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Author Tomkins, L.M.; Williams, K.A.; Thomson, P.C.; McGreevy, P.D. url  doi
openurl 
  Title Sensory Jump Test as a measure of sensory (visual) lateralization in dogs (Canis familiaris) Type Journal Article
  Year 2010 Publication Journal of Veterinary Behavior Abbreviated Journal  
  Volume (up) 5 Issue 5 Pages 256-267  
  Keywords sensory lateralization; monocular vision; binocular vision; jump kinematics; dog  
  Abstract Sensory lateralization in dogs (n = 74) was investigated in this study using our innovation, the Sensory Jump Test. This required the modification of head halters to create three different ocular treatments (binocular, right, and left monocular vision) for eye preference assessment in a jumping task. Ten jumps were recorded as a jump set for each treatment. Measurements recorded included (i) launch and landing paws, (ii) type of jump, (iii) approach distance, (iv) clearance height of the forepaw, hindpaw, and the lowest part of the body to clear the jump, and (v) whether the jump was successful. Factors significantly associated with these jump outcomes included ocular treatment, jump set number, and replication number. Most notably, in the first jump set, findings indicated a left hemispheric dominance for the initial navigation of the Sensory Jump Test, as left monocular vision (LMV) compromised of jumping more than right monocular (RMV) and binocular vision, with a significantly reduced approach distance and forepaw clearance observed in dogs with LMV. However, by the third jump set, dogs undergoing LMV launched from a greater approach distance and with a higher clearance height, corresponding to an increase in success rate of the jump, in comparison with RMV and binocular vision dogs. A marginally non-significant RMV bias was observed for eye preference based on the laterality indices for approach distance (P = 0.060) and lowest body part clearance height (P = 0.067). A comparison between eye preference and launching or landing paws showed no association between these measures of sensory and motor laterality. To our knowledge, this is the first study to report on sensory lateralization in the dog, and furthermore, to compare both motor and sensory laterality in dogs.  
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  Series Volume Series Issue Edition  
  ISSN 1558-7878 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ S1558-7878(10)00019-5 Serial 5379  
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Author McGreevy, P.D.; Harman, A.; McLean, A.; Hawson, L. url  doi
openurl 
  Title Over-flexing the horse's neck: A modern equestrian obsession? Type Journal Article
  Year 2010 Publication Journal of Veterinary Behavior: Clinical Applications and Research Abbreviated Journal Journal of Veterinary Behavior: Clinical Applications and Research  
  Volume (up) 5 Issue 4 Pages 180-186  
  Keywords  
  Abstract We used an opportunistic review of photographs of different adult and juvenile horses walking, trotting, and cantering (n = 828) to compare the angle of the nasal plane relative to vertical in feral and domestic horses at liberty (n = 450) with ridden horses advertised in a popular Australian horse magazine (n = 378). We assumed that horses in advertisements were shown at, what was perceived by the vendors to be, their best. Of the ridden horses, 68% had their nasal plane behind the vertical. The mean angle of the unridden horses at walk, trot, and canter (30.7 ± 11.5; 27.3 ± 12.0; 25.5 ± 11.0) was significantly greater than those of the ridden horses (1.4 ± 14.1; ?5.1 ± ?11.1; 3.1 ± 15.4, P < 0.001). Surprisingly, unridden domestic horses showed greater angles than feral horses or domestic horses at liberty. We compared adult and juvenile horses in all 3 gaits and found no significant difference. Taken together, these findings demonstrate that the longitudinal neck flexion of the degree desirable by popular opinion in ridden horses is not a common feature of unridden horses moving naturally. Moreover, they suggest that advertised horses in our series are generally being ridden at odds with their natural carriage and contrary to the international rules of dressage (as published by the International Equestrian Federation). These findings are discussed against the backdrop of the established doctrine, which states that carrying a rider necessitates changes in longitudinal flexion, and in the context of the current debate around hyperflexion.  
