Avoidance, maze learning and social dominance in ponies. J. Anim. Sci., 50, 329–335.
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Houpt, K. A. (1990). Ingestive behavior. Vet Clin North Am Equine Pract, 6(2), 319–337.
Abstract: In summary, horses spend 60% or more of their time eating when grazing or when feed is available free choice. Grasses are their preferred food, but they supplement the grass with herbs and woody plants. Sweetened mixtures of oats and corn are the most preferred concentrate. Horses can increase or decrease the time spent eating and amount eaten to maintain caloric intake. Their intake is stimulated by drugs such as diazepam and by the presence of other horses. Horses stop eating when gastric osmolality increases; increases in plasma osmolality, protein, and glucose accompany digestion. Foals eat several times an hour and begin sampling solid food at the same time that their dam is eating. Several areas of particular importance to the equine industry have not been investigated. These areas include the effect of exercise on short- and long-term food intake and the influence of reproductive state on the feeding of mares.
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Houpt, K. A. (2002). Formation and dissolution of the mare-foal bond. Appl. Anim. Behav. Sci., 78(2-4), 319–328.
Abstract: The behavior of mares at foaling and their behavior during subsequent weekly brief separations from their foals was quantified. Thirteen multiparous pony mares were observed for the first 30 min after foaling. Activities directed toward the foal and toward the fetal membranes were recorded. There were 16+/-8 contacts with the fetal membranes, most in the first 10 min post-partum. There were 81+/-12 activities directed toward the foal. There was a downward trend in foal contacts over the first 30 min. Eight of the 13 mares were observed for the 30-60 min post-partum during which licking and touching the foal continued. Only one of the foals suckled in the first 30 min, but half had suckled by 60 min. Ten mares and foals were separated for 5 min each week for 9 weeks. Although mare responses (neighs and steps) decreased with age of the foal, the foals' responses increased from weeks 1 to 2 and then decreased with foal age. This indicates that foal attachment to the mare is not complete until it is 2 weeks old.
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Houpt, K. A., Law, K., & Martinisi, V. (1978). Dominance hierarchies in domestic horses. Appl. Animal. Ethol., 4(3), 273–283.
Abstract: Dominance hierarchies were studied in 11 herds of domestic horses and ponies (Equus caballus). A paired feeding test was utilized to establish the dominance--subordination relationship between each pair of animals in a herd. Aggressive actions, threats, bites, kicks and chases were also recorded. In small herds linear hierarchies were formed, but in large herds triangular relationships were observed. Aggression was correlated with dominance rank. Body weight, but not age, appear to affect rank in the equine hierarchy. Juvenile horses were more likely to share feed with each other than were adult horses and were usually subordinate to adult horses. The daughters of a dominant mare were dominant within their own herds.
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Virga, V., & Houpt, K. A. (2001). Prevalence of placentophagia in horses. Equine Vet J, 33(2), 208–210.
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