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  Corporate Author Thesis  
  Publisher Elsevier Place of Publication Editor  
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  ISSN 1558-7878 ISBN Medium  
  Area Expedition Conference  
  Notes doi: 10.1016/j.jveb.2010.03.004 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6501  
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Author Huron, D. url  doi
openurl 
  Title Voice Denumerability of Homogeneous Timbres Type Journal Article
  Year 2010 Publication Music Percept Interdiscip J Abbreviated Journal  
  Volume (up) 6 Issue Pages  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Huron2010 Serial 6472  
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Author Maros, K.; Boross, B.; Kubinyi, E. url  doi
openurl 
  Title Approach and follow behaviour – possible indicators of the human-horse relationship Type Abstract
  Year 2010 Publication Interaction Studies Abbreviated Journal  
  Volume (up) 11 Issue 3 Pages 410-427  
  Keywords Approach; Follow; Human&#8211; Horse Interaction  
  Abstract The aim of our study was to analyze the behavioural responses of horses (N = 51) to familiar humans and to find factors that may affect these responses in three tests: (1) approach to, (2) standing beside, and (3) following the familiar person. We investigated the impacts of horse-related factors (gender and age) and human-related factors (type of work, housing management, amount of handling, number of handlers and training to follow).<br xmlns=“http://pub2web.metastore.ingenta.com/ns/”></br> Horses with one handler needed less time to approach the human than horses with more handlers. Standing beside the human correlated positively with following. Following was mainly affected by training.<br xmlns=“http://pub2web.metastore.ingenta.com/ns/”></br> According to our results, the number of handlers has an important effect on horses' responses to familiar humans, especially regarding approach and follow behaviour. However, following behaviour is fundamentally determined by training.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5728  
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Author Farmer, K.; Krueger, K.; Byrne, R. pdf  url
doi  openurl
  Title Visual laterality in the domestic horse (Equus caballus) interacting with humans Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue Pages 229-238  
  Keywords Horse – Laterality – Eye preference – Emotion – Vision  
  Abstract Most horses have a side on which they are easier to handle and a direction they favour when working on a circle, and recent studies have suggested a correlation between emotion and visual laterality when horses observe inanimate objects. As such lateralisation could provide important clues regarding the horse’s cognitive processes, we investigated whether horses also show laterality in association with people. We gave horses the choice of entering a chute to left or right, with and without the passive, non-interactive presence of a person unknown to them. The left eye was preferred for scanning under both conditions, but significantly more so when a person was present. Traditionally, riders handle horses only from the left, so we repeated the experiment with horses specifically trained on both sides. Again, there was a consistent preference for left eye scanning in the presence of a person, whether known to the horses or not. We also examined horses interacting with a person, using both traditionally and bilaterally trained horses. Both groups showed left eye preference for viewing the person, regardless of training and test procedure. For those horses tested under both passive and interactive conditions, the left eye was preferred significantly more during interaction. We suggest that most horses prefer to use their left eye for assessment and evaluation, and that there is an emotional aspect to the choice which may be positive or negative, depending on the circumstances. We believe these results have important practical implications and that emotional laterality should be taken into account in training methods.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4953  
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Author Proops, L.; McComb, K. url  doi
openurl 
  Title Attributing attention: the use of human-given cues by domestic horses (Equus caballus) Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue 2 Pages 197-205  
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  Abstract Abstract  Recent research has shown that domestic dogs are particularly good at determining the focus of human attention, often outperforming chimpanzees and hand-reared wolves. It has been suggested that the close evolutionary relationship between humans and dogs has led to the development of this ability; however, very few other domestic species have been studied. We tested the ability of 36 domestic horses to discriminate between an attentive and inattentive person in determining whom to approach for food. The cues provided were body orientation, head orientation or whether the experimenters’ eyes were open or closed. A fourth, mixed condition was included where the attentive person stood with their body facing away from the subjects but their head turned towards the subject while the inattentive person stood with their body facing the subject but their head turned away. Horses chose the attentive person significantly more often using the body cue, head cue, and eye cue but not the mixed cue. This result suggests that domestic horses are highly sensitive to human attentional cues, including gaze. The possible role of evolutionary and environmental factors in the development of this ability is discussed.  
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  Call Number Equine Behaviour @ team @ Serial 4984  
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Author Stone, S. url  doi
openurl 
  Title Human facial discrimination in horses: can they tell us apart? Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue Pages 51-61  
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  Abstract Abstract  The human–horse relationship has a long evolutionary history. Horses continue to play a pivotal role in the lives of humans and it is common for humans to think their horses recognize them by face. If a horse can distinguish his/her human companion from other humans, then evolution has supplied the horse with a very adaptive cognitive ability. The current study used operant conditioning trials to examine whether horses could discriminate photographed human faces and transfer this facial recognition ability a novel setting. The results indicated the horses (a) learned to discriminate photographs of the unrelated individuals, fraternal twins, and identical twins and (b) demonstrated transfer of facial recognition by spending more time with their S+ woman in the field test.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5004  
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Author Hothersall, B.; Gale, E.; Harris, P.; Nicol, C. doi  openurl
  Title Cue use by foals (Equus caballus) in a discrimination learning task Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue 1 Pages 63-74  
  Keywords  
  Abstract Abstract  Discrimination learning studies suggest that horses learn more easily using spatial than visible object-specific (OS) cues. However, spatial cues have generally confounded intra-array, distal and/or egocentric spatial information. It is also unclear whether conflicting cues compete for association or are redundantly encoded, and furthermore, the influence of prior experiences or training has not been quantified so far. We examined the effect of cue modality on unweaned foals’ performance in a discrimination learning task. After a pilot study confirmed that horses could perform the required OS cue discrimination, nine foals learnt to find food in one of three covered buckets, in any of four positions within a test arena. In Stage 1 the rewarded bucket was signified both by OS cues (pattern) and by relative spatial cues (position). On reaching criterion, cues were separated and foals had to ignore the inappropriate cue (Stage 2). Foals assigned to follow spatial cues (n = 5) completed Stage 2 faster than foals for whom OS cues remained consistent (n = 4). Spatial group foals all reached criterion without delay; no foal in the OS group reached criterion within the testing period. OS group foals initially persisted in responding to the previously correct position, adopting spatially-based strategies when this proved unsuccessful. The findings show for the first time that, even in the absence of distal spatial information, intra-array spatial cues were more salient than OS cues for foals in a food-finding task and learning appeared rather inflexible.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5082  
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Author Hopewell, L.; Leaver, L.; Lea, S.; Wills, A. url  doi
openurl 
  Title Grey squirrels (Sciurus carolinensis) show a feature-negative effect specific to social learning Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue 2 Pages 219-227  
  Keywords Feature-negative effect – Grey squirrels – Social learning – Caching  
  Abstract Abstract  Previous laboratory studies on social learning suggest that some animals can learn more readily if they first observe a conspecific demonstrator perform the task unsuccessfully and so fail to obtain a food reward than if they observe a successful demonstrator that obtains the food. This effect may indicate a difference in how easily animals are able to associate different outcomes with the conspecific or could simply be the result of having food present in only some of the demonstrations. To investigate we tested a scatter-hoarding mammal, the eastern grey squirrel, on its ability to learn to choose between two pots of food after watching a conspecific remove a nut from one of them on every trial. Squirrels that were rewarded for choosing the opposite pot to the conspecific chose correctly more frequently than squirrels rewarded for choosing the same pot (a feature-negative effect). Another group of squirrels was tested on their ability to choose between the two pots when the rewarded option was indicated by a piece of card. This time, squirrels showed no significant difference in their ability to learn to choose the same or the opposite pot. The results add to anecdotal reports that grey squirrels can learn by observing a conspecific and suggest that even when all subjects are provided with demonstrations with the same content, not all learning occurs equally. Prior experience or expectations of the association between a cue (a conspecific) and food influences what can be learned through observation whilst previously unfamiliar cues (the card) can be associated more readily with any outcome.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5108  
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Author Gaunet, F. url  doi
openurl 
  Title How do guide dogs and pet dogs (Canis familiaris) ask their owners for their toy and for playing? Type Journal Article
  Year 2010 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume (up) 13 Issue 2 Pages 311-323  
  Keywords Dog – Deictic behaviour – Intentional communication – Guide dogs – Socialisation – Play – Social cognition  
  Abstract Abstract  When apes are not fully understood by humans, they persist with attempts to communicate, elaborating their behaviours to better convey their meaning. Such abilities have never been investigated in dogs. The present study aimed to clarify any effect of the visual attentional state of the owner on dogs’ (Canis familiaris) social-communicative signals for interacting with humans, and to determine whether dogs persist and elaborate their behaviour in the face of failure to communicate a request. Gaze at a hidden target or at the owner, gaze alternation between a hidden target and the owner, vocalisations and contacts in 12 guide and 12 pet dogs were analysed (i) when the dogs were asked by their owners (blind or sighted) to fetch their inaccessible toy and (ii) when the dogs were subsequently given an unfamiliar object (apparent unsuccessful communication) or their toy (apparent successful communication). No group differences were found, indicating no effect of the visual status of the owner on the dogs’ socio-communicative modes (i.e. no sensitivity to human visual attention). Results, however, suggest that the dogs exhibited persistence (but not elaboration) in their “showing” behaviours in each condition, except that in which the toy was returned. Thus, their communication was about a specific item in space (the toy). The results suggest that dogs possess partially intentional non-verbal deictic abilities: (i) to get their inaccessible toy, the dogs gazed at their owners as if to trigger their attention; gaze alternation between the owner and the target direction, and two behaviours directed at the target were performed, apparently to indicate the location of the hidden toy; (ii) after the delivery of the toy, the dogs behaved as if they returned to the play routine, gazing at their owner whilst holding their toy. In conclusion, this study shows that dogs possess partially intentional non-verbal deictic abilities: they exhibit successive visual orienting between a partner and objects, apparent attention-getting behaviours, no sensitivity to the visual status of humans for communication, and persistence in (but no elaboration of) communicative behaviours when apparent attempts to “manipulate” the human partner fail.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5113  
